292 lines
39 KiB
XML
292 lines
39 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.973.53940" ID-GBIF-Dataset="fee87159-37fa-48c9-a745-e1656a387087" ID-PMC="PMC7550394" ID-Pensoft-Pub="1313-2970-973-89" ID-Pensoft-UUID="5D78426E113D5C2FADE90EA6039CE67D" ID-PubMed="33110374" ID-ZooBank="9471AA9B4FED45A4800FB79B9AA39949" ModsDocID="1313-2970-973-89" checkinTime="1601957130496" checkinUser="pensoft" docAuthor="Shavrin, Alexey V. & Yamamoto, Shuhei" docDate="2020" docId="F63B56709B345E019B4B7FD3C750BD7A" docLanguage="en" docName="ZooKeys 973: 89-101" docOrigin="ZooKeys 973" docSource="http://dx.doi.org/10.3897/zookeys.973.53940" docTitle="Anthobium alekseevi Shavrin & Yamamoto 2020, sp. nov." docType="treatment" docUuid="A1797770-BE2C-4B9A-99B1-5D93D54B7943" docUuidSource="ZooBank" docVersion="4" id="5D78426E113D5C2FADE90EA6039CE67D" lastPageNumber="89" masterDocId="5D78426E113D5C2FADE90EA6039CE67D" masterDocTitle="A remarkable new species of the rove beetle genus Anthobium Leach, 1819 from Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)" masterLastPageNumber="101" masterPageNumber="89" pageNumber="89" updateTime="1668169574041" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A remarkable new species of the rove beetle genus Anthobium Leach, 1819 from Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shavrin, Alexey V.</mods:namePart>
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<mods:affiliation>Institute of Life Sciences and Technologies, Daugavpils University, Vienibas 13, Daugavpils, LV- 5401, Latvia</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6487-4530</mods:nameIdentifier>
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<mods:nameIdentifier type="email">ashavrin@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yamamoto, Shuhei</mods:namePart>
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<mods:affiliation>The Hokkaido University Museum, Kita 8, Nishi 5, Kita-ku, Sapporo 060 - 0808, Japan</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4162-8457</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>973</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>89</mods:start>
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<mods:end>101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.973.53940</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.973.53940</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-973-89</mods:identifier>
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<mods:identifier type="ZooBank">9471AA9B4FED45A4800FB79B9AA39949</mods:identifier>
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<mods:identifier type="Pensoft-UUID">5D78426E113D5C2FADE90EA6039CE67D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="168237436" LSID="urn:lsid:zoobank.org:act:A1797770-BE2C-4B9A-99B1-5D93D54B7943" httpUri="http://treatment.plazi.org/id/F63B56709B345E019B4B7FD3C750BD7A" lastPageNumber="89" pageId="0" pageNumber="89">
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<subSubSection pageId="0" pageNumber="89" type="nomenclature">
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<paragraph pageId="0" pageNumber="89">
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†
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<taxonomicName LSID="F63B5670-9B34-5E01-9B4B-7FD3C750BD7A" authority="Shavrin & Yamamoto" authorityName="Shavrin & Yamamoto" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthobium alekseevi" order="Coleoptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="alekseevi" status="sp. nov.">Anthobium alekseevi Shavrin & Yamamoto</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="89">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">Figures 1-4</figureCitation>
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<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. Anthobium alekseevi sp. nov., habitus (lateral view). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures5-6" httpUri="https://binary.pensoft.net/fig/459301" pageId="0" pageNumber="89">, 5, 6</figureCitation>
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<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. Anthobium alekseevi sp. nov. 7 legs 8 protarsus. Scale bars: 1.0 mm (7); 0.1 mm (8). Abbreviations: pt 1 - pt 5 = protarsomeres 1 - 5." figureDoi="10.3897/zookeys.973.53940.figures7-8" httpUri="https://binary.pensoft.net/fig/459302" pageId="0" pageNumber="89">, 7, 8</figureCitation>
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<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">, 9-13</figureCitation>
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<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">, 14-19</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="type material">
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<paragraph pageId="0" pageNumber="89">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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<emphasis bold="true" pageId="0" pageNumber="89">Holotype</emphasis>
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: male, complete specimen as inclusion in a piece of small yellow Baltic amber, 11.0 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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0.7 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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0.5 mm in size (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">1</figureCitation>
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,
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">2</figureCitation>
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), with glued small paper on side of an amber labeled
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<normalizedToken originalValue="“AWI148”">"AWI148"</normalizedToken>
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and additional labels within a plastic envelope: "AWI-148 |
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<taxonomicName authorityName="MacLeay" authorityYear="1825" lsidName="" pageId="0" pageNumber="89" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
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| gen. nov. | (mesotibia!)" <handwritten>, "
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<emphasis bold="true" pageId="0" pageNumber="89">HOLOTYPE</emphasis>
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<taxonomicName authorityName="Leach" authorityYear="1819" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthobium" order="Coleoptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="89">Anthobium</emphasis>
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</taxonomicName>
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<taxonomicName authorityName="Shavrin & Yamamoto" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="alekseevi" order="Coleoptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="alekseevi">
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<emphasis italics="true" pageId="0" pageNumber="89">alekseevi</emphasis>
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</taxonomicName>
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sp. nov. | Shavrin A. & Yamamoto S. des. 2020" <red rectangular label, printed> (PIN).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="preservation">
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<paragraph pageId="0" pageNumber="89">Preservation.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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The specimen is poorly visible because it is partially covered with white microbubbles, and some details of the structure of the body are not visible: head, median portion of pronotum and scutellum, ventral side of the body and abdomen (Figs
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">1</figureCitation>
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,
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">2</figureCitation>
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). The basal part of the abdomen ventrally is covered by left hind wing.
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</paragraph>
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<caption doi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89" start="Figures 1–4" startId="F1">
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<paragraph pageId="0" pageNumber="89">
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<emphasis bold="true" pageId="0" pageNumber="89">Figures 1-4.</emphasis>
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<taxonomicName authorityName="Shavrin & Yamamoto" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthobium alekseevi" order="Coleoptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="alekseevi">
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<emphasis italics="true" pageId="0" pageNumber="89">Anthobium alekseevi</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="89">1, 2</emphasis>
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amber specimen with inclusion
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<emphasis bold="true" pageId="0" pageNumber="89">3</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="89">4</emphasis>
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habitus, dorsoventral view. Scale bar: 1.0 mm (
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<emphasis bold="true" pageId="0" pageNumber="89">3, 4</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="locality and horizon">
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<paragraph pageId="0" pageNumber="89">Locality and horizon.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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Baltic amber from Kaliningrad Area, westernmost Russia; mid-Eocene (ca 44 Ma;
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<bibRefCitation author="Wappler, T" journalOrPublisher="University of Kwazulu-Natal, South African National Biodiversity Institute, Pretoria" pageId="0" pageNumber="89" refId="B18" refString="Wappler, T, 2005. The age of Baltic amber: could Eckfeld resolve this problem? In: Brothers D, Mostovski M (Eds) Fossils X3, 3rd International Congress of Palaeoentomology with 2nd International Meeting on Palaeoarthropodology and 2nd World Congress on Amber and its Inclusions. Programme and Abstracts, 7th to 11th Feb 2005, Pretoria South Africa. University of Kwazulu-Natal, South African National Biodiversity Institute, Pretoria" title="The age of Baltic amber: could Eckfeld resolve this problem? In: Brothers D, Mostovski M (Eds) Fossils X 3, 3 rd International Congress of Palaeoentomology with 2 nd International Meeting on Palaeoarthropodology and 2 nd World Congress on Amber and its Inclusions. Programme and Abstracts, 7 th to 11 th Feb 2005, Pretoria South Africa." year="2005">Wappler 2005</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="89" type="description">
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<paragraph pageId="0" pageNumber="89">Description.</paragraph>
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<paragraph pageId="0" pageNumber="89">
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Measurements: maximum width of head including eyes: 1.30; length of head (from base of labrum to neck constriction along head midline in dorsal view): 0.75; ocular length: 0.40; length
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<normalizedToken originalValue="×">x</normalizedToken>
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width of segments III and IV of maxillary palpi: III: 0.15
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<normalizedToken originalValue="×">x</normalizedToken>
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0.10, IV: 0.30
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<normalizedToken originalValue="×">x</normalizedToken>
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0.10; length of antenna: 2.70; length of pronotum: 1.35; maximum width of pronotum: 1.75; sutural length of elytra from apex of scutellum to posterior margin of sutural angle: 2.60; length of elytron from basal to apical margin: 2.95; maximum width of elytra: 2.30; length of metatibia: 1.60; length of metatarsus: 0.80; maximum width of abdomen (at segment IV): 2.10; total length (from anterior margin of clypeus to apex of abdomen): ~5.40.
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</paragraph>
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<paragraph pageId="0" pageNumber="89">
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Body oblong, moderately wide, shiny (Fig.
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">3</figureCitation>
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); body laterally as in Figures
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<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. Anthobium alekseevi sp. nov., habitus (lateral view). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures5-6" httpUri="https://binary.pensoft.net/fig/459301" pageId="0" pageNumber="89">5</figureCitation>
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,
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<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. Anthobium alekseevi sp. nov., habitus (lateral view). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures5-6" httpUri="https://binary.pensoft.net/fig/459301" pageId="0" pageNumber="89">6</figureCitation>
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,
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<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">9</figureCitation>
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, and
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<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">10</figureCitation>
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; body dorsolaterally as in Figure
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<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">4</figureCitation>
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; body ventrally as in Figures
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<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">11</figureCitation>
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and
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<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">12</figureCitation>
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; forebody as in Figure
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<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">14</figureCitation>
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. Body and antennomeres 3-11, brown; legs and mouthparts reddish-brown; antennomeres 1 and 2 yellow-brown.
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</paragraph>
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<caption doi="10.3897/zookeys.973.53940.figures5-6" httpUri="https://binary.pensoft.net/fig/459301" pageId="0" pageNumber="89" start="Figures 5, 6" startId="F2">
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<paragraph pageId="0" pageNumber="89">
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<emphasis bold="true" pageId="0" pageNumber="89">Figures 5, 6.</emphasis>
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<taxonomicName authorityName="Shavrin & Yamamoto" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthobium alekseevi" order="Coleoptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="alekseevi">
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<emphasis italics="true" pageId="0" pageNumber="89">Anthobium alekseevi</emphasis>
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</taxonomicName>
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sp. nov., habitus (lateral view). Scale bar: 1.0 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="89">
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Head transverse (Figs
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<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">15</figureCitation>
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,
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<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">16</figureCitation>
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), 1.7 times as wide as long; anterior portion of frons with obliquely elevated supra-antennal prominences and moderately wide impressions behind them; middle portion slightly elevated, with distinct, deep, narrow dorsal tentorial pits (grooves) in front of ocelli, diagonally stretching apicad to level of middle length of eyes; basal portion with distinct, narrow impression between ocelli; postocular ridge distinct, acute, located relatively close to posterior margin of eye, if see laterally (Figs
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<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. Anthobium alekseevi sp. nov., habitus (lateral view). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures5-6" httpUri="https://binary.pensoft.net/fig/459301" pageId="0" pageNumber="89">5</figureCitation>
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,
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<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">9</figureCitation>
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,
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">18</figureCitation>
|
||
). Eyes large, convex. Ocelli large, situated at about level of postocular ridges; distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye (Figs
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">15</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">16</figureCitation>
|
||
). Labrum wide, transverse. Mentum and labium narrow, with distinctly elongate apical labial palpomere (Fig.
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">12</figureCitation>
|
||
). Preapical segment of maxillary palpus moderately long, 1.5 times as long as wide, and as wide as apical segment; apical segment about twice as long as preceding segment, from middle gradually narrowing toward apex (Figs
|
||
<figureCitation captionStart="Figures 5, 6" captionStartId="F2" captionText="Figures 5, 6. Anthobium alekseevi sp. nov., habitus (lateral view). Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures5-6" httpUri="https://binary.pensoft.net/fig/459301" pageId="0" pageNumber="89">5</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">11</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">18</figureCitation>
|
||
). Gular sutures with shortest distance located at level of posterior third of eyes (Fig.
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">12</figureCitation>
|
||
). Antenna moderately short, exceeding apical third of elytra, with elongate antennomeres 5-10 and long preapical setae, antennomeres 3-11 covered by dense pubescence; basal antennomere moderately wide, 1.5 times as long as 2, antennomere 2 ovoid, about twice as long as wide and slightly shorter than 3, 3 and 4 as wide as 2, 5-7 slightly shorter than 4, 8 and 9 slightly shorter than 7, 10 distinctly shorter than 9, apical antennomere 1.6 times as long as 10, from middle gradually narrowed toward apex (Fig.
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="89">
|
||
Pronotum subrectangular, 1.2 times as wide as long, 1.3 times as wide as head, widest in middle, evenly rounded both anteriad and posteriad (Fig.
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">14</figureCitation>
|
||
); apical margin rounded, distinctly narrower than posterior margin; anterior angles widely rounded, slightly protruded anteriad; posterior angles obtuse; lateral edges bordered, without visible crenulation; disc with middle portion widely elevated, with indistinct, wide, semioval impression on mediobasal third, and lateral portions moderately wide, slightly explanate, each with deep pit in middle (Fig.
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">16</figureCitation>
|
||
). Laterobasal and basal portions of pronotum with dense and fine punctation (Fig.
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">3</figureCitation>
|
||
). Pronotal hypomeron and postcoxal process well developed; intercoxal process elongate and moderately wide; prosternum with distinct median carina (Figs
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">11</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">18</figureCitation>
|
||
). Scutellum large and wide, with rounded apex. Metaventrite wide, convex.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="89">
|
||
Elytra convex, slightly longer than wide, about twice as long as pronotum, indistinctly widened in middle, reaching basal margin of abdominal tergite VI, with widely rounded apical margins (Fig.
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">17</figureCitation>
|
||
); shoulders of elytra rounded; lateral portions narrow, explanate; surface of elytra without visible elevations. Punctation sparse and moderately small, each elytron with longitudinal rows of vague serial punctures in middle (Fig.
|
||
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Anthobium alekseevi sp. nov. 1, 2 amber specimen with inclusion 3 habitus, dorsal view 4 habitus, dorsoventral view. Scale bar: 1.0 mm (3, 4)." figureDoi="10.3897/zookeys.973.53940.figures1-4" httpUri="https://binary.pensoft.net/fig/459300" pageId="0" pageNumber="89">3</figureCitation>
|
||
). Hind wings fully developed.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="89">
|
||
Legs long (Fig.
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. Anthobium alekseevi sp. nov. 7 legs 8 protarsus. Scale bars: 1.0 mm (7); 0.1 mm (8). Abbreviations: pt 1 - pt 5 = protarsomeres 1 - 5." figureDoi="10.3897/zookeys.973.53940.figures7-8" httpUri="https://binary.pensoft.net/fig/459302" pageId="0" pageNumber="89">7</figureCitation>
|
||
); procoxae wide, protruding ventrad, contiguous; mesocoxae large, convex; metacoxae strongly transverse; pro-, meso-, and metatrochanter relatively narrow, elongate (Figs
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">11</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">12</figureCitation>
|
||
); all femora widest at about middle, profemora slightly wider than meso- and metafemora; pro- and mesotibiae about as long as femora; protibiae covered with dense long setae; protarsomeres 1-5 as in Fig.
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. Anthobium alekseevi sp. nov. 7 legs 8 protarsus. Scale bars: 1.0 mm (7); 0.1 mm (8). Abbreviations: pt 1 - pt 5 = protarsomeres 1 - 5." figureDoi="10.3897/zookeys.973.53940.figures7-8" httpUri="https://binary.pensoft.net/fig/459302" pageId="0" pageNumber="89">8</figureCitation>
|
||
; mesotibiae indistinctly curved in middle, inner side of each mesotibia with large and wide subtriangular, fin-shaped tooth in middle (Fig.
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. Anthobium alekseevi sp. nov. 7 legs 8 protarsus. Scale bars: 1.0 mm (7); 0.1 mm (8). Abbreviations: pt 1 - pt 5 = protarsomeres 1 - 5." figureDoi="10.3897/zookeys.973.53940.figures7-8" httpUri="https://binary.pensoft.net/fig/459302" pageId="0" pageNumber="89">7</figureCitation>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="89">arrows</emphasis>
|
||
); meso- and metatibiae covered by very dense, long and strong setae (including apex of tooth on each mesotibia); metatibiae with additional very long setae; metatibiae distinctly longer than metafemora, slightly widened in about middle, covered by dense, strong and long setae, with a few additional spines around apical margin; all tarsi 5-segmented, all tarsi combined shorter than tibia; tarsomeres 1-4 with long and dense lateral setation; apical tarsomere about as long as preceding three tarsomeres together; tarsal claws simple, without modifications (Fig.
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. Anthobium alekseevi sp. nov. 7 legs 8 protarsus. Scale bars: 1.0 mm (7); 0.1 mm (8). Abbreviations: pt 1 - pt 5 = protarsomeres 1 - 5." figureDoi="10.3897/zookeys.973.53940.figures7-8" httpUri="https://binary.pensoft.net/fig/459302" pageId="0" pageNumber="89">7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.973.53940.figures7-8" httpUri="https://binary.pensoft.net/fig/459302" pageId="0" pageNumber="89" start="Figures 7, 8" startId="F3">
|
||
<paragraph pageId="0" pageNumber="89">
|
||
<emphasis bold="true" pageId="0" pageNumber="89">Figures 7, 8.</emphasis>
|
||
<taxonomicName authorityName="Shavrin & Yamamoto" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthobium alekseevi" order="Coleoptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="alekseevi">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">Anthobium alekseevi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="89">7</emphasis>
|
||
legs
|
||
<emphasis bold="true" pageId="0" pageNumber="89">8</emphasis>
|
||
protarsus. Scale bars: 1.0 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="89">7</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="89">8</emphasis>
|
||
). Abbreviations: pt1-pt5 = protarsomeres 1-5.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="89">
|
||
Abdomen distinctly narrower than elytra (Fig.
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">17</figureCitation>
|
||
), from segment IV significantly narrowing apicad (Fig.
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">11</figureCitation>
|
||
); abdominal segment IX elongate. Abdomen ventrally as in Figures
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">11</figureCitation>
|
||
and
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">12</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="89">
|
||
Male. Protarsomeres 1-4 distinctly wide, with very long lateral setae (Fig.
|
||
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. Anthobium alekseevi sp. nov. 7 legs 8 protarsus. Scale bars: 1.0 mm (7); 0.1 mm (8). Abbreviations: pt 1 - pt 5 = protarsomeres 1 - 5." figureDoi="10.3897/zookeys.973.53940.figures7-8" httpUri="https://binary.pensoft.net/fig/459302" pageId="0" pageNumber="89">8</figureCitation>
|
||
). Apical margin of abdominal tergite VIII rounded (Fig.
|
||
<figureCitation captionStart="Figures 14–19" captionStartId="F5" captionText="Figures 14 - 19. Anthobium alekseevi sp. nov. 14 forebody, dorsal view 15 head, apical portion of pronotum and legs, frontal view 16 head, pronotum and antenna, dorsal view 17 elytra and abdomen, dorsal view 18 head and thorax, lateroventral view 19 apical portion of elytra, abdomen and legs, posterodorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures14-19" httpUri="https://binary.pensoft.net/fig/459304" pageId="0" pageNumber="89">17</figureCitation>
|
||
). Apical margin of sternite VIII sinuate (Fig.
|
||
<figureCitation captionStart="Figures 9–13" captionStartId="F4" captionText="Figures 9 - 13. Habitus of Anthobium alekseevi sp. nov. 9, 10 lateral view 11, 12 ventral view 13 dorsal view. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89">12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="89">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="89" type="etymology">
|
||
<paragraph pageId="0" pageNumber="89">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="89">Patronymic, the species is named to honor our colleague Vitalii I. Alekseev (Kaliningrad), great contributor to the knowledge of the fossil beetle fauna from Baltic amber.</paragraph>
|
||
<caption doi="10.3897/zookeys.973.53940.figures9-13" httpUri="https://binary.pensoft.net/fig/459303" pageId="0" pageNumber="89" start="Figures 9–13" startId="F4">
|
||
<paragraph pageId="0" pageNumber="89">
|
||
<emphasis bold="true" pageId="0" pageNumber="89">Figures 9-13.</emphasis>
|
||
Habitus of
|
||
<taxonomicName authorityName="Shavrin & Yamamoto" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthobium alekseevi" order="Coleoptera" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="alekseevi">
|
||
<emphasis italics="true" pageId="0" pageNumber="89">Anthobium alekseevi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="89">9, 10</emphasis>
|
||
lateral view
|
||
<emphasis bold="true" pageId="0" pageNumber="89">11, 12</emphasis>
|
||
ventral view
|
||
<emphasis bold="true" pageId="0" pageNumber="89">13</emphasis>
|
||
dorsal view. Scale bar: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |