205 lines
25 KiB
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205 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="E64A04E36A995430AE3BE8BEA130A88A" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole lavasoa Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="603A3805-0A35-4BAA-9AAA-8BB0882809D3" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>949</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="165090157" LSID="urn:lsid:zoobank.org:act:603A3805-0A35-4BAA-9AAA-8BB0882809D3" httpUri="http://treatment.plazi.org/id/E64A04E36A995430AE3BE8BEA130A88A" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="E64A04E3-6A99-5430-AE3B-E8BEA130A88A" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole lavasoa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lavasoa" status="sp. nov.">Pheidole lavasoa</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">Figs 37A-F</figureCitation>
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63S</figureCitation>
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<figureCitation captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65S</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. •1 major worker; Toliara; Grand Lavasoa, 25.9km W Tolagnaro;
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<geoCoordinate degrees="25.08767" direction="south" orientation="latitude" precision="1" value="-25.08767">-25.08767</geoCoordinate>
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,
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<geoCoordinate degrees="46.749" direction="east" orientation="longitude" precision="55" value="46.749">46.749</geoCoordinate>
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; alt. 450 m; 30 Nov 2006; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF15392; CASENT0122898 (CASC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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•1w.; same data as for holotype, CASENT0923282 (CASC).
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</paragraph>
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<caption doi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole lavasoa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lavasoa">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole lavasoa</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0923282) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0122898).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Minute species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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HL <0.95 mm and WL <0.8 mm; head in full-face view sub-oval; body dark brown; sides of head with very dense, short, suberect pilosity; entire head distinctly sculptured; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal teeth distinct, large, with rounded apex; outer hypostomal teeth dentate with apex directed indistinctly outward; inner and outer hypostomal teeth not closely spaced and not connected by concavity.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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HL <0.5 mm and WL <0.6 mm, scape, when laid back, surpassing the posterior head margin by two-fifths of its length, propodeal spines reduced to small lobes, head elongate, and oval and body yellowish brown; head sculpture shiny and predominantly smooth, frons with sparse, short, and longitudinal rugulae; mesosoma shiny and smooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 0.94; HW: 0.98; SL: 0.66; EL: 0.12; WL: 0.83; PSL: 0.13; MTL: 0.57; PNW: 0.36; PTW: 0.13; PPW: 0.28; CI: 95.6; SI: 67.0; PSLI: 13.9; PPI: 44.0; PNI: 36.3; MTI: 57.9.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, slightly widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with very dense, short, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, dense, longitudinal and interrupted rugae and smooth interspaces, posteromedial frons with rugae more irregular and directed slightly outward; lateral sides with thick, dense, and irregular rugae, interspaces shiny and smooth or indistinctly rugulate. Occipital lobes with thick, irregular rugae and smooth to rugoreticulate interspaces. Area posterolateral from eyes rugoreticulate to rugofoveolate with additional longitudinal thick rugae. Gena with relatively dense, thick, and longitudinal rugae and smooth to indistinctly rugoreticulate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by three-fifths of its length; pilosity subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37B, D</figureCitation>
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). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth dentate and lower than inner teeth with sharp apex directed slightly outward; inner and outer hypostomal teeth not closely spaced and not connected by concavity (Fig.
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63S</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process very indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderately short, with moderately wide base and acute apex; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37D</figureCitation>
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). Surface shiny with fine rugofoveolae; medial parts of lateral sides of pronotum with reduced sculpture. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with fine foveolae; node smooth to finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally straight to slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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Shiny and smooth; pilosity dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Dark brown; legs, gaster and antennae yellowish brown (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37D, F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 1): HL: 0.48; HW: 0.42; SL: 0.54; EL: 0.1; WL: 0.58; PSL: 0.05; MTL: 0.4; PNW: 0.27; PTW: 0.07; PPW: 0.11; CI: 116.4; SI: 130.8; PSLI: 9.7; PPI: 65.5; PNI: 65.1; MTI: 95.4.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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Cephalic margin indistinctly convex (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37A</figureCitation>
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). Pilosity relatively dense, long, decumbent to suberect. Sculpture shiny and smooth; frons with sparse, short, and longitudinal rugulae; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37A, C</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines reduced to small lobes (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37C</figureCitation>
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). Sculpture shiny and smooth. Pilosity moderately sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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With sparse, erect pilosity (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37C, E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Yellowish brown (Fig.
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<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Pheidole lavasoa sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923282) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0122898)." figureDoi="10.3897/zookeys.949.51269.figure37" httpUri="https://binary.pensoft.net/fig/432206" pageId="0" pageNumber="1">37C, E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
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<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
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<paragraph pageId="0" pageNumber="1">The species was collected at 450 m in elevation, in rainforest. Nest was located in a rotten log.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="comments">
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<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole lavasoa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lavasoa">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole lavasoa</emphasis>
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</taxonomicName>
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sp. nov., described from Grand Lavasoa, Toliara, resembles the parapatric species
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<taxonomicName lsidName="P. andohahela" pageId="0" pageNumber="1" rank="species" species="andohahela">
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<emphasis italics="true" pageId="0" pageNumber="1">P. andohahela</emphasis>
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</taxonomicName>
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sp. nov. known to date from Col de Tanatana in Parc National Andohahela, Toliara. Major workers of both taxa are very similar and can be separated based on slight difference in setosity of lateral sides of head (which is denser and shorter in
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole lavasoa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lavasoa">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole lavasoa</emphasis>
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</taxonomicName>
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sp. nov.) and shape of hypostomal teeth (
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<taxonomicName lsidName="P. lavasoa" pageId="0" pageNumber="1" rank="species" species="lavasoa">
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<emphasis italics="true" pageId="0" pageNumber="1">P. lavasoa</emphasis>
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</taxonomicName>
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sp. nov. has inner hypostomal teeth with rounded apex and outer hypostomal teeth are dentate and indistinctly directed outward). A better resource for species separation are minor workers. Minor workers of
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole lavasoa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lavasoa">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole lavasoa</emphasis>
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</taxonomicName>
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sp. nov. lack transverse rugulae on vertex and frons and their frons have sparse, short, and longitudinal rugulae, mesosoma is entirely smooth, and body is yellowish brown.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |