164 lines
42 KiB
XML
164 lines
42 KiB
XML
<document id="9BB396F706EC54EAA6EC72F4C9966743" ID-DOI="10.11646/zootaxa.4508.1.5" ID-GBIF-Dataset="735a2fc4-2383-4e07-89cc-b4387bebd396" ID-ISSN="1175-5326" ID-Zenodo-Dep="3713918" ID-ZooBank="7C816EF3-658C-437D-82C0-E259782AB9A0" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584478799513" checkinUser="plazi" docAuthor="Waringer, Johann, Karaouzas, Ioannis & Malicky, Hans" docDate="2018" docId="F60D87F9A651E270BB8DEF8CFBDAFCA0" docLanguage="en" docName="zootaxa.4508.1.5.pdf.imf" docOrigin="Zootaxa 4508 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila gudrunae Malicky 1972" docType="treatment" docVersion="7" lastPageNumber="87" masterDocId="0A34FF81A650E272BB1AEC4AFFD8FFB4" masterDocTitle="The larvae of Rhyacophila tsurakiana Malicky 1984, Rhyacophila gudrunae Malicky 1972, and Rhyacophila biegelmeieri Malicky 1984, including an update for the larval key to the Greek species of genus Rhyacophila Stephens 1836 (Rhyacophilidae, Trichoptera)" masterLastPageNumber="100" masterPageNumber="85" pageNumber="86" updateTime="1698729507662" updateUser="plazi">
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<mods:title id="9E52C545A87ED8CEDFCAC0E0496BE69F">The larvae of Rhyacophila tsurakiana Malicky 1984, Rhyacophila gudrunae Malicky 1972, and Rhyacophila biegelmeieri Malicky 1984, including an update for the larval key to the Greek species of genus Rhyacophila Stephens 1836 (Rhyacophilidae, Trichoptera)</mods:title>
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<mods:namePart id="930F56ADF89DB78E5A91033C5CB4FDCE">Waringer, Johann</mods:namePart>
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<mods:namePart id="BE5C37551AD48D13FE0475AA05FF80FE">Karaouzas, Ioannis</mods:namePart>
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<mods:namePart id="97271450FC754169368A0A6283DB684F">Malicky, Hans</mods:namePart>
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<mods:date id="95BCE67FDF74F526B99F4DF39FD8E55D">2018</mods:date>
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<treatment id="F60D87F9A651E270BB8DEF8CFBDAFCA0" ID-DOI="http://doi.org/10.5281/zenodo.3717673" ID-GBIF-Taxon="162629280" ID-Zenodo-Dep="3717673" LSID="urn:lsid:plazi:treatment:F60D87F9A651E270BB8DEF8CFBDAFCA0" httpUri="http://treatment.plazi.org/id/F60D87F9A651E270BB8DEF8CFBDAFCA0" lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86">
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<subSubSection id="36BE6564A651E273BB8DEF8CFBE4FC54" box="[151,1084,966,992]" pageId="1" pageNumber="86" type="nomenclature">
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<paragraph id="7E1B36EFA651E273BB8DEF8CFBE4FC54" blockId="1.[151,1084,966,992]" box="[151,1084,966,992]" pageId="1" pageNumber="86">
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<heading id="25538183A651E273BB8DEF8CFBE4FC54" bold="true" box="[151,1084,966,992]" fontSize="11" level="1" pageId="1" pageNumber="86" reason="1">
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<emphasis id="4CD0EAFDA651E273BB8DEF8CFBE4FC54" bold="true" box="[151,1084,966,992]" pageId="1" pageNumber="86">
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Description of the final instar larva of
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<taxonomicName id="B9A44D6CA651E273B96CEF8CFBE4FC54" authority="Malicky 1972" authorityName="Malicky" authorityYear="1972" box="[630,1084,966,992]" class="Magnoliopsida" family="Lythraceae" genus="Rhyacophila" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="gudrunae">
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<emphasis id="4CD0EAFDA651E273B96CEF8CFC53FC54" bold="true" box="[630,907,966,992]" italics="true" pageId="1" pageNumber="86">Rhyacophila gudrunae</emphasis>
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<bibRefCitation id="1A354B1EA651E273B889EF8CFBE4FC54" author="Malicky, H." box="[915,1084,966,992]" pageId="1" pageNumber="86" pagination="25 - 68" refId="ref6339" refString="Malicky, H. (1972) Weitere neue Arten und Fundorte von westpalaarktischen Kocherfliegen (Trichoptera), vor allem aus dem ostlichen Mediterrangebiet. Mitteilungen der Entomologischen Gesellschaft Basel, 22, 25 - 68." type="journal article" year="1972">Malicky 1972</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="36BE6564A651E270BB8DE847FBDAFCA0" lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86" type="description">
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<paragraph id="7E1B36EFA651E273BB8DE847FF31FBDA" blockId="1.[151,1437,1037,2034]" pageId="1" pageNumber="86">
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<emphasis id="4CD0EAFDA651E273BB8DE847FED4FB92" bold="true" box="[151,268,1037,1062]" pageId="1" pageNumber="86">Biometry.</emphasis>
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Body length ranging from 20.0 to
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<quantity id="B95C9B0AA651E273B988E844FD20FB91" box="[658,760,1038,1062]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="1" pageNumber="86" unit="mm" value="21.7">21.7 mm</quantity>
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, head width from
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<quantity id="B95C9B0AA651E273B8D9E844FBA4FB91" box="[963,1148,1037,1062]" metricMagnitude="-3" metricUnit="m" metricValue="1.2149999999999999" metricValueMax="1.26" metricValueMin="1.17" pageId="1" pageNumber="86" unit="mm" value="1.2149999999999999" valueMax="1.26" valueMin="1.17">1.17 to 1.26 mm</quantity>
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(n = 3). Chaetotaxonomy according to
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<bibRefCitation id="1A354B1EA651E273BA37E87BFDB5FBFE" author="Williams, N. E. & Wiggins, G. B." box="[301,621,1073,1098]" pageId="1" pageNumber="86" pagination="421 - 429" refId="ref7928" refString="Williams, N. E. & Wiggins, G. B. (1981) A proposed setal nomenclature and homology for larval Trichoptera. In: Moretti, G. P. (Ed.), Proceedings of the Third International Symposium on Trichoptera. Series Entomologica. Vol. 20. Springer, Dordrecht, pp. 421 - 429. https: // doi. org / 10.1007 / 978 - 94 - 009 - 8641 - 1 _ 52" type="book chapter" year="1981">Williams & Wiggins (1981)</bibRefCitation>
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and
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<bibRefCitation id="1A354B1EA651E273B9B2E87BFC70FBFE" author="Friedrich, F. & Schulz, J. & Kubiak, M. & Beckmann, F. & Wilde, F." box="[680,936,1073,1098]" pageId="1" pageNumber="86" pagination="1505 - 1524" refId="ref5404" refString="Friedrich, F., Schulz, J., Kubiak, M., Beckmann, F. & Wilde, F. (2015) The larval head anatomy of Rhyacophila (Rhyacophilidae) with discussion on mouthpart homology and the groundplan of Trichoptera. Journal of Morphology, 276, 1505 - 1524. https: // doi. org / 10.1002 / jmor. 20475" type="journal article" year="2015">
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Friedrich
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<emphasis id="4CD0EAFDA651E273B80DE878FC93FBFD" box="[791,843,1073,1097]" italics="true" pageId="1" pageNumber="86">et al</emphasis>
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. (2015)
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</bibRefCitation>
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, anal proleg terminology following
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<bibRefCitation id="1A354B1EA651E273BE5CE87BFF3CFBDA" author="Nielsen, A." pageId="1" pageNumber="86" pagination="255 - 631" refId="ref7218" refString="Nielsen, A. (1942) Uber die Entwicklung und Biologie der Trichopteren mit besonderer Berucksichtigung der Quelltrichopteren Himmerlands. Archiv fur Hydrobiologie, 17 (Supplement), 255 - 631." type="journal article" year="1942">Nielsen (1942)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="7E1B36EFA651E273BBDDE833FDE4F966" blockId="1.[151,1437,1037,2034]" pageId="1" pageNumber="86">
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<emphasis id="4CD0EAFDA651E273BBDDE833FED5FB26" bold="true" box="[199,269,1145,1170]" pageId="1" pageNumber="86">Head.</emphasis>
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Head capsule finely wrinkled, rectangular, with lateral margins almost parallel, slightly convergent anteriorly, posterior third evenly rounded (
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<figureCitation id="E69F2A6AA651E273B96AE8D7FD0CFB02" box="[624,724,1181,1206]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Figs 1–2</figureCitation>
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). Coloration pale yellowish, with conspicuous dark brown, oval muscle attachment spots at dorsal, ventral, and lateral sides of parietalia, at frontoclypeus, and along coronal suture (
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<figureCitation id="E69F2A6AA651E273BB85E8AFFEF8FB4A" box="[159,288,1253,1278]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Figs 1–2, 5</figureCitation>
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). In addition, with median brown pigmented areas (1) at posterior third of frontoclypeus, including 3– 4 light muscle attachment spots; (2) on parietalia, dorsally from posterior frontoclypeal suture beside posterior coronal suture to foramen occipitale (
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<figureCitation id="E69F2A6AA651E273B951E967FD57FAF2" box="[587,655,1325,1350]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
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); (3) laterally, at eye level, from seta #12 to apophysis of foramen occipitale (
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<figureCitation id="E69F2A6AA651E273BA09E91BFE8EFADE" box="[275,342,1361,1386]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 5</figureCitation>
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), and (4) ventrally, from foramen occipitale to center of parietalia (
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<figureCitation id="E69F2A6AA651E273BF5DE91BFB54FADE" box="[1095,1164,1361,1386]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 2</figureCitation>
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). With brownish-yellow anterior margin of frontoclypeus and pale brown patches immediately posterior of frontoclypeal margin and around setae #5 (
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<figureCitation id="E69F2A6AA651E273BA1BE9D3FE9BFA06" box="[257,323,1433,1458]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
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). On each parietal, 13 dorsal and 5 ventral primary setae present, with setae #9, 12, 14 and 15 long and conspicuous (
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<figureCitation id="E69F2A6AA651E273BA45E9F7FE47FA62" box="[351,415,1469,1494]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
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). In addition, 17 pairs and a single sensory pit (sensilla campaniformia) present, 3 pairs and the single pit situated on frontocylpeus. Frontoclypeus bell-shaped, with deep central constriction and 5 pairs of primary setae, 2 of them near anterior border (#2–3) (
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<figureCitation id="E69F2A6AA651E273B9E7EA4FFC98F9AA" box="[765,832,1541,1566]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
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). Antennae each midway between anterior border of eye and lateral base of mandible, antennal base conical, antenna short, with short flagellum. Labrum pale yellow, with 6 pairs of primary setae (only 3 pairs visible in
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<figureCitation id="E69F2A6AA651E273B9E2EA07FCE3F9D2" box="[760,827,1613,1638]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
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). Ventral apotome triangular, with thickened anterior border (
|
||
<figureCitation id="E69F2A6AA651E273BBEBEA3BFEEFF93E" box="[241,311,1649,1674]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 2</figureCitation>
|
||
). Mandibles medium brown, almost black apically, and asymmetrical (
|
||
<figureCitation id="E69F2A6AA651E273BF45EA3BFB7DF93E" box="[1119,1189,1649,1674]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 3</figureCitation>
|
||
): right mandible with short, stout basal tooth, lacking on left mandible and replaced there by straight cutting edge. Each mandible base with 2 setae laterally (
|
||
<figureCitation id="E69F2A6AA651E273BA95EAF3FE0BF966" box="[399,467,1721,1746]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 3</figureCitation>
|
||
, arrows).
|
||
</paragraph>
|
||
<paragraph id="7E1B36EFA651E270BBDDEA97FCA4FE64" blockId="1.[151,1437,1037,2034]" lastBlockId="2.[151,1437,151,789]" lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86">
|
||
<emphasis id="4CD0EAFDA651E273BBDDEA97FEFEF942" bold="true" box="[199,294,1757,1782]" pageId="1" pageNumber="86">Thorax.</emphasis>
|
||
Pronotum fully covered by 2 large sclerites, tapering posteriorly. Pronotal sclerites yellowish, each with reddish brown muscle attachment spots creating arc–like pattern marked also by light brown pigmentation pointing toward anterolateral corner. In addition, circles of muscle attachment spots situated on posterolateral halves of sclerites, each with seta #5 on anterior border of circle. Dark round spot anterolateral of each seta #5; further muscles attachment spots and narrow stripes of pigment bordering posterior half of median suture (
|
||
<figureCitation id="E69F2A6AA651E273BE26EB27FF7FF81E" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Figs. 4a, 5</figureCitation>
|
||
). Posterior pronotal corners prolonged ventrally (
|
||
<figureCitation id="E69F2A6AA651E273B9F6EBDBFCECF81E" box="[748,820,1937,1962]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="1" pageNumber="86">Fig. 5</figureCitation>
|
||
). Posterior and lateral margins thickened and darkly striped, including anterolateral corners; posterior dark border interrupted by pale patch dorsal of posterior bulge; lateral borders straight to slightly concave. Pronotal notch at anterolateral corner large, deep and semicircular, with 2 long setae (
|
||
<figureCitation id="E69F2A6AA652E270BA2CECDDFE52FF04" box="[310,394,151,176]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="2" pageNumber="87">Fig. 4a</figureCitation>
|
||
, #22, #23) and 3 tiny (
|
||
<figureCitation id="E69F2A6AA652E270B980ECDDFD36FF04" box="[666,750,151,176]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="2" pageNumber="87">Fig. 4a</figureCitation>
|
||
, dotted oval). Prosternal horn lacking, prosternite narrowly quadrangular, with four medium-brown patches (
|
||
<figureCitation id="E69F2A6AA652E270B9D4ECF6FCFDFF61" box="[718,805,188,213]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="2" pageNumber="87">Fig. 4b</figureCitation>
|
||
). Meso- and metathoraces totally unsclerotized, white, with pale purplish-blue coloured areas subdivided by whitish bands; each with single pair of anterior
|
||
<emphasis id="4CD0EAFDA652E270BE1CECABFAC7FF4C" box="[1286,1311,225,248]" italics="true" pageId="2" pageNumber="87">sa</emphasis>
|
||
1 setae and groups of one long and two tiny posterior
|
||
<emphasis id="4CD0EAFDA652E270B96CED4FFD57FEA8" box="[630,655,261,284]" italics="true" pageId="2" pageNumber="87">sa</emphasis>
|
||
2 setae. Foretrochantins large, each with dark central stripe and large rounded anterior process bearing terminal seta #10 (
|
||
<figureCitation id="E69F2A6AA652E270B9EAED6DFCBFFEF4" box="[752,871,295,320]" captionStart="FIGURES 1–5" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[217,1378,195,1903]" captionTargetId="figure@6.[214,1380,193,1905]" captionTargetPageId="6" captionText="FIGURES 1–5. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 1, head, dorsal (lb = labrum; small numbers refer to setal positions); 2, head, ventral (small numbers refer to setal positions); 3, mandibles, ventral (maxillolabium removed; arrows= lateral setae); 4a, prothorax, dorsal (small numbers refer to setal positions; dotted oval= tiny setae of pronotal notch); 4b, prosternum (p= prosternite); 5, head and prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713922" httpUri="https://zenodo.org/record/3713922/files/figure.png" pageId="2" pageNumber="87">Figs 4a, 5</figureCitation>
|
||
,
|
||
<figureCitation id="E69F2A6AA652E270B86FED6DFC5DFEF4" box="[885,901,295,320]" captionStart="FIGURES 6–10" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[153,1407,195,1853]" captionTargetId="figure@7.[151,1435,193,1855]" captionTargetPageId="7" captionText="FIGURES 6–10. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 6, left foreleg, anterior (small numbers refer to setal positions); 7, left mesothoracic gill and midleg, anterior; 8, left metathoracic gill and hind leg, anterior; 9, detail of right mesothoracic gill, right lateral; 10, gills of abdominal segment VII, anterolateral (small numbers refer to setal positions). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713924" httpUri="https://zenodo.org/record/3713924/files/figure.png" pageId="2" pageNumber="87">6</figureCitation>
|
||
). Pleura of meso- and metathoraces each with upright, single-filament gill; episterna and epimera elongate, black (
|
||
<figureCitation id="E69F2A6AA652E270B8BAED06FBD1FED0" box="[928,1033,332,357]" captionStart="FIGURES 6–10" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[153,1407,195,1853]" captionTargetId="figure@7.[151,1435,193,1855]" captionTargetPageId="7" captionText="FIGURES 6–10. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 6, left foreleg, anterior (small numbers refer to setal positions); 7, left mesothoracic gill and midleg, anterior; 8, left metathoracic gill and hind leg, anterior; 9, detail of right mesothoracic gill, right lateral; 10, gills of abdominal segment VII, anterolateral (small numbers refer to setal positions). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713924" httpUri="https://zenodo.org/record/3713924/files/figure.png" pageId="2" pageNumber="87">Figs 7–9</figureCitation>
|
||
). Legs pale brown, with borders of sclerites darker and with numerous setae on coxae, trochanters, and femora; tibiae and tarsi with only small number of setae (
|
||
<figureCitation id="E69F2A6AA652E270BBE6EDDEFEBAFE19" box="[252,354,404,429]" captionStart="FIGURES 6–10" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[153,1407,195,1853]" captionTargetId="figure@7.[151,1435,193,1855]" captionTargetPageId="7" captionText="FIGURES 6–10. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 6, left foreleg, anterior (small numbers refer to setal positions); 7, left mesothoracic gill and midleg, anterior; 8, left metathoracic gill and hind leg, anterior; 9, detail of right mesothoracic gill, right lateral; 10, gills of abdominal segment VII, anterolateral (small numbers refer to setal positions). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713924" httpUri="https://zenodo.org/record/3713924/files/figure.png" pageId="2" pageNumber="87">Figs 6–8</figureCitation>
|
||
). Forefemora distinctly wider than mid- and hind femora. Tarsal claws long and slender, curved, pointed, with basal spur originating from conical base (
|
||
<figureCitation id="E69F2A6AA652E270B818EDFDFCB6FE64" box="[770,878,439,464]" captionStart="FIGURES 6–10" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[153,1407,195,1853]" captionTargetId="figure@7.[151,1435,193,1855]" captionTargetPageId="7" captionText="FIGURES 6–10. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 6, left foreleg, anterior (small numbers refer to setal positions); 7, left mesothoracic gill and midleg, anterior; 8, left metathoracic gill and hind leg, anterior; 9, detail of right mesothoracic gill, right lateral; 10, gills of abdominal segment VII, anterolateral (small numbers refer to setal positions). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713924" httpUri="https://zenodo.org/record/3713924/files/figure.png" pageId="2" pageNumber="87">Figs. 6–8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E1B36EFA652E270BBDDED96FBDAFCA0" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="87">
|
||
<emphasis id="4CD0EAFDA652E270BBDDED96FE9BFE41" bold="true" box="[199,323,476,501]" pageId="2" pageNumber="87">Abdomen.</emphasis>
|
||
Abdomen slightly depressed (flattened dorsoventrally); dorsally pale purplish–blue, with coloured areas subdivided by pale bands. Laterally and ventrally pale cream coloured. Each segment with 2 pairs of long dorsal primary setae (
|
||
<figureCitation id="E69F2A6AA652E270BA93EE6EFE03FD88" box="[393,475,548,573]" captionStart="FIGURES 6–10" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[153,1407,195,1853]" captionTargetId="figure@7.[151,1435,193,1855]" captionTargetPageId="7" captionText="FIGURES 6–10. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 6, left foreleg, anterior (small numbers refer to setal positions); 7, left mesothoracic gill and midleg, anterior; 8, left metathoracic gill and hind leg, anterior; 9, detail of right mesothoracic gill, right lateral; 10, gills of abdominal segment VII, anterolateral (small numbers refer to setal positions). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713924" httpUri="https://zenodo.org/record/3713924/files/figure.png" pageId="2" pageNumber="87">Fig. 10</figureCitation>
|
||
, setae #2, 5), a pair of lateral gills consisting of four filaments, originating from short, truncate base (
|
||
<figureCitation id="E69F2A6AA652E270BA20EE0DFE55FDD4" box="[314,397,583,608]" captionStart="FIGURES 6–10" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[153,1407,195,1853]" captionTargetId="figure@7.[151,1435,193,1855]" captionTargetPageId="7" captionText="FIGURES 6–10. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 6, left foreleg, anterior (small numbers refer to setal positions); 7, left mesothoracic gill and midleg, anterior; 8, left metathoracic gill and hind leg, anterior; 9, detail of right mesothoracic gill, right lateral; 10, gills of abdominal segment VII, anterolateral (small numbers refer to setal positions). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713924" httpUri="https://zenodo.org/record/3713924/files/figure.png" pageId="2" pageNumber="87">Fig. 10</figureCitation>
|
||
), and pair of ventral setae (
|
||
<figureCitation id="E69F2A6AA652E270B9D8EE0DFCCCFDD4" box="[706,788,583,608]" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1874,1897]" captionTargetBox="[217,1377,195,1861]" captionTargetId="figure@8.[213,1379,193,1863]" captionTargetPageId="8" captionText="FIGURES 11–15. Figures 11–12. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 11, abdominal segments IX, X and anal proleg, right lateral (ac = anal claw; bd = distal ‘b’ sclerite; bp = proximal ‘b’ sclerite; bv = ventral ‘b’ sclerite; c = ‘c’ sclerite; h = proximal hook of sclerite ‘b’; small numbers refer to setal positions; arrow = teeth on anal claw); 12, abdominal segments IX, X and anal proleg, dorsal. Figures 13–15. Rhyacophila tsurakiana Malicky 1984, 5th instar larva. 13, head, dorsal; 14, head, ventral; 15, prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713926" httpUri="https://zenodo.org/record/3713926/files/figure.png" pageId="2" pageNumber="87">Fig. 11</figureCitation>
|
||
, #10). Lateral fringe lacking. Abdominal dorsum IX with large, yellow quadrangular sclerite with black anterior and posterior borders (
|
||
<figureCitation id="E69F2A6AA652E270BF3BEE26FBA1FD30" box="[1057,1145,620,645]" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1874,1897]" captionTargetBox="[217,1377,195,1861]" captionTargetId="figure@8.[213,1379,193,1863]" captionTargetPageId="8" captionText="FIGURES 11–15. Figures 11–12. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 11, abdominal segments IX, X and anal proleg, right lateral (ac = anal claw; bd = distal ‘b’ sclerite; bp = proximal ‘b’ sclerite; bv = ventral ‘b’ sclerite; c = ‘c’ sclerite; h = proximal hook of sclerite ‘b’; small numbers refer to setal positions; arrow = teeth on anal claw); 12, abdominal segments IX, X and anal proleg, dorsal. Figures 13–15. Rhyacophila tsurakiana Malicky 1984, 5th instar larva. 13, head, dorsal; 14, head, ventral; 15, prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713926" httpUri="https://zenodo.org/record/3713926/files/figure.png" pageId="2" pageNumber="87">Fig. 12</figureCitation>
|
||
); anterior border slightly narrower than posterior border, with outermost setae inserted on lateral borders of this sclerite (
|
||
<figureCitation id="E69F2A6AA652E270BFECEEC5FA95FD1C" box="[1270,1357,655,680]" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1874,1897]" captionTargetBox="[217,1377,195,1861]" captionTargetId="figure@8.[213,1379,193,1863]" captionTargetPageId="8" captionText="FIGURES 11–15. Figures 11–12. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 11, abdominal segments IX, X and anal proleg, right lateral (ac = anal claw; bd = distal ‘b’ sclerite; bp = proximal ‘b’ sclerite; bv = ventral ‘b’ sclerite; c = ‘c’ sclerite; h = proximal hook of sclerite ‘b’; small numbers refer to setal positions; arrow = teeth on anal claw); 12, abdominal segments IX, X and anal proleg, dorsal. Figures 13–15. Rhyacophila tsurakiana Malicky 1984, 5th instar larva. 13, head, dorsal; 14, head, ventral; 15, prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713926" httpUri="https://zenodo.org/record/3713926/files/figure.png" pageId="2" pageNumber="87">Fig. 12</figureCitation>
|
||
). Anal prolegs each consisting of proximal sclerite ’b’ with long sword process and ventral basal hook (
|
||
<figureCitation id="E69F2A6AA652E270BFCFEEFEFAFDFD78" box="[1237,1317,692,717]" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1874,1897]" captionTargetBox="[217,1377,195,1861]" captionTargetId="figure@8.[213,1379,193,1863]" captionTargetPageId="8" captionText="FIGURES 11–15. Figures 11–12. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 11, abdominal segments IX, X and anal proleg, right lateral (ac = anal claw; bd = distal ‘b’ sclerite; bp = proximal ‘b’ sclerite; bv = ventral ‘b’ sclerite; c = ‘c’ sclerite; h = proximal hook of sclerite ‘b’; small numbers refer to setal positions; arrow = teeth on anal claw); 12, abdominal segments IX, X and anal proleg, dorsal. Figures 13–15. Rhyacophila tsurakiana Malicky 1984, 5th instar larva. 13, head, dorsal; 14, head, ventral; 15, prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713926" httpUri="https://zenodo.org/record/3713926/files/figure.png" pageId="2" pageNumber="87">Fig. 11</figureCitation>
|
||
bp, h, sp), pointed distal sclerite ‘b’, ventral ‘c’ sclerite (
|
||
<figureCitation id="E69F2A6AA652E270B9B9EE9DFD2BFD44" box="[675,755,727,752]" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1874,1897]" captionTargetBox="[217,1377,195,1861]" captionTargetId="figure@8.[213,1379,193,1863]" captionTargetPageId="8" captionText="FIGURES 11–15. Figures 11–12. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 11, abdominal segments IX, X and anal proleg, right lateral (ac = anal claw; bd = distal ‘b’ sclerite; bp = proximal ‘b’ sclerite; bv = ventral ‘b’ sclerite; c = ‘c’ sclerite; h = proximal hook of sclerite ‘b’; small numbers refer to setal positions; arrow = teeth on anal claw); 12, abdominal segments IX, X and anal proleg, dorsal. Figures 13–15. Rhyacophila tsurakiana Malicky 1984, 5th instar larva. 13, head, dorsal; 14, head, ventral; 15, prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713926" httpUri="https://zenodo.org/record/3713926/files/figure.png" pageId="2" pageNumber="87">Fig. 11</figureCitation>
|
||
bd, c), and long, curved anal claw (ac) partially divided by ventral membranous section fitted with two tiny ventral teeth (
|
||
<figureCitation id="E69F2A6AA652E270B848EEB6FC7CFCA0" box="[850,932,764,789]" captionStart="FIGURES 11–15" captionStartId="8.[151,264,1874,1897]" captionTargetBox="[217,1377,195,1861]" captionTargetId="figure@8.[213,1379,193,1863]" captionTargetPageId="8" captionText="FIGURES 11–15. Figures 11–12. Rhyacophila gudrunae Malicky 1972, 5th instar larva. 11, abdominal segments IX, X and anal proleg, right lateral (ac = anal claw; bd = distal ‘b’ sclerite; bp = proximal ‘b’ sclerite; bv = ventral ‘b’ sclerite; c = ‘c’ sclerite; h = proximal hook of sclerite ‘b’; small numbers refer to setal positions; arrow = teeth on anal claw); 12, abdominal segments IX, X and anal proleg, dorsal. Figures 13–15. Rhyacophila tsurakiana Malicky 1984, 5th instar larva. 13, head, dorsal; 14, head, ventral; 15, prothorax, right lateral. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3713926" httpUri="https://zenodo.org/record/3713926/files/figure.png" pageId="2" pageNumber="87">Fig. 11</figureCitation>
|
||
, arrow).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |