174 lines
19 KiB
XML
174 lines
19 KiB
XML
<document id="27A1F83E9F3397072024B67EC1AB6CF3" ID-CLB-Dataset="29175" ID-DOI="10.11646/zootaxa.4453.1.1" ID-GBIF-Dataset="348e1a52-72a5-4809-b533-a158fc7a298a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1454920" ID-ZooBank="866690A9-0462-4892-AE29-9AAC623F87B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539178978648" checkinUser="plazi" docAuthor="Paweł Jałoszyński" docDate="2018" docId="2161879CFF808A55FF7A307C627FDFC0" docLanguage="en" docName="zootaxa.4453.1.1.pdf.imf" docOrigin="Zootaxa 4453 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Papusus Casey" docType="treatment" docVersion="8" lastPageNumber="71" masterDocId="DD58FFE4FFC68A13FFED3418614DD933" masterDocTitle="World genera of Mastigitae: review of morphological structures and new ecological data (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="119" masterPageNumber="1" pageNumber="71" updateTime="1698572836684" updateUser="ExternalLinkService">
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<mods:titleInfo id="9C9B323B7284149422AAA148FBDFA0FD">
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<mods:title id="8FE22ED9D855162A80AE587CB78A53BF">World genera of Mastigitae: review of morphological structures and new ecological data (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="FB3C4990CEF47C92929F3681D9C47165" type="personal">
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<mods:namePart id="C9EEFF24D197A34849B92EBB046AE385">Paweł Jałoszyński</mods:namePart>
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<mods:title id="665C132A3E3677C07726A7F4B52E9DAB">Zootaxa</mods:title>
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<mods:part id="2C45189AD09BC0E1563A1682CECD1782">
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<mods:date id="CA225C92C98EE2DBC4DF4BC5508C6450">2018</mods:date>
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<mods:number id="C00069213FF5C85221770DA968AFDEC0">2018-07-30</mods:number>
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<mods:detail id="F6C7078296F677F30CFF28B7156A4A03" type="volume">
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<mods:number id="B530169791A3573F08DBB64DE6CBF088">4453</mods:number>
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<mods:identifier id="628349CFBD62312BB9B3A4DBEAD27402" type="CLB-Dataset">29175</mods:identifier>
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<mods:identifier id="E6784BBB45593B386B44DDD4D7B4A59F" type="DOI">10.11646/zootaxa.4453.1.1</mods:identifier>
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<mods:identifier id="A7C76BB2B217457DE5A429F5D169ECB5" type="GBIF-Dataset">348e1a52-72a5-4809-b533-a158fc7a298a</mods:identifier>
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<mods:identifier id="1723C66E1070E82B806639DA418C8F75" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8CED1A4BA94271BDACDD557DF349BEC7" type="Zenodo-Dep">1454920</mods:identifier>
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<treatment id="2161879CFF808A55FF7A307C627FDFC0" ID-DOI="http://doi.org/10.5281/zenodo.5976952" ID-GBIF-Taxon="148669760" ID-Zenodo-Dep="5976952" LSID="urn:lsid:plazi:treatment:2161879CFF808A55FF7A307C627FDFC0" httpUri="http://treatment.plazi.org/id/2161879CFF808A55FF7A307C627FDFC0" lastPageNumber="71" pageId="70" pageNumber="71">
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<subSubSection id="E1D26501FF808A55FF7A307C601DDD4D" box="[151,336,1124,1150]" pageId="70" pageNumber="71" type="nomenclature">
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<paragraph id="A977368AFF808A55FF7A307C601DDD4D" blockId="70.[151,336,1124,1150]" box="[151,336,1124,1150]" pageId="70" pageNumber="71">
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<heading id="F23F81E6FF808A55FF7A307C601DDD4D" bold="true" box="[151,336,1124,1150]" fontSize="11" level="1" pageId="70" pageNumber="71" reason="1">
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<taxonomicName id="6EC84D09FF808A55FF7A307C601DDD4D" ID-CoL="8XNK4" authority="Casey" authorityName="Casey" box="[151,336,1124,1150]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
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||
<emphasis id="9BBCEA98FF808A55FF7A307C601DDD4D" bold="true" box="[151,336,1124,1150]" pageId="70" pageNumber="71">
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<emphasis id="9BBCEA98FF808A55FF7A307C61B0DD4D" bold="true" box="[151,253,1124,1150]" italics="true" pageId="70" pageNumber="71">Papusus</emphasis>
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Casey
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E1D26501FF808A55FF7A30B36022DDCD" pageId="70" pageNumber="71" type="reference_group">
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<paragraph id="A977368AFF808A55FF7A30B36022DDCD" blockId="70.[151,1436,1194,1278]" pageId="70" pageNumber="71">
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<taxonomicName id="6EC84D09FF808A55FF7A30B3603FDDF3" ID-CoL="8XNK4" authority="CASEY, 1897" authorityName="Casey" authorityYear="1897" box="[151,370,1194,1216]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="4" phylum="Arthropoda" rank="genus">
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<emphasis id="9BBCEA98FF808A55FF7A30B361A0DDF3" box="[151,237,1195,1216]" italics="true" pageId="70" pageNumber="71">Papusus</emphasis>
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<bibRefCitation id="CD594B7BFF808A55FF1E30B2603FDDF3" box="[243,370,1194,1216]" pageId="70" pageNumber="71" refId="ref51905" refString="CASEY, T. L. (1897) COLEOPTEROLOGICAL NOTICES, VII. Annals of the New York Academy of Sciences, 9, 285 - 684." type="journal article">Casey, 1897</bibRefCitation>
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</taxonomicName>
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: 542. Type species:
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<taxonomicName id="6EC84D09FF808A55FDD030B36217DDF3" ID-CoL="4CLVR" authority="CASEY, 1897" authorityName="Casey" authorityYear="1897" box="[573,858,1194,1216]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="112" phylum="Arthropoda" rank="species" species="macer">
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<emphasis id="9BBCEA98FF808A55FDD030B36395DDF3" box="[573,728,1195,1216]" italics="true" pageId="70" pageNumber="71">Papusus macer</emphasis>
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<bibRefCitation id="CD594B7BFF808A55FD3330B26217DDF3" box="[734,858,1194,1216]" pageId="70" pageNumber="71" refId="ref51905" refString="CASEY, T. L. (1897) COLEOPTEROLOGICAL NOTICES, VII. Annals of the New York Academy of Sciences, 9, 285 - 684." type="journal article">Casey, 1897</bibRefCitation>
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</taxonomicName>
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(monotypy). Note: Genus transferred to
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<taxonomicName id="6EC84D09FF808A55FB1530B26431DDF3" ID-CoL="8VRB7" authorityName="Leach" authorityYear="1815" box="[1272,1404,1194,1216]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="tribe" tribe="Scydmaenini">Scydmaenini</taxonomicName>
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by
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<bibRefCitation id="CD594B7BFF808A55FF2A30D2601DDDEC" box="[199,336,1225,1247]" pageId="70" pageNumber="71" refId="ref52418" refString="FRANz, H. (1985) REVISION CASEYSCHER SCYdMAENIdENTYPEN. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, ABTEILUNG I, 194, 149 - 186." type="journal article">Franz (1985)</bibRefCitation>
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; synonymized with
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<taxonomicName id="6EC84D09FF808A55FDC930D163F9DDEC" ID-CoL="8XM9V" box="[548,692,1225,1247]" class="Insecta" family="Staphylinidae" genus="Leptochromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis id="9BBCEA98FF808A55FDC930D163F9DDEC" box="[548,692,1225,1247]" italics="true" pageId="70" pageNumber="71">Leptochromus</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="CD594B7BFF808A55FD0D30D26224DDEC" box="[736,873,1225,1247]" pageId="70" pageNumber="71" refId="ref52474" refString="FRANz, H. (1995) REVISION dER VON SHARP (1887) AUS ZENTRALAMERIkA BESCHRIEBEN SCYdMAENIdEN (COLEOPTERA: SCYdMAENIdAE). Koleopterologische Rundschau, 65, 47 - 83." type="journal article">Franz (1995)</bibRefCitation>
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; resurrected as valid name and placed in Clidicini by O’Keefe (1998).
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</paragraph>
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</subSubSection>
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<subSubSection id="E1D26501FF808A55FF7A313263E3DCB8" pageId="70" pageNumber="71" type="diagnosis">
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<paragraph id="A977368AFF808A55FF7A31326457DC71" blockId="70.[151,1436,1322,1779]" box="[151,1306,1322,1347]" pageId="70" pageNumber="71">
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<emphasis id="9BBCEA98FF808A55FF7A3132605CDC70" bold="true" box="[151,273,1322,1347]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
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<taxonomicName id="6EC84D09FF808A55FEF53134603ADC70" box="[280,375,1324,1347]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis id="9BBCEA98FF808A55FEF53134603ADC70" box="[280,375,1324,1347]" italics="true" pageId="70" pageNumber="71">Papusus</emphasis>
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</taxonomicName>
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is the only genus of Papusini; diagnosis and characteristics as for tribe,
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<emphasis id="9BBCEA98FF808A55FB7331326459DC70" box="[1182,1300,1322,1347]" italics="true" pageId="70" pageNumber="71">vide supra</emphasis>
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.
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</paragraph>
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<paragraph id="A977368AFF808A55FF2A315663E3DCB8" blockId="70.[151,1436,1322,1779]" pageId="70" pageNumber="71">
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<emphasis id="9BBCEA98FF808A55FF2A3156637BDC54" bold="true" box="[199,566,1358,1383]" pageId="70" pageNumber="71">Composition and distribution.</emphasis>
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<taxonomicName id="6EC84D09FF808A55FDAD315763D2DC55" box="[576,671,1359,1382]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis id="9BBCEA98FF808A55FDAD315763D2DC55" box="[576,671,1359,1382]" italics="true" pageId="70" pageNumber="71">Papusus</emphasis>
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</taxonomicName>
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comprises nine species distributed in
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<collectingRegion id="6B0CF868FF808A55FBB0315665FEDC54" box="[1117,1203,1358,1383]" country="Mexico" name="Mexico" pageId="70" pageNumber="71">
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<collectingCountry id="D1DF761AFF808A55FBB0315665FEDC54" box="[1117,1203,1358,1383]" name="Mexico" pageId="70" pageNumber="71">Mexico</collectingCountry>
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</collectingRegion>
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and
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<collectingCountry id="D1DF761AFF808A55FB023157646ADC54" box="[1263,1319,1359,1383]" name="United States of America" pageId="70" pageNumber="71">USA</collectingCountry>
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(
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<collectingRegion id="6B0CF868FF808A55FAD4315664D5DC54" box="[1337,1432,1358,1383]" country="United States of America" name="Arizona" pageId="70" pageNumber="71">Arizona</collectingRegion>
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,
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<collectingRegion id="6B0CF868FF808A55FF7A316A6041DCB8" box="[151,268,1394,1419]" country="United States of America" name="California" pageId="70" pageNumber="71">California</collectingRegion>
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,
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<collectingRegion id="6B0CF868FF808A55FEFB316A60CCDCB8" box="[278,385,1394,1419]" country="United States of America" name="Colorado" pageId="70" pageNumber="71">Colorado</collectingRegion>
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,
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<collectingRegion id="6B0CF868FF808A55FE60316A60A8DCB8" box="[397,485,1394,1419]" country="United States of America" name="Nevada" pageId="70" pageNumber="71">Nevada</collectingRegion>
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,
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<collectingRegion id="6B0CF868FF808A55FE1D316A6365DCB8" box="[496,552,1394,1419]" country="United States of America" name="Utah" pageId="70" pageNumber="71">Utah</collectingRegion>
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) (
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<figureCitation id="31F32A0FFF808A55FDD2316A63EFDCB8" box="[575,674,1394,1419]" captionStart="FIGURE 140" captionStartId="63.[151,250,1907,1929]" captionTargetBox="[151,1436,1338,1885]" captionTargetId="figure@63.[151,1436,1338,1885]" captionTargetPageId="63" captionText="FIGURE 140. Papusini, distribution of extant species of Papusus." httpUri="https://zenodo.org/record/1454998/files/figure.png" pageId="70" pageNumber="71">Fig. 140</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="E1D26501FF808A55FF2A318E628EDFFC" pageId="70" pageNumber="71" type="description">
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<paragraph id="A977368AFF808A55FF2A318E628EDFFC" blockId="70.[151,1436,1322,1779]" pageId="70" pageNumber="71">
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<emphasis id="9BBCEA98FF808A55FF2A318E60CADC9C" bold="true" box="[199,391,1430,1455]" pageId="70" pageNumber="71">Natural history.</emphasis>
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Species of
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<taxonomicName id="6EC84D09FF808A55FDFD318F6322DC9D" box="[528,623,1431,1454]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
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<emphasis id="9BBCEA98FF808A55FDFD318F6322DC9D" box="[528,623,1431,1454]" italics="true" pageId="70" pageNumber="71">Papusus</emphasis>
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</taxonomicName>
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are unique among all
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<taxonomicName id="6EC84D09FF808A55FC99318E655ADC9C" authorityName="Leach" authorityYear="1815" box="[884,1047,1430,1455]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
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in inhabiting the driest deserts of North America (Great Basin Desert, including the Death and Saline Valleys, Mojave Desert,
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<collectingRegion id="6B0CF868FF808A55FB5E31A26456DCE0" box="[1203,1307,1466,1491]" country="United States of America" name="Colorado" pageId="70" pageNumber="71">Colorado</collectingRegion>
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Desert and
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<collectingRegion id="6B0CF868FF808A55FF7A31C761ABDCC4" box="[151,230,1503,1527]" country="Mexico" name="Sonora" pageId="70" pageNumber="71">Sonora</collectingRegion>
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Desert); moreover, adults were collected only during the hottest seasons of the year (O’Keefe 2003). Beetles were collected predominantly by pitfall traps set in the Shedscale Scrub and Creosote Bush Scrub communities, and in deserts with
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<taxonomicName id="6EC84D09FF808A55FDD9323F63A0DF0D" authority=", Amrosia" authorityName="Amrosia" box="[564,749,1575,1598]" class="Magnoliopsida" family="Asparagaceae" genus="Agave" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
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<emphasis id="9BBCEA98FF808A55FDD9323F6337DF0D" box="[564,634,1575,1598]" italics="true" pageId="70" pageNumber="71">Agave</emphasis>
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,
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<emphasis id="9BBCEA98FF808A55FD62323F63A0DF0D" box="[655,749,1575,1598]" italics="true" pageId="70" pageNumber="71">Amrosia</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="6EC84D09FF808A55FCEF323E6211DF0C" box="[770,860,1574,1599]" class="Magnoliopsida" family="Amaranthaceae" genus="Atriplex" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
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<emphasis id="9BBCEA98FF808A55FCEF323E6211DF0C" box="[770,860,1574,1599]" italics="true" pageId="70" pageNumber="71">Atriplex</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="6EC84D09FF808A55FC9F323F62F9DF0D" box="[882,948,1575,1598]" class="Magnoliopsida" family="Asparagaceae" genus="Yucca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
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<emphasis id="9BBCEA98FF808A55FC9F323F62F9DF0D" box="[882,948,1575,1598]" italics="true" pageId="70" pageNumber="71">Yucca</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="6EC84D09FF808A55FC24323F650ADF0D" box="[969,1095,1575,1598]" class="Magnoliopsida" family="Fouquieriaceae" genus="Fouquieria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
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<emphasis id="9BBCEA98FF808A55FC24323F650ADF0D" box="[969,1095,1575,1598]" italics="true" pageId="70" pageNumber="71">Fouquieria</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="6EC84D09FF808A55FBB1323E65A7DF0C" authorityName="N.L.Britton & J.N.Rose" authorityYear="1909" box="[1116,1258,1574,1599]" class="Magnoliopsida" family="Cactaceae" genus="Pachycereus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
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<emphasis id="9BBCEA98FF808A55FBB1323E65A7DF0C" box="[1116,1258,1574,1599]" italics="true" pageId="70" pageNumber="71">Pachycereus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="6EC84D09FF808A55FB12323E6428DF0C" authorityName="Bentham" authorityYear="1844" box="[1279,1381,1574,1599]" class="Magnoliopsida" family="Fabaceae" genus="Lysiloma" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
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<emphasis id="9BBCEA98FF808A55FB12323E6428DF0C" box="[1279,1381,1574,1599]" italics="true" pageId="70" pageNumber="71">Lysiloma</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="6EC84D09FF808A55FF7A32536051DF51" authorityName="Riccobono" authorityYear="1909" box="[151,284,1611,1634]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="9BBCEA98FF808A55FF7A32536051DF51" box="[151,284,1611,1634]" italics="true" pageId="70" pageNumber="71">Stenocereus</emphasis>
|
||
</taxonomicName>
|
||
. Adults were found most often in alkali scrub or sand habitats with scattered vegetation, but not on open sand dunes. O'Keefe (2003) observed beetles walking on sand at night and found individuals under volcanic rocks
|
||
<quantity id="6E309B6FFF808A55FF36328B6006DF99" box="[219,331,1683,1707]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="70" pageNumber="71" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 cm</quantity>
|
||
in diameter. He concluded that "in the Great Basin Desert adults of
|
||
<taxonomicName id="6EC84D09FF808A55FBA3328C65E0DF98" box="[1102,1197,1684,1707]" class="Insecta" family="Staphylinidae" genus="Papusus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9BBCEA98FF808A55FBA3328C65E0DF98" box="[1102,1197,1684,1707]" italics="true" pageId="70" pageNumber="71">Papusus</emphasis>
|
||
</taxonomicName>
|
||
are active during the warmest time of the year and are most likely nocturnal" (O'Keefe 2003).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E1D26501FF808A55FF2A32C2627FDFC0" box="[199,818,1754,1779]" pageId="70" pageNumber="71" type="discussion">
|
||
<paragraph id="A977368AFF808A55FF2A32C2627FDFC0" blockId="70.[151,1436,1322,1779]" box="[199,818,1754,1779]" pageId="70" pageNumber="71">
|
||
<emphasis id="9BBCEA98FF808A55FF2A32C26076DFC0" bold="true" box="[199,315,1754,1779]" pageId="70" pageNumber="71">Remarks.</emphasis>
|
||
All species were revised by O'Keefe (2003).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |