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<document id="3782F146B10DF6ABAFA41EA8E09E46F6" ID-DOI="10.21248/contrib.entomol.69.1.091-173" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742724" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638269330072" checkinUser="felipe" docAuthor="Assing, Volker &amp; Schülke, Michael" docDate="2019" docId="F078561B7215710FFC846792FAB9FE71" docLanguage="en" docName="ContribEntomol.69.1.91-173.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 69 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Anotylus hamatoides SCHULKE 2019, spec. nov." docType="treatment" docUuid="0167E052-EBAB-421A-BB72-39E33C1EDC97" docUuidSource="ZooBank" docVersion="3" lastPageNumber="159" masterDocId="0C412E637256714BFFD26338F915FFC6" masterDocTitle="The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera)" masterLastPageNumber="173" masterPageNumber="91" pageNumber="158" updateTime="1699437206044" updateUser="plazi">
<mods:mods id="03CF06CB8008D10EAC29AE6A48E2AFC9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7492C9F1DF606743ABC4006A7414E452">
<mods:title id="4DD63E5E0A8657CF08427C2256F8B40B">The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="99D29EE838B3E373DEA53BF930FCC667" type="personal">
<mods:role id="BD52F36611A0171DB8FA97F4F509E40C">
<mods:roleTerm id="9A6C46BF655CFF510B68170C2C859AEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1FD8EB50645611A444CBBCD8109A4A8C">Assing, Volker</mods:namePart>
</mods:name>
<mods:name id="695A24FE64ED244B1D83E6902A5044FB" type="personal">
<mods:role id="E40E6194DC139ED8C1598F9AA616F84E">
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<mods:namePart id="534752622B22CBFB0EDA757E7802B1DF">Schülke, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="054E6CBE65209AD5ABCD5D2FA939940D">text</mods:typeOfResource>
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<mods:titleInfo id="6F9192F331253DD1BA465D847E94A4A3">
<mods:title id="61A8D6D1B05947C50D5839AC7B4A9891">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="A2AF92E1A50EDDD49359FFA5D3562D87">
<mods:date id="B867F8167747EE4FFB0EE3F651E95284">2019</mods:date>
<mods:detail id="6B117D71F958201421E5128A881D8408" type="pubDate">
<mods:number id="CF0B2B5560D9483D0846EE401C751DE1">2019-06-24</mods:number>
</mods:detail>
<mods:detail id="5D12563C40659A55C327CD0DF1F83DAE" type="volume">
<mods:number id="A70170733ADA553BD2D077E39694B0D6">69</mods:number>
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<mods:number id="9588E47FF1431DCE4A62E752D0E7E147">1</mods:number>
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<mods:start id="7BD2CF2A9590F202D63F644E52D4DEFB">91</mods:start>
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</mods:extent>
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<mods:classification id="79C140131B4DEE067BC98B3281D39F03">journal article</mods:classification>
<mods:identifier id="BB804517F40F759701A5F9E54BA264A4" type="DOI">10.21248/contrib.entomol.69.1.091-173</mods:identifier>
<mods:identifier id="0C45D8E9A2F729C706F1C64FB73F2D57" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="F231CF09665DB99F7E6E04A82AE0A3B6" type="Zenodo-Dep">5742724</mods:identifier>
<mods:identifier id="C867A15E6C1238767594FA60ADADD9A2" type="ZooBank">9B8BCFC0-C491-45EF-9BC8-04A6B6B7DB8C</mods:identifier>
</mods:mods>
<treatment id="F078561B7215710FFC846792FAB9FE71" LSID="urn:lsid:zoobank.org:act:0167E052-EBAB-421A-BB72-39E33C1EDC97" httpUri="http://treatment.plazi.org/id/F078561B7215710FFC846792FAB9FE71" lastPageId="68" lastPageNumber="159" pageId="67" pageNumber="158">
<subSubSection id="30CBB48672157108FC846792FC75FB23" pageId="67" pageNumber="158" type="nomenclature">
<paragraph id="786EE70D72157108FC846792FDEBFB00" blockId="67.[854,1376,1189,1285]" box="[854,1278,1189,1222]" pageId="67" pageNumber="158">
<heading id="2326506172157108FC846792FDEBFB00" box="[854,1278,1189,1222]" fontSize="12" level="2" pageId="67" pageNumber="158" reason="2">
<taxonomicName id="BFD19C8E72157108FC846792FD9BFB03" ID-CoL="8XTRN" authority="SCHULKE" authorityName="SCHULKE" authorityYear="2019" box="[854,1166,1189,1222]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="158" phylum="Arthropoda" rank="species" species="hamatoides" status="sp. nov.">
<emphasis id="4AA53B1F72157108FC846792FD3BFB00" box="[854,1070,1194,1222]" italics="true" pageId="67" pageNumber="158">Anotylus hamatoides</emphasis>
SCHÜLKE
</taxonomicName>
<taxonomicNameLabel id="5196866472157108FB466792FDEBFB00" box="[1172,1278,1194,1222]" pageId="67" pageNumber="158" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="786EE70D72157108FC8467E9FC75FB23" blockId="67.[854,1376,1189,1285]" box="[854,1376,1233,1253]" pageId="67" pageNumber="158">
<uri id="0C40EB0F72157108FC8467E9FC75FB23" box="[854,1376,1233,1253]" pageId="67" pageNumber="158">
urn:lsid:zoobank.org:act:
<uuid id="0C77DDD872157108FBD767E9FC75FB23" box="[1029,1376,1233,1253]" pageId="67" pageNumber="158">0167E052-EBAB-421A-BB72-39E33C1EDC97</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="30CBB48672157108FC8467C9FAA6FAC3" box="[854,947,1265,1285]" pageId="67" pageNumber="158" type="description">
<paragraph id="786EE70D72157108FC8467C9FAA6FAC3" blockId="67.[854,1376,1189,1285]" box="[854,947,1265,1285]" pageId="67" pageNumber="158">
(
<figureCitation id="E0EAFB8872157108FC8867C9FABAFAC3" box="[858,943,1265,1285]" captionStart="Figs 41–56" captionStartId="59.[174,214,2000,2022]" captionTargetBox="[174,1421,255,1984]" captionTargetId="figure-27@59.[173,1421,254,1994]" captionTargetPageId="59" captionText="Figs 41–56: Tachyusa unguis (41–46), Anotylus hamatoides (47–52), Platystethus cephalotes (53–54), and P. laevis (55–56): forebody (41); abdomen (42); (median lobe of) aedeagus in lateral and in ventral view (43–44, 51–52); apex of ventral process of aedeagus in ventral view (45); spermatheca (46); male sternite VII (47); posterior process of male sternite VII (48); male sternite VIII (49, 53, 55); posterior process of male sternite VIII (50); male tergite X (54, 56). Scale bars: 41–42: 0.5 mm; 43–56: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742748" httpUri="https://zenodo.org/record/5742748/files/figure.png" pageId="67" pageNumber="158">Figs 47–52</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="30CBB4867215710FFCE26618FB13FE3E" lastPageId="68" lastPageNumber="159" pageId="67" pageNumber="158" type="materials_examined">
<paragraph id="786EE70D7215710FFCE26618FB13FE3E" blockId="67.[816,1428,1311,2104]" lastBlockId="68.[212,823,256,504]" lastPageId="68" lastPageNumber="159" pageId="67" pageNumber="158">
<emphasis id="4AA53B1F72157108FCE26618FAC5FAF1" bold="true" box="[816,976,1311,1336]" pageId="67" pageNumber="158">Type material</emphasis>
:
<typeStatus id="A76A59AF72157108FC346618FD5AFAFE" box="[998,1103,1312,1336]" pageId="67" pageNumber="158" type="holotype">Holotype</typeStatus>
: “
<collectingCountry id="00C6A79D72157108FB596618FC16FAFE" box="[1163,1283,1312,1336]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
,
<collectingRegion id="BA1529EF72157108FAC46618FC9AFAF1" box="[1302,1423,1312,1336]" country="Armenia" name="Lorri" pageId="67" pageNumber="158">Lori prov.</collectingRegion>
, Teghut mine, near artef. pond 41.0907° 044.8117° (soil traps)
<quantity id="BF294AE872157108FCAB6658FAABFAB1" box="[889,958,1376,1400]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="67" pageNumber="158" unit="m" value="990.0">990 m</quantity>
, 23.07.–
<date id="0C6FC1CD72157108FBC46658FD99FABE" box="[1046,1164,1376,1400]" pageId="67" pageNumber="158" value="2015-08-23">23.08.2015</date>
Karagyan &amp; Kalashian leg. / +6 / Holotypus-
<taxonomicName id="BFD19C8E72157108FBED66B8FC08FA5E" authority="Assing &amp; Schülke, 2019" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[1087,1309,1408,1432]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="158" phylum="Arthropoda" rank="species" species="hamatoides" status="sp. nov.">
<emphasis id="4AA53B1F72157108FBED66B8FC08FA5E" box="[1087,1309,1408,1432]" italics="true" pageId="67" pageNumber="158">Anotylus hamatoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5196866472157108FAF466B8FC81FA51" box="[1318,1428,1408,1431]" pageId="67" pageNumber="158" rank="species">spec. nov.</taxonomicNameLabel>
det. M. Schülke 2018” (
<collectionCode id="1EC07FC872157108FBFB6698FD7EFA7E" box="[1065,1131,1440,1464]" pageId="67" pageNumber="158">IZAY</collectionCode>
).
<typeStatus id="A76A59AF72157108FBAF6698FDF2FA7E" box="[1149,1255,1440,1464]" pageId="67" pageNumber="158" type="paratype">Paratypes</typeStatus>
: 1: same data as
<typeStatus id="A76A59AF72157108FC8066F8FAA5FA1E" box="[850,944,1472,1496]" pageId="67" pageNumber="158" type="holotype">holotype</typeStatus>
but “+14” (
<collectionCode id="1EC07FC872157108FBEC66F8FD6AFA1E" box="[1086,1151,1472,1496]" pageId="67" pageNumber="158">IZAY</collectionCode>
); 1: “
<collectingCountry id="00C6A79D72157108FB0566F8FC5AFA1E" box="[1239,1359,1472,1496]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
,
<collectingRegion id="BA1529EF72157108FAB266F8FA7DFA31" country="Armenia" name="Lorri" pageId="67" pageNumber="158">Lori prov.</collectingRegion>
, Teghut mine, N41.0967°, E44.8218°
<quantity id="BF294AE872157108FB2366D8FC23FA31" box="[1265,1334,1504,1528]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="67" pageNumber="158" unit="m" value="920.0">920 m</quantity>
, 22.08.–
<date id="0C6FC1CD72157108FCE26538FAB0F9DE" box="[816,933,1536,1560]" pageId="67" pageNumber="158" value="2013-01-10">01.10.2013</date>
, Kalashian leg. / (soil traps Ser.2)” (cSch); 1: “
<collectingCountry id="00C6A79D72157108FCEB6518FAA4F9FE" box="[825,945,1568,1592]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
,
<collectingRegion id="BA1529EF72157108FC126518FD25F9F1" box="[960,1072,1568,1592]" country="Armenia" name="Lorri" pageId="67" pageNumber="158">Lori prov.</collectingRegion>
, Teghut mine, near artef. pond 41.0907° 044.8117° (soil traps)
<quantity id="BF294AE872157108FBAC6578FDD7F991" box="[1150,1218,1600,1624]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="67" pageNumber="158" unit="m" value="990.0">990 m</quantity>
, 23.08.–
<date id="0C6FC1CD72157108FAC56578FC98F99E" box="[1303,1421,1600,1624]" pageId="67" pageNumber="158" value="2015-09-23">23.09.2015</date>
Karagyan &amp; Kalashian leg. / +2” (cSch); 1: “
<collectingCountry id="00C6A79D72157108FAC36558FC9AF9BE" box="[1297,1423,1632,1656]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
,
<collectingRegion id="BA1529EF72157108FCE265B8FA8AF951" box="[816,927,1664,1688]" country="Armenia" name="Lorri" pageId="67" pageNumber="158">Lori prov.</collectingRegion>
, Teghut mine, N
<geoCoordinate id="1DE581CA72157108FB8565B8FDD8F95E" box="[1111,1229,1664,1688]" degrees="41.0743" direction="east" orientation="longitude" pageId="67" pageNumber="158" precision="5" value="41.0743">41.0743° E</geoCoordinate>
44.8399°,
<quantity id="BF294AE872157108FAEA65B8FC9CF951" box="[1336,1417,1664,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.105" pageId="67" pageNumber="158" unit="m" value="1105.0">1105 m</quantity>
, 22.08.–
<date id="0C6FC1CD72157108FCA96598FAE7F97E" box="[891,1010,1696,1720]" pageId="67" pageNumber="158" value="2013-01-10">01.10.2013</date>
, Kalashian leg. / (soil traps Ser.3) /
<taxonomicName id="BFD19C8E72157108FCE265F8FD44F91E" authority="Pand." authorityName="Pand." box="[816,1105,1728,1752]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="158" phylum="Arthropoda" rank="species" species="fairmairei">
<emphasis id="4AA53B1F72157108FCE265F8FD10F91E" box="[816,1029,1728,1752]" italics="true" pageId="67" pageNumber="158">Anotylus fairmairei</emphasis>
Pand.
</taxonomicName>
” (
<collectionCode id="1EC07FC872157108FBA365F8FDA7F91E" box="[1137,1202,1728,1752]" pageId="67" pageNumber="158">IZAY</collectionCode>
); 1: “
<collectingCountry id="00C6A79D72157108FAC365F8FC9AF91E" box="[1297,1423,1728,1752]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
,
<collectingRegion id="BA1529EF72157108FCE265D8FAB4F931" box="[816,929,1760,1784]" country="Armenia" name="Lorri" pageId="67" pageNumber="158">Lori prov.</collectingRegion>
, Teghut mine, N
<geoCoordinate id="1DE581CA72157108FB8D65D8FDCDF93E" box="[1119,1240,1760,1784]" degrees="41.0910" direction="east" orientation="longitude" pageId="67" pageNumber="158" precision="5" value="41.091">41.0910° E</geoCoordinate>
44.8514°,
<quantity id="BF294AE872157108FA9765D8FC9CF931" box="[1349,1417,1760,1784]" metricMagnitude="2" metricUnit="m" metricValue="8.87" pageId="67" pageNumber="158" unit="m" value="887.0">887 m</quantity>
, 22.08.–
<date id="0C6FC1CD72157108FCA96438FAE7F8DE" box="[891,1010,1792,1816]" pageId="67" pageNumber="158" value="2013-01-10">01.10.2013</date>
, Kalashian leg. / (soil traps Ser.4)”; 1: “
<collectingCountry id="00C6A79D72157108FCA36418FAFCF8FE" box="[881,1001,1824,1848]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
,
<collectingRegion id="BA1529EF72157108FC286418FD79F8F1" box="[1018,1132,1824,1848]" country="Armenia" name="Lorri" pageId="67" pageNumber="158">Lori prov.</collectingRegion>
, Teghut mine, N
<geoCoordinate id="1DE581CA72157108FAE36418FA55F89E" degrees="41.0967" direction="east" orientation="longitude" pageId="67" pageNumber="158" precision="5" value="41.0967">41.0967° E</geoCoordinate>
44.8218°,
<quantity id="BF294AE872157108FC676478FAEFF891" box="[949,1018,1856,1880]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="67" pageNumber="158" unit="m" value="920.0">920 m</quantity>
, 20.06.–
<date id="0C6FC1CD72157108FB8F6478FDC1F89E" box="[1117,1236,1856,1880]" pageId="67" pageNumber="158" value="2013-01-08">01.08.2013</date>
, Kalashian leg. / (soil traps Ser. 2)”; 1: “
<collectingCountry id="00C6A79D72157108FBBC6458FDF1F8BE" box="[1134,1252,1888,1912]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
– Teghut env. (Akhtala),
<geoCoordinate id="1DE581CA72157108FC6764B8FD0FF85E" box="[949,1050,1920,1944]" degrees="41.091" direction="north" orientation="latitude" pageId="67" pageNumber="158" precision="55" value="41.091">41.091°N</geoCoordinate>
,
<geoCoordinate id="1DE581CA72157108FBE464B8FD82F85E" box="[1078,1175,1920,1944]" degrees="44.813" direction="east" orientation="longitude" pageId="67" pageNumber="158" precision="55" value="44.813">44.813°E</geoCoordinate>
,
<quantity id="BF294AE872157108FB6064B8FC16F851" box="[1202,1283,1920,1944]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="67" pageNumber="158" unit="m" value="1020.0">1020 m</quantity>
,
<date id="0C6FC1CD72157108FACD64B8FC9CF85E" box="[1311,1417,1920,1944]" pageId="67" pageNumber="158" value="2016-05-20">20.V.2016</date>
, leg. A. &amp; J. Müller” (cAss); 1: “T6 BF
<collectingCountry id="00C6A79D72157108FB3C6498FC5AF87E" box="[1262,1359,1952,1976]" name="Armenia" pageId="67" pageNumber="158">Armenia</collectingCountry>
, near Teghut (Akhtala)
<quantity id="BF294AE872157108FC2264F8FD54F811" box="[1008,1089,1984,2008]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="67" pageNumber="158" unit="m" value="1000.0">1000 m</quantity>
a. s. l., N
<geoCoordinate id="1DE581CA72157108FB7D64F8FC33F811" box="[1199,1318,1984,2008]" degrees="41.09099" direction="east" orientation="longitude" pageId="67" pageNumber="158" precision="1" value="41.09099">41.09099 E</geoCoordinate>
44.81202
<date id="0C6FC1CD72157108FCE264D8FAA2F83E" box="[816,951,2016,2040]" pageId="67" pageNumber="158" value="2016-08-26">26.VIII.2016</date>
leg. J. Müller” (cSch); 1, 1: “
<collectingCountry id="00C6A79D72157108FAC564D8FC86F83E" box="[1303,1427,2016,2040]" name="Armenia" pageId="67" pageNumber="158">ARMENIA</collectingCountry>
–
<collectingRegion id="BA1529EF72157108FC966B38FABAF7D1" box="[836,943,2048,2072]" country="Armenia" name="Lorri" pageId="67" pageNumber="158">Lori prov.</collectingRegion>
, Teghut mine, N
<geoCoordinate id="1DE581CA72157108FB8C6B38FDD1F7DE" box="[1118,1220,2048,2072]" degrees="41.091" direction="north" orientation="latitude" pageId="67" pageNumber="158" precision="55" value="41.091">41.091°N</geoCoordinate>
, E044.812°,
<quantity id="BF294AE872157108FA976B38FC9CF7D1" box="[1349,1417,2048,2072]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="67" pageNumber="158" unit="m" value="990.0">990 m</quantity>
,
<date id="0C6FC1CD72157108FCE26B18FAEAF7FE" box="[816,1023,2080,2104]" pageId="67" pageNumber="158" value="2017-08-13" valueMax="2017-10-13" valueMin="2017-08-13">13.VIII.–13.X.2017</date>
, leg. Kalashian et al.” (cAss); 1: “
<collectingCountry id="00C6A79D7212710FFF0E6238F847FEDE" box="[220,338,256,280]" name="Armenia" pageId="68" pageNumber="159">ARMENIA</collectingCountry>
–
<collectingRegion id="BA1529EF7212710FFE506238F8EDFED1" box="[386,504,256,280]" country="Armenia" name="Lorri" pageId="68" pageNumber="159">Lori prov.</collectingRegion>
, Teghut mine, N
<geoCoordinate id="1DE581CA7212710FFD1A6238FA27FEDE" box="[712,818,256,280]" degrees="41.096" direction="north" orientation="latitude" pageId="68" pageNumber="159" precision="55" value="41.096">41.096°N</geoCoordinate>
, E044.822°,
<quantity id="BF294AE87212710FFE836218F880FEF1" box="[337,405,288,312]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="68" pageNumber="159" unit="m" value="910.0">910 m</quantity>
,
<date id="0C6FC1CD7212710FFE776218FB7DFEFE" box="[421,616,288,312]" pageId="68" pageNumber="159" value="2017-05-25" valueMax="2017-07-23" valueMin="2017-05-25">25.V.–23.VII.2017</date>
, leg. Kalashian et al.” (cAss); 2: “
<collectingCountry id="00C6A79D7212710FFE486278FB05FE9E" box="[410,528,320,344]" name="Armenia" pageId="68" pageNumber="159">ARMENIA</collectingCountry>
–
<collectingRegion id="BA1529EF7212710FFDFF6278FB8CFE91" box="[557,665,320,344]" country="Armenia" name="Lorri" pageId="68" pageNumber="159">Lori prov.</collectingRegion>
, Teghut mine, N
<geoCoordinate id="1DE581CA7212710FFF356258F858FEBE" box="[231,333,352,376]" degrees="41.091" direction="north" orientation="latitude" pageId="68" pageNumber="159" precision="55" value="41.091">41.091°N</geoCoordinate>
, E044.851°,
<quantity id="BF294AE87212710FFE346258FB3FFEB1" box="[486,554,352,376]" metricMagnitude="2" metricUnit="m" metricValue="8.9" pageId="68" pageNumber="159" unit="m" value="890.0">890 m</quantity>
,
<date id="0C6FC1CD7212710FFD926258FBE3FEBE" box="[576,758,352,376]" pageId="68" pageNumber="159" value="2017-04-28" valueMax="2017-05-25" valueMin="2017-04-28">28.IV.–25.V.2017</date>
, leg. Kalashian et al.” (cAss);
<specimenCount id="6ED72C847212710FFE0F62B8FB06FE51" box="[477,531,384,408]" pageId="68" pageNumber="159" type="generic">1 ex.</specimenCount>
: “Lenkoran Leder (Reitter / Coll. Reitter /
<taxonomicName id="BFD19C8E7212710FFE5A6298FBEBFE7E" authority="(Fairm. &amp; Lab.)" baseAuthorityName="FAIRMAIRE &amp; LABOULBENE" baseAuthorityYear="1856" box="[392,766,416,440]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4AA53B1F7212710FFE5A6298FB5CFE7E" box="[392,585,416,440]" italics="true" pageId="68" pageNumber="159">Anotylus hamatus</emphasis>
(Fairm. &amp; Lab.)
</taxonomicName>
det. Makranczy, 1997 / misidentification!
<taxonomicName id="BFD19C8E7212710FFDBA62F8FBD0FE1E" authority="Assing &amp; Schülke, 2019" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[616,709,448,472]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="4AA53B1F7212710FFDBA62F8FBD0FE1E" box="[616,709,448,472]" italics="true" pageId="68" pageNumber="159">Anotylus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="519686647212710FFD1962F8FA17FE11" box="[715,770,448,471]" pageId="68" pageNumber="159" rank="species">sp. n.</taxonomicNameLabel>
det. Makranczy, 1998” (
<collectionCode id="1EC07FC87212710FFE7362D8F8EEFE3E" box="[417,507,480,504]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="68" pageNumber="159">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="30CBB4867212710FFF066118FB2AFDB1" pageId="68" pageNumber="159" type="etymology">
<paragraph id="786EE70D7212710FFF066118FB2AFDB1" blockId="68.[212,823,544,632]" pageId="68" pageNumber="159">
<emphasis id="4AA53B1F7212710FFF066118F85EFDFE" bold="true" box="[212,331,544,568]" pageId="68" pageNumber="159">Etymology</emphasis>
: The specific epithet alludes to the strong resemblance to
<taxonomicName id="BFD19C8E7212710FFEAE6178F806FDBE" authority="(FAIRMAIRE &amp; LABOULBENE, 1856)" baseAuthorityName="FAIRMAIRE &amp; LABOULBENE" baseAuthorityYear="1856" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4AA53B1F7212710FFEAE6178F8E0FD9E" box="[380,501,576,600]" italics="true" pageId="68" pageNumber="159">A. hamatus</emphasis>
(FAIRMAIRE &amp; LABOULBÈNE, 1856)
</taxonomicName>
, most likely the sister taxon.
</paragraph>
</subSubSection>
<subSubSection id="30CBB4867212710FFF0661A7F8F9F7F1" pageId="68" pageNumber="159" type="description">
<paragraph id="786EE70D7212710FFF0661A7F843FCDE" blockId="68.[212,823,671,1688]" pageId="68" pageNumber="159">
<emphasis id="4AA53B1F7212710FFF0661A7F843FD71" bold="true" box="[212,342,671,695]" pageId="68" pageNumber="159">Description</emphasis>
: Body length
<quantity id="BF294AE87212710FFE296198FB68FD71" box="[507,637,672,696]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="68" pageNumber="159" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
; length of forebody 1.0–
<quantity id="BF294AE87212710FFE9661F8F882FD11" box="[324,407,704,728]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="68" pageNumber="159" unit="mm" value="1.2">1.2 mm</quantity>
. Coloration: whole body black; legs yellow; antennae black; mandibles and maxillary palpi dark brown.
</paragraph>
<paragraph id="786EE70D7212710FFF066018F8A2FB91" blockId="68.[212,823,671,1688]" pageId="68" pageNumber="159">
Measurements: males: head width
<quantity id="BF294AE87212710FFD866018FBFAFCF1" box="[596,751,800,824]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="68" pageNumber="159" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47–0.50 mm</quantity>
, head length
<quantity id="BF294AE87212710FFEF46078F8D4FC91" box="[294,449,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="68" pageNumber="159" unit="mm" value="0.325" valueMax="0.33" valueMin="0.32">0.32–0.33 mm</quantity>
, eye length
<quantity id="BF294AE87212710FFD846078FBA2FC91" box="[598,695,832,856]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="68" pageNumber="159" unit="mm" value="0.14">0.14 mm</quantity>
, length of temples
<quantity id="BF294AE87212710FFEFD6058F8DFFCB1" box="[303,458,864,888]" metricMagnitude="-4" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="68" pageNumber="159" unit="mm" value="0.135" valueMax="0.14" valueMin="0.13">0.13–0.14 mm</quantity>
, pronotum width
<quantity id="BF294AE87212710FFD406058FA38FCB1" box="[658,813,864,888]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.1" metricValueMin="4.9" pageId="68" pageNumber="159" unit="mm" value="0.5" valueMax="0.51" valueMin="0.49">0.49–0.51 mm</quantity>
, pronotum length
<quantity id="BF294AE87212710FFE7560B8FB12FC51" box="[423,519,896,920]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="68" pageNumber="159" unit="mm" value="0.35">0.35 mm</quantity>
, sutural length of elytra
<quantity id="BF294AE87212710FFF066098F87AFC71" box="[212,367,928,952]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.5999999999999996" pageId="68" pageNumber="159" unit="mm" value="0.37" valueMax="0.38" valueMin="0.36">0.36–0.38 mm</quantity>
, elytral width
<quantity id="BF294AE87212710FFDFB6098FBD1FC71" box="[553,708,928,952]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="68" pageNumber="159" unit="mm" value="0.625" valueMax="0.64" valueMin="0.61">0.61–0.64 mm</quantity>
. females: head width
<quantity id="BF294AE87212710FFE8960F8F8E3FC11" box="[347,502,960,984]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.8" metricValueMin="3.7" pageId="68" pageNumber="159" unit="mm" value="0.375" valueMax="0.38" valueMin="0.37">0.37–0.38 mm</quantity>
, head length
<quantity id="BF294AE87212710FFD4060F8FA38FC11" box="[658,813,960,984]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="68" pageNumber="159" unit="mm" value="0.245" valueMax="0.25" valueMin="0.24">0.24–0.25 mm</quantity>
, eye length
<quantity id="BF294AE87212710FFE8060D8F8F8FC31" box="[338,493,992,1016]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="68" pageNumber="159" unit="mm" value="0.11499999999999999" valueMax="0.12" valueMin="0.11">0.11–0.12 mm</quantity>
, length of temples
<quantity id="BF294AE87212710FFD1E60D8FA38FC31" box="[716,813,992,1016]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="68" pageNumber="159" unit="mm" value="0.08">0.08 mm</quantity>
, pronotum width
<quantity id="BF294AE87212710FFE5D6738FB3CFBD1" box="[399,553,1024,1048]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="68" pageNumber="159" unit="mm" value="0.41" valueMax="0.42" valueMin="0.4">0.40–0.42 mm</quantity>
, pronotum length 0,28–
<quantity id="BF294AE87212710FFF066718F821FBF1" box="[212,308,1056,1080]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="68" pageNumber="159" unit="mm" value="0.29">0.29 mm</quantity>
, sutural length of elytra
<quantity id="BF294AE87212710FFD946718FBF5FBF1" box="[582,736,1056,1080]" metricMagnitude="-4" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.5" metricValueMin="3.2" pageId="68" pageNumber="159" unit="mm" value="0.33499999999999996" valueMax="0.35" valueMin="0.32">0.32–0.35 mm</quantity>
, elytral width
<quantity id="BF294AE87212710FFECA6778F8A7FB91" box="[280,434,1088,1112]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.3" metricValueMin="5.1" pageId="68" pageNumber="159" unit="mm" value="0.52" valueMax="0.53" valueMin="0.51">0.51–0.53 mm</quantity>
.
</paragraph>
<paragraph id="786EE70D7212710FFF066758FA35FB5E" blockId="68.[212,823,671,1688]" pageId="68" pageNumber="159">
In size, body shape, punctuation, and microsculpture indistinguishable from
<taxonomicName id="BFD19C8E7212710FFE1967B8FB51FB5E" baseAuthorityName="FAIRMAIRE &amp; LABOULBENE" baseAuthorityYear="1856" box="[459,580,1152,1176]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4AA53B1F7212710FFE1967B8FB51FB5E" box="[459,580,1152,1176]" italics="true" pageId="68" pageNumber="159">A. hamatus</emphasis>
</taxonomicName>
and similar species.
</paragraph>
<paragraph id="786EE70D7212710FFF3A6798FB19F9F1" blockId="68.[212,823,671,1688]" pageId="68" pageNumber="159">
: head distinctly larger, transverse (width/length 1.45– 1.57), eyes as long as, or slightly longer than temples (eye length/temple length 1.0–1.1); sternite VII (
<figureCitation id="E0EAFB887212710FFD6067D8FA3CFB3E" box="[690,809,1248,1272]" captionStart="Figs 41–56" captionStartId="59.[174,214,2000,2022]" captionTargetBox="[174,1421,255,1984]" captionTargetId="figure-27@59.[173,1421,254,1994]" captionTargetPageId="59" captionText="Figs 41–56: Tachyusa unguis (41–46), Anotylus hamatoides (47–52), Platystethus cephalotes (53–54), and P. laevis (55–56): forebody (41); abdomen (42); (median lobe of) aedeagus in lateral and in ventral view (43–44, 51–52); apex of ventral process of aedeagus in ventral view (45); spermatheca (46); male sternite VII (47); posterior process of male sternite VII (48); male sternite VIII (49, 53, 55); posterior process of male sternite VIII (50); male tergite X (54, 56). Scale bars: 41–42: 0.5 mm; 43–56: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742748" httpUri="https://zenodo.org/record/5742748/files/figure.png" pageId="68" pageNumber="159">Figs 47–48</figureCitation>
) similar to that of
<taxonomicName id="BFD19C8E7212710FFE436638FB1FFADE" baseAuthorityName="FAIRMAIRE &amp; LABOULBENE" baseAuthorityYear="1856" box="[401,522,1280,1304]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4AA53B1F7212710FFE436638FB1FFADE" box="[401,522,1280,1304]" italics="true" pageId="68" pageNumber="159">A. hamatus</emphasis>
</taxonomicName>
, with prominent spine and a small tubercle with two larger spines in front of it; sternite VIII highly modified (
<figureCitation id="E0EAFB887212710FFDE26678FBBDFA9E" box="[560,680,1344,1368]" captionStart="Figs 41–56" captionStartId="59.[174,214,2000,2022]" captionTargetBox="[174,1421,255,1984]" captionTargetId="figure-27@59.[173,1421,254,1994]" captionTargetPageId="59" captionText="Figs 41–56: Tachyusa unguis (41–46), Anotylus hamatoides (47–52), Platystethus cephalotes (53–54), and P. laevis (55–56): forebody (41); abdomen (42); (median lobe of) aedeagus in lateral and in ventral view (43–44, 51–52); apex of ventral process of aedeagus in ventral view (45); spermatheca (46); male sternite VII (47); posterior process of male sternite VII (48); male sternite VIII (49, 53, 55); posterior process of male sternite VIII (50); male tergite X (54, 56). Scale bars: 41–42: 0.5 mm; 43–56: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742748" httpUri="https://zenodo.org/record/5742748/files/figure.png" pageId="68" pageNumber="159">Figs 49–50</figureCitation>
), in general respect similar to that of
<taxonomicName id="BFD19C8E7212710FFE2B6658FB67FABE" baseAuthorityName="FAIRMAIRE &amp; LABOULBENE" baseAuthorityYear="1856" box="[505,626,1376,1400]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4AA53B1F7212710FFE2B6658FB67FABE" box="[505,626,1376,1400]" italics="true" pageId="68" pageNumber="159">A. hamatus</emphasis>
</taxonomicName>
, with broad and deep emargination at base, broad emargination at apical border, interrupted in the middle by a short two-lobed process with 4–5 setae at apex of each lobe; aedeagus (
<figureCitation id="E0EAFB887212710FFF0966D8F846FA3E" box="[219,339,1504,1528]" captionStart="Figs 41–56" captionStartId="59.[174,214,2000,2022]" captionTargetBox="[174,1421,255,1984]" captionTargetId="figure-27@59.[173,1421,254,1994]" captionTargetPageId="59" captionText="Figs 41–56: Tachyusa unguis (41–46), Anotylus hamatoides (47–52), Platystethus cephalotes (53–54), and P. laevis (55–56): forebody (41); abdomen (42); (median lobe of) aedeagus in lateral and in ventral view (43–44, 51–52); apex of ventral process of aedeagus in ventral view (45); spermatheca (46); male sternite VII (47); posterior process of male sternite VII (48); male sternite VIII (49, 53, 55); posterior process of male sternite VIII (50); male tergite X (54, 56). Scale bars: 41–42: 0.5 mm; 43–56: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742748" httpUri="https://zenodo.org/record/5742748/files/figure.png" pageId="68" pageNumber="159">Figs 51–52</figureCitation>
),
<quantity id="BF294AE87212710FFEBC66D8F8D8FA31" box="[366,461,1504,1528]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="68" pageNumber="159" unit="mm" value="0.47">0.47 mm</quantity>
long, nearly symmetric, elongate, with the apical flag-like part of the paramere more extended than in
<taxonomicName id="BFD19C8E7212710FFE5F6518FB13F9FE" baseAuthorityName="FAIRMAIRE &amp; LABOULBENE" baseAuthorityYear="1856" box="[397,518,1568,1592]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4AA53B1F7212710FFE5F6518FB13F9FE" box="[397,518,1568,1592]" italics="true" pageId="68" pageNumber="159">A. hamatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="786EE70D7212710FFF3A6578F8F6F95E" blockId="68.[212,823,671,1688]" pageId="68" pageNumber="159">: head distinctly smaller, transverse (width/length 1.47– 1.59), eyes distinctly longer than temples (eye length/ temple length 1.45–1.50).</paragraph>
<paragraph id="786EE70D7212710FFF066587F8F9F7F1" blockId="68.[212,823,1727,2104]" pageId="68" pageNumber="159">
<emphasis id="4AA53B1F7212710FFF066587F8B7F91E" bold="true" box="[212,418,1727,1752]" pageId="68" pageNumber="159">Comparative notes</emphasis>
:
<taxonomicName id="BFD19C8E7212710FFE7F65F8FB93F91E" authorityName="Assing &amp; Schülke" authorityYear="2019" box="[429,646,1728,1752]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatoides">
<emphasis id="4AA53B1F7212710FFE7F65F8FB93F91E" box="[429,646,1728,1752]" italics="true" pageId="68" pageNumber="159">Anotylus hamatoides</emphasis>
</taxonomicName>
is distinguished from all other representatives of the
<taxonomicName id="BFD19C8E7212710FFD8065D8F9E6F8D1" baseAuthorityName="BLOCK" baseAuthorityYear="1799" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="tetracarinatus">
<emphasis id="4AA53B1F7212710FFD8065D8F9E6F8D1" italics="true" pageId="68" pageNumber="159">Anotylus tetracarinatus</emphasis>
</taxonomicName>
group with the outer edge of the front tibiae apically strongly indented only by the male primary and secondary sexual characters. The pronounced similarity to the widespread
<taxonomicName id="BFD19C8E7212710FFE816458F8D9F8BE" baseAuthorityName="FAIRMAIRE &amp; LABOULBENE" baseAuthorityYear="1856" box="[339,460,1888,1912]" class="Insecta" family="Staphylinidae" genus="Anotylus" kingdom="Animalia" order="Coleoptera" pageId="68" pageNumber="159" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4AA53B1F7212710FFE816458F8D9F8BE" box="[339,460,1888,1912]" italics="true" pageId="68" pageNumber="159">A. hamatus</emphasis>
</taxonomicName>
in the shape of the male sternites VII and VIII and in the shape of the aedeagus suggests a close relationship; most likely they represent sister taxa. Both species are distinguished especially by the shape of the apical margin of the male sternite VIII. Differences in the shape of sternite VII and the aedeagus are less pronounced, but constant.
</paragraph>
</subSubSection>
<subSubSection id="30CBB4867212710FFC8463C7FAB9FE71" pageId="68" pageNumber="159" type="distribution">
<paragraph id="786EE70D7212710FFC8463C7FAB9FE71" blockId="68.[854,1465,255,440]" pageId="68" pageNumber="159">
<emphasis id="4AA53B1F7212710FFC8463C7FDA1FED1" bold="true" box="[854,1204,255,280]" pageId="68" pageNumber="159">Distribution and natural history</emphasis>
: The species is currently known from the environs of Teghut in
<collectingRegion id="BA1529EF7212710FFAC56218FCA0FEFE" box="[1303,1461,288,312]" country="Armenia" name="Lorri" pageId="68" pageNumber="159">Lori province</collectingRegion>
, North
<collectingCountry id="00C6A79D7212710FFC4D6278FD15FE9E" box="[927,1024,320,344]" name="Armenia" pageId="68" pageNumber="159">Armenia</collectingCountry>
, and from the
<collectingRegion id="BA1529EF7212710FFB726278FC46FE9E" box="[1184,1363,320,344]" country="Azerbaijan" name="Lankaran" pageId="68" pageNumber="159">Lenkoran region</collectingRegion>
in
<collectingCountry id="00C6A79D7212710FFAA56278FA8FFEBE" name="Azerbaijan" pageId="68" pageNumber="159">Azerbaijan</collectingCountry>
. The Armenian specimens were collected with pitfall traps at elevations between approximately 890 and
<quantity id="BF294AE87212710FFC846298FAB2FE71" box="[854,935,416,440]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="68" pageNumber="159" unit="m" value="1100.0">1100 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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