664 lines
79 KiB
XML
664 lines
79 KiB
XML
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<mods:title id="353FDBE5DE68A47F2FD882139601582C">The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae)</mods:title>
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<mods:namePart id="F126364D5E89F3DBDC4A0564DE806138">Bosmans, Robert</mods:namePart>
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<mods:namePart id="C0B1B194C7DF876002BEE32B24F6CD6B">Kherbouche-Abrous, Ourida</mods:namePart>
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<mods:namePart id="0CF445A25DD94E9C3BDD5669F5503A9B">Benhalima, Souâd</mods:namePart>
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<mods:namePart id="42E756197B5D42AEBFE3C2F6A652101F">Hervé, Christophe</mods:namePart>
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<mods:title id="E694973912FC1AFA426EC68A636E8C96">Zootaxa</mods:title>
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<mods:date id="AC63A74F06D3A89E18905F50EBE3193B">2018</mods:date>
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<mods:number id="26746392B1CE17D7541EE7EB8FB1AEAE">4451</mods:number>
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<paragraph id="564636B0C020B303FF397537FD47FA8A" blockId="53.[151,648,1381,1440]" box="[151,648,1381,1407]" pageId="53" pageNumber="54">
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<heading id="0D0E81DCC020B303FF397537FD47FA8A" bold="true" box="[151,648,1381,1407]" fontSize="11" level="1" pageId="53" pageNumber="54" reason="1">
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<taxonomicName id="91F94D33C020B303FF397537FD47FA8A" ID-CoL="3JMJR" authority="(Thorell, 1871)" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[151,648,1381,1407]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="macellinus">
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<emphasis id="648DEAA2C020B303FF397537FD47FA8A" bold="true" box="[151,648,1381,1407]" pageId="53" pageNumber="54">
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<emphasis id="648DEAA2C020B303FF397537FE07FA8A" bold="true" box="[151,456,1381,1407]" italics="true" pageId="53" pageNumber="54">Haplodrassus macellinus</emphasis>
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(
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<bibRefCitation id="32684B41C020B303FE797537FD4FFA8A" box="[471,640,1381,1407]" pageId="53" pageNumber="54" refId="ref45901" refString="Thorell, T. (1871) Remarks on synonyms of European spiders. Part II. C. J. Lundstrom, Uppsala, 312 pp. [pp. 97 - 228]" type="book chapter">Thorell, 1871</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="564636B0C020B303FF3975D5FE4CFA55" blockId="53.[151,648,1381,1440]" box="[151,387,1415,1440]" pageId="53" pageNumber="54">
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<figureCitation id="CEC22A35C020B303FF3975D5FEFCFA55" box="[151,307,1415,1440]" captionStart="FIGURES 232–240" captionStartId="56.[151,264,1915,1938]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@56.[214,1372,193,1894]" captionTargetPageId="56" captionText="FIGURES 232–240. Haplodrassus macellinus (France, male from Crozes-Hermitage, female from La Ciotat) 232. Male, dorsal aspect; 233. Female, dorsal aspect; 234. Idem, ventral aspect; 235. Male palp, retrolateral view; 236. Idem, ventral view; 237. Male palpal tibia, dorsolateral view; 238. Epigyne, ventral view; 239. Vulva, ventral view; 240. Idem, dorsal view. LS = Lateral sclerite." httpUri="https://zenodo.org/record/1445132/files/figure.png" pageId="53" pageNumber="54">Figs 232‒240</figureCitation>
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, map 8
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</paragraph>
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</subSubSection>
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<subSubSection id="1EE3653BC020B303FF397598FD46F893" pageId="53" pageNumber="54" type="reference_group">
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<paragraph id="564636B0C020B303FF397598FDAAFA2A" blockId="53.[151,1116,1481,1687]" box="[151,613,1481,1503]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="76E9119EC020B303FF397598FDAAFA2A" box="[151,613,1481,1503]" pageId="53" pageNumber="54">
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<taxonomicName id="91F94D33C020B303FF397598FDEEFA2A" authority="Thorell, 1871: 185" authorityName="Thorell" authorityPageNumber="185" authorityYear="1871" box="[151,545,1481,1503]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="macellinus">
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<emphasis id="648DEAA2C020B303FF397598FE91FA2A" box="[151,350,1481,1503]" italics="true" pageId="53" pageNumber="54">Drassus macellinus</emphasis>
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<treatmentCitation id="D75810A1C020B303FECA759BFDEEFA2A" author="" box="[356,545,1481,1503]" page="185" pageId="53" pageNumber="54">
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: 185
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</treatmentCitation>
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</taxonomicName>
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(♂♀).
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</paragraph>
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<paragraph id="564636B0C020B303FF3975BBFC43FA0B" blockId="53.[151,1116,1481,1687]" box="[151,908,1512,1534]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="76E9119EC020B303FF3975BBFC43FA0B" box="[151,908,1512,1534]" pageId="53" pageNumber="54">
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<taxonomicName id="91F94D33C020B303FF3975BBFDF6FA0B" authority="Thorell, 1871: 184" authorityName="Thorell" authorityPageNumber="184" authorityYear="1871" box="[151,569,1512,1534]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="occcidentalis">
|
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<emphasis id="648DEAA2C020B303FF3975BBFEB9FA0B" box="[151,374,1512,1534]" italics="true" pageId="53" pageNumber="54">Drassus occcidentalis</emphasis>
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<treatmentCitation id="D75810A1C020B303FED275BAFDF6FA0B" author="" box="[380,569,1512,1534]" page="184" pageId="53" pageNumber="54">
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: 184
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</treatmentCitation>
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</taxonomicName>
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(
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<specimenCount id="40FFFD39C020B303FDE775BAFD96FA0B" box="[585,601,1512,1534]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
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) synonymy by
|
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<bibRefCitation id="32684B41C020B303FD5775BAFC47FA0B" box="[761,904,1512,1534]" pageId="53" pageNumber="54" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon (1914)</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="564636B0C020B303FF39765AFCA2F9E9" blockId="53.[151,1116,1481,1687]" box="[151,877,1542,1565]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="76E9119EC020B303FF39765AFCA2F9E9" box="[151,877,1542,1565]" pageId="53" pageNumber="54">
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<taxonomicName id="91F94D33C020B303FF39765AFDD5F9E9" authority="Thorell, 1871: 185" authorityName="Thorell" authorityPageNumber="185" authorityYear="1871" box="[151,538,1542,1565]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="nyctalicus">
|
||
<emphasis id="648DEAA2C020B303FF39765AFE98F9E9" box="[151,343,1542,1565]" italics="true" pageId="53" pageNumber="54">Drassus nyctalicus</emphasis>
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<treatmentCitation id="D75810A1C020B303FEF37654FDD5F9E9" author="" box="[349,538,1542,1564]" page="185" pageId="53" pageNumber="54">
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: 185
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</treatmentCitation>
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</taxonomicName>
|
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(
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<specimenCount id="40FFFD39C020B303FD847654FDF5F9E9" box="[554,570,1542,1564]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
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) synonymy by
|
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<bibRefCitation id="32684B41C020B303FD747654FCA6F9E9" box="[730,873,1542,1564]" pageId="53" pageNumber="54" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon (1878)</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="564636B0C020B303FF397674FC98F9CE" blockId="53.[151,1116,1481,1687]" box="[151,855,1573,1595]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="76E9119EC020B303FF397674FC98F9CE" box="[151,855,1573,1595]" pageId="53" pageNumber="54">
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<taxonomicName id="91F94D33C020B303FF397674FEE5F9CE" box="[151,298,1573,1595]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="hebes">
|
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<emphasis id="648DEAA2C020B303FF397674FEE5F9CE" box="[151,298,1573,1595]" italics="true" pageId="53" pageNumber="54">Drassus hebes</emphasis>
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</taxonomicName>
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O.P.-
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<treatmentCitation id="D75810A1C020B303FEF37677FD87F9CE" author="" box="[349,584,1573,1595]" page="390" pageId="53" pageNumber="54">
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: 390
|
||
</treatmentCitation>
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, pl. 51, fig. 15 (
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<specimenCount id="40FFFD39C020B303FD447677FD30F9CE" box="[746,767,1573,1595]" pageId="53" pageNumber="54" type="male">♂</specimenCount>
|
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),
|
||
<emphasis id="648DEAA2C020B303FCBC7674FC9EF9CE" bold="true" box="[786,849,1574,1595]" pageId="53" pageNumber="54">syn. n</emphasis>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="564636B0C020B303FF397617FB93F9AF" blockId="53.[151,1116,1481,1687]" box="[151,1116,1604,1626]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="76E9119EC020B303FF397617FB93F9AF" box="[151,1116,1604,1626]" pageId="53" pageNumber="54">
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<taxonomicName id="91F94D33C020B303FF397617FE32F9AF" authority="Simon, 1878: 132" authorityName="Simon" authorityPageNumber="132" authorityYear="1878" box="[151,509,1604,1626]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="648DEAA2C020B303FF397617FE8DF9AF" box="[151,322,1605,1626]" italics="true" pageId="53" pageNumber="54">Drassus vicarius</emphasis>
|
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<treatmentCitation id="D75810A1C020B303FEE67616FE32F9AF" author="" box="[328,509,1604,1626]" page="132" pageId="53" pageNumber="54">
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<bibRefCitation id="32684B41C020B303FEE67616FE02F9AF" box="[328,461,1604,1626]" pageId="53" pageNumber="54" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon, 1878</bibRefCitation>
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: 132
|
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</treatmentCitation>
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</taxonomicName>
|
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(
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<specimenCount id="40FFFD39C020B303FDA37616FDD1F9AF" box="[525,542,1604,1626]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
|
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), synonymy with
|
||
<taxonomicName id="91F94D33C020B303FD797617FC6BF9AF" box="[727,932,1604,1626]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="subSpecies" species="macellinus" subSpecies="hebes">
|
||
<emphasis id="648DEAA2C020B303FD797617FC6BF9AF" box="[727,932,1604,1626]" italics="true" pageId="53" pageNumber="54">D. macellinus hebes</emphasis>
|
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</taxonomicName>
|
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by
|
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<bibRefCitation id="32684B41C020B303FC677616FB97F9AF" box="[969,1112,1604,1626]" pageId="53" pageNumber="54" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon (1914)</bibRefCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="564636B0C020B303FF397630FC8DF98D" blockId="53.[151,1116,1481,1687]" box="[151,834,1634,1656]" pageId="53" pageNumber="54">
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<treatmentCitationGroup id="76E9119EC020B303FF397630FC8DF98D" box="[151,834,1634,1656]" pageId="53" pageNumber="54">
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<taxonomicName id="91F94D33C020B303FF397630FDBCF98D" authority=": Simon 1914: 140" authorityName="Simon" authorityPageNumber="140" authorityYear="1914" box="[151,627,1634,1656]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="subSpecies" species="macellinus" subSpecies="hebes">
|
||
<emphasis id="648DEAA2C020B303FF397630FE79F98D" box="[151,438,1634,1656]" italics="true" pageId="53" pageNumber="54">Drassodes macellinus hebes</emphasis>
|
||
:
|
||
<treatmentCitation id="D75810A1C020B303FE6D7630FDBCF98D" author="" box="[451,627,1634,1656]" page="140" pageId="53" pageNumber="54">
|
||
<bibRefCitation id="32684B41C020B303FE6D7630FD8EF98D" box="[451,577,1634,1656]" pageId="53" pageNumber="54" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon 1914</bibRefCitation>
|
||
: 140
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, 209, fig. 248 (
|
||
<specimenCount id="40FFFD39C020B303FCA17630FCE9F98D" box="[783,806,1634,1656]" pageId="53" pageNumber="54" type="male">♂</specimenCount>
|
||
<specimenCount id="40FFFD39C020B303FC8B7630FCFAF98D" box="[805,821,1634,1656]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="564636B0C020B303FF3976D3FCF9F962" blockId="53.[151,1116,1481,1687]" box="[151,822,1665,1687]" pageId="53" pageNumber="54">
|
||
<treatmentCitationGroup id="76E9119EC020B303FF3976D3FCF9F962" box="[151,822,1665,1687]" pageId="53" pageNumber="54">
|
||
<taxonomicName id="91F94D33C020B303FF3976D3FD85F962" ID-CoL="3JML8" authority="Di Franco, 1996: 173" authorityName="Di Franco" authorityPageNumber="173" authorityYear="1996" box="[151,586,1665,1687]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C020B303FF3976D3FEA6F962" box="[151,361,1665,1687]" italics="true" pageId="53" pageNumber="54">Haplodrassus vignai</emphasis>
|
||
<treatmentCitation id="D75810A1C020B303FEC176D3FD85F962" author="" box="[367,586,1665,1687]" page="173" pageId="53" pageNumber="54">
|
||
<bibRefCitation id="32684B41C020B303FEC176D3FDD7F962" box="[367,536,1665,1687]" pageId="53" pageNumber="54" refId="ref43938" refString="Di Franco, F. (1996) Gnaphosidae dei Colli Albani (Arachnida, Araneae). Fragmenta Entomologica, 28, 169 - 187." type="journal article">Di Franco, 1996</bibRefCitation>
|
||
: 173
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, figs 1‒4 (
|
||
<specimenCount id="40FFFD39C020B303FD1876D3FD03F962" box="[694,716,1665,1687]" pageId="53" pageNumber="54" type="male">♂</specimenCount>
|
||
<specimenCount id="40FFFD39C020B303FD6376D3FD13F962" box="[717,732,1665,1687]" pageId="53" pageNumber="54" type="female">♀</specimenCount>
|
||
),
|
||
<emphasis id="648DEAA2C020B303FD5F76D3FCFFF963" bold="true" box="[753,816,1665,1686]" pageId="53" pageNumber="54">syn. n</emphasis>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="564636B0C020B303FF3976EFFD46F893" blockId="53.[151,1437,1725,2038]" pageId="53" pageNumber="54">
|
||
<emphasis id="648DEAA2C020B303FF3976EFFF2CF923" bold="true" box="[151,227,1725,1750]" pageId="53" pageNumber="54">Types:</emphasis>
|
||
Type series of
|
||
<taxonomicName id="91F94D33C020B303FE2276ECFDAEF923" authorityName="Thorell" authorityYear="1871" box="[396,609,1725,1750]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C020B303FE2276ECFDAEF923" box="[396,609,1725,1750]" italics="true" pageId="53" pageNumber="54">Drassus macellinus</emphasis>
|
||
</taxonomicName>
|
||
from France, Nice, containing 7♂ and 8 juveniles; NHRS 179a; examined. Holotype ♀ 1 subadult ♀ paratype of
|
||
<taxonomicName id="91F94D33C020B303FDEB76B0FCD4F90F" authorityName="Thorell" authorityYear="1871" box="[581,795,1761,1786]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="nyctalicus">
|
||
<emphasis id="648DEAA2C020B303FDEB76B0FCD4F90F" box="[581,795,1761,1786]" italics="true" pageId="53" pageNumber="54">Drassus nyctalicus</emphasis>
|
||
</taxonomicName>
|
||
from France, Nice; NHRS 179b; examined. Holotype ♀ of
|
||
<taxonomicName id="91F94D33C020B303FF187754FEBCF8E8" authorityName="Simon" authorityYear="1878" box="[182,371,1798,1821]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="648DEAA2C020B303FF187754FEBCF8E8" box="[182,371,1798,1821]" italics="true" pageId="53" pageNumber="54">Drassus vicarius</emphasis>
|
||
</taxonomicName>
|
||
from Corsica; not examined, inavailable. Holotype ♂ of
|
||
<taxonomicName id="91F94D33C020B303FC547754FB52F8EB" box="[1018,1181,1797,1822]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="hebes">
|
||
<emphasis id="648DEAA2C020B303FC547754FB52F8EB" box="[1018,1181,1797,1822]" italics="true" pageId="53" pageNumber="54">Drassus hebes</emphasis>
|
||
</taxonomicName>
|
||
from France, Menton, J.T. Moggridge leg.,
|
||
<bibRefCitation id="32684B41C020B303FE3E7778FE2AF8B7" box="[400,485,1834,1858]" pageId="53" pageNumber="54" refId="ref45079" refString="Pickard-Cambridge, O. (1874) On some new species of Drassides. Proceedings of the Zoological Society of London, 42 (3), 370 - 419." type="journal article">
|
||
<date id="22471070C020B303FE3E7778FE2AF8B7" box="[400,485,1834,1858]" pageId="53" pageNumber="54" value="1874-02">II.1874</date>
|
||
</bibRefCitation>
|
||
; HECO; not examined. Holotype ♂, paratype ♀ of
|
||
<taxonomicName id="91F94D33C020B303FBFA7778FB0DF8B4" ID-CoL="3JML8" authorityName="Di Franco" authorityYear="1996" box="[1108,1218,1834,1857]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C020B303FBFA7778FB0DF8B4" box="[1108,1218,1834,1857]" italics="true" pageId="53" pageNumber="54">H. vignai</emphasis>
|
||
</taxonomicName>
|
||
from Italy, Lazio, Tuscolo; deposited at MZUR; not examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EE3653BC020B303FF697723FB88F803" pageId="53" pageNumber="54" type="diagnosis">
|
||
<paragraph id="564636B0C020B303FF697723FB88F803" blockId="53.[151,1437,1725,2038]" pageId="53" pageNumber="54">
|
||
<emphasis id="648DEAA2C020B303FF697723FEF2F87F" bold="true" box="[199,317,1905,1930]" pageId="53" pageNumber="54">Diagnosis.</emphasis>
|
||
Males are closely related to
|
||
<taxonomicName id="91F94D33C020B303FD2F7720FCCAF87F" authority="Bosmans & Kherbouche-Abrous & Benhalima & Hervé, 2018" authorityName="Bosmans & Kherbouche-Abrous & Benhalima & Hervé" authorityYear="2018" box="[641,773,1906,1930]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="securifer" status="sp. nov.">
|
||
<emphasis id="648DEAA2C020B303FD2F7720FCCAF87F" box="[641,773,1906,1930]" italics="true" pageId="53" pageNumber="54">H. securifer</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7FBE57D9C020B303FCA07723FC82F87F" box="[782,845,1905,1930]" pageId="53" pageNumber="54" rank="species">
|
||
<emphasis id="648DEAA2C020B303FCA07723FC82F87F" bold="true" box="[782,845,1905,1930]" pageId="53" pageNumber="54">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
but are easily distinguished by the much shorter and wider embolic apophysis (
|
||
<figureCitation id="CEC22A35C020B303FE1277C7FDD0F85B" box="[444,543,1941,1966]" captionStart="FIGURES 232–240" captionStartId="56.[151,264,1915,1938]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@56.[214,1372,193,1894]" captionTargetPageId="56" captionText="FIGURES 232–240. Haplodrassus macellinus (France, male from Crozes-Hermitage, female from La Ciotat) 232. Male, dorsal aspect; 233. Female, dorsal aspect; 234. Idem, ventral aspect; 235. Male palp, retrolateral view; 236. Idem, ventral view; 237. Male palpal tibia, dorsolateral view; 238. Epigyne, ventral view; 239. Vulva, ventral view; 240. Idem, dorsal view. LS = Lateral sclerite." httpUri="https://zenodo.org/record/1445132/files/figure.png" pageId="53" pageNumber="54">Figs 237</figureCitation>
|
||
versus 128). Females are distinguished by the elongated epigyne with rounded, narrow hood, a relatively small areola and a longitudinal internal sclerite accompanying laterally the pro-fovea (
|
||
<figureCitation id="CEC22A35C020B303FAC277EBFF37F803" captionStart="FIGURES 232–240" captionStartId="56.[151,264,1915,1938]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@56.[214,1372,193,1894]" captionTargetPageId="56" captionText="FIGURES 232–240. Haplodrassus macellinus (France, male from Crozes-Hermitage, female from La Ciotat) 232. Male, dorsal aspect; 233. Female, dorsal aspect; 234. Idem, ventral aspect; 235. Male palp, retrolateral view; 236. Idem, ventral view; 237. Male palpal tibia, dorsolateral view; 238. Epigyne, ventral view; 239. Vulva, ventral view; 240. Idem, dorsal view. LS = Lateral sclerite." httpUri="https://zenodo.org/record/1445132/files/figure.png" pageId="53" pageNumber="54">Figs 238‒240</figureCitation>
|
||
, LS). However, this lateral sclerite was found to be absent in a few specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="02866638C023B300FF3977C4FBACF813" httpUri="https://zenodo.org/record/1445128/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,258,1942,1964]" targetBox="[206,1381,193,1921]" targetPageId="54">
|
||
<paragraph id="564636B0C023B300FF3977C4FBACF813" blockId="54.[151,1437,1942,2022]" pageId="54" pageNumber="55">
|
||
<emphasis id="648DEAA2C023B300FF3977C4FE94F859" bold="true" box="[151,347,1942,1964]" pageId="54" pageNumber="55">FIGURES 214–222</emphasis>
|
||
.
|
||
<taxonomicName id="91F94D33C023B300FEC877C4FDF3F85E" box="[358,572,1942,1963]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="creticus">
|
||
<emphasis id="648DEAA2C023B300FEC877C4FDF3F85E" box="[358,572,1942,1963]" italics="true" pageId="54" pageNumber="55">Haplodrassus creticus</emphasis>
|
||
</taxonomicName>
|
||
(Greece, Crete, male from Agia Galini, female from Elafonisos). 214. Male, dorsal aspect. 215. Female, dorsal aspect. 216. Idem, ventral aspect; 217. Male palpal tibia, dorsolateral view; 218. Male palp, retrolateral view; 219. Idem, ventral view; 220. Idem, prolateral view; 221. Epigyne, ventral view; 222. Vulva, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="02866638C022B301FF3977C4FC85F813" httpUri="https://zenodo.org/record/1445130/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,258,1942,1964]" targetBox="[206,1381,198,1921]" targetPageId="55">
|
||
<paragraph id="564636B0C022B301FF3977C4FC85F813" blockId="55.[151,1436,1942,2022]" pageId="55" pageNumber="56">
|
||
<emphasis id="648DEAA2C022B301FF3977C4FE93F859" bold="true" box="[151,348,1942,1964]" pageId="55" pageNumber="56">FIGURES 223–231</emphasis>
|
||
.
|
||
<taxonomicName id="91F94D33C022B301FEC477C4FD80F85E" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[362,591,1942,1963]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="invalidus">
|
||
<emphasis id="648DEAA2C022B301FEC477C4FD80F85E" box="[362,591,1942,1963]" italics="true" pageId="55" pageNumber="56">Haplodrassus invalidus</emphasis>
|
||
</taxonomicName>
|
||
(Cyprus, Mercinlik). 223. Male, dorsal aspect; 224. Female, dorsal aspect. 225. Idem, ventral aspect. 226. Male palp, retrolateral view; 227. Idem, ventral view; 228. Idem, prolateral view; 229. Male palpal tibia, dorsolateral view; 230. Epigyne, ventral view; 231. Vulva, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="02866638C02DB30EFF397729FEFAF818" httpUri="https://zenodo.org/record/1445132/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,264,1915,1938]" targetBox="[214,1372,193,1894]" targetPageId="56">
|
||
<paragraph id="564636B0C02DB30EFF397729FEFAF818" blockId="56.[151,1436,1915,2029]" pageId="56" pageNumber="57">
|
||
<emphasis id="648DEAA2C02DB30EFF397729FEA5F865" bold="true" box="[151,362,1915,1938]" pageId="56" pageNumber="57">FIGURES 232–240</emphasis>
|
||
.
|
||
<taxonomicName id="91F94D33C02DB30EFED57729FDB0F864" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[379,639,1915,1937]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02DB30EFED57729FDB0F864" box="[379,639,1915,1937]" italics="true" pageId="56" pageNumber="57">Haplodrassus macellinus</emphasis>
|
||
</taxonomicName>
|
||
(France, male from Crozes-Hermitage, female from La Ciotat) 232. Male, dorsal aspect; 233. Female, dorsal aspect; 234. Idem, ventral aspect; 235. Male palp, retrolateral view; 236. Idem, ventral view; 237. Male palpal tibia, dorsolateral view; 238. Epigyne, ventral view; 239. Vulva, ventral view; 240. Idem, dorsal view. LS = Lateral sclerite.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="1EE3653BC02CB30FFF6970C5FD0FFD38" pageId="57" pageNumber="58" type="discussion">
|
||
<paragraph id="564636B0C02CB30FFF6970C5FD95FE25" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
<emphasis id="648DEAA2C02CB30FFF6970C5FEF4FF45" bold="true" box="[199,315,151,176]" pageId="57" pageNumber="58">Remarks.</emphasis>
|
||
According to the
|
||
<bibRefCitation id="32684B41C02CB30FFDBE70C5FC96FF45" box="[528,857,151,176]" pageId="57" pageNumber="58" refId="ref46054" refString="World Spider Catalog (2017) World Spider Catalog. Version 18.0. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 15 June 2017)" type="url">World Spider Catalog (2017)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="91F94D33C02CB30FFCC670C5FBB0FF45" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[872,1151,151,176]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFCC670C5FBB0FF45" box="[872,1151,151,176]" italics="true" pageId="57" pageNumber="58">Haplodrassus macellinus</emphasis>
|
||
</taxonomicName>
|
||
is a West Mediterranean species. Various authors cited it frequently in
|
||
<collectingCountry id="2EEE7620C02CB30FFDD070EFFD07FF21" box="[638,712,189,212]" name="France" pageId="57" pageNumber="58">France</collectingCountry>
|
||
and
|
||
<collectingCountry id="2EEE7620C02CB30FFD5570EEFCE2FF20" box="[763,813,188,213]" name="Italy" pageId="57" pageNumber="58">Italy</collectingCountry>
|
||
, and in the Maghreb (
|
||
<bibRefCitation id="32684B41C02CB30FFBB870EEFB5AFF21" box="[1046,1173,188,213]" pageId="57" pageNumber="58" refId="ref43743" refString="Denis J., (1937) On a collection of spiders from Algeria. Proceedings of the zoological Society of London, 1937, 1027 - 1060." type="journal article">Denis 1937</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="32684B41C02CB30FFB0E70EEFA84FF21" box="[1184,1355,188,213]" pageId="57" pageNumber="58" refId="ref43909" refString="Di Franco, F. (1994) Contributo alla conoscenza degli Gnaphosidae (Arachnida, Araneae) del Maghreb. Animalia, 19, 193 - 211." type="journal article">Di Franco 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="32684B41C02CB30FFAF870EEFF1FFF0D" pageId="57" pageNumber="58" refId="ref44992" refString="Pavesi, P. (1884) Materiali per lo studio della fauna tunisina raccolti da e L. Doria: Aracnidi. Annali del Museo civico di Storia naturale di Genova, 20, 446 - 486." type="journal article">Pavesi 1884</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="32684B41C02CB30FFF73708DFEABFF0D" box="[221,356,223,248]" pageId="57" pageNumber="58" refId="ref45591" refString="Simon, E. (1885) Etudes sur les Arachnides recueillis en Tunisie en 1883 et 1884 par MM. A. Letourneux, M. Sedillot et Valery Mayet, membres de la mission de l'Exploration scientifique de la Tunisie. In: Exploration scientifique de la Tunisie, publiee sous les auspices du Ministere de l'instruction publique. Zoologie-Arachnides, Paris, pp. 1 - 55." type="book chapter">Simon 1885</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="32684B41C02CB30FFEDE70B2FE65FF0D" box="[368,426,224,248]" pageId="57" pageNumber="58" refId="ref45701" refString="Simon, E. (1908) Arachnides. In: Gadeau de Kerville, H. (Ed.), Voyage zoologique en Khroumirie. University of Michigan Library, Paris, pp. 51 - 56." type="book chapter">1908</bibRefCitation>
|
||
). There are also two citations from
|
||
<collectingCountry id="2EEE7620C02CB30FFC9F70B2FCB1FF0D" box="[817,894,224,248]" name="Greece" pageId="57" pageNumber="58">Greece</collectingCountry>
|
||
and
|
||
<collectingCountry id="2EEE7620C02CB30FFC1B708DFBCDFF0D" box="[949,1026,223,248]" name="Turkey" pageId="57" pageNumber="58">Turkey</collectingCountry>
|
||
(Caporiacco 1948;
|
||
<bibRefCitation id="32684B41C02CB30FFB75708DFA94FF0D" box="[1243,1371,223,248]" pageId="57" pageNumber="58" refId="ref44189" refString="Karol, S. (1966) Spiders of Ankara and environs with a description of a new species Xysticus turcicus (Araneae, Thomisidae). Communications de la Faculte des Sciences de l'Universite d'Ankara, 11 (C), 15 - 32. https: // doi. org / 10.1501 / Commuc _ 0000000151" type="journal article">Karol 1966</bibRefCitation>
|
||
). One can wonder however how these specimens where identified, when the only available figure of
|
||
<taxonomicName id="91F94D33C02CB30FFB237157FAECFEE8" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[1165,1315,260,285]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFB237157FAECFEE8" box="[1165,1315,260,285]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
is a simple sketch of a part of the male palp by
|
||
<bibRefCitation id="32684B41C02CB30FFDB57175FD7CFEB5" box="[539,691,295,320]" pageId="57" pageNumber="58" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon (1914)</bibRefCitation>
|
||
, and the epigyne has never been figured at all.
|
||
<bibRefCitation id="32684B41C02CB30FFB1E7175FA89FEB5" box="[1200,1350,295,320]" pageId="57" pageNumber="58" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon (1914)</bibRefCitation>
|
||
already stated that the species is very variable, when he declared
|
||
<taxonomicName id="91F94D33C02CB30FFCA6711FFC69FE90" box="[776,934,332,357]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFCA6711FFC69FE90" box="[776,934,332,357]" italics="true" pageId="57" pageNumber="58">Drassus hebes</emphasis>
|
||
</taxonomicName>
|
||
O.P.-
|
||
<bibRefCitation id="32684B41C02CB30FFC70711EFB6BFE91" box="[990,1188,332,357]" pageId="57" pageNumber="58" refId="ref45079" refString="Pickard-Cambridge, O. (1874) On some new species of Drassides. Proceedings of the Zoological Society of London, 42 (3), 370 - 419." type="journal article">Cambridge, 1874</bibRefCitation>
|
||
,
|
||
<taxonomicName id="91F94D33C02CB30FFB01711FFF00FE7D" authority="Thorell, 1871" authorityName="Thorell" authorityYear="1871" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nyctalicus">
|
||
<emphasis id="648DEAA2C02CB30FFB01711FFAF1FE90" box="[1199,1342,332,357]" italics="true" pageId="57" pageNumber="58">D. nyctalicus</emphasis>
|
||
<bibRefCitation id="32684B41C02CB30FFAE8711EFF00FE7D" pageId="57" pageNumber="58" refId="ref45901" refString="Thorell, T. (1871) Remarks on synonyms of European spiders. Part II. C. J. Lundstrom, Uppsala, 312 pp. [pp. 97 - 228]" type="book chapter">Thorell, 1871</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="91F94D33C02CB30FFF727123FDEEFE7D" authority="Thorell, 1871" authorityName="Thorell" authorityYear="1871" box="[220,545,367,392]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="648DEAA2C02CB30FFF727123FE4FFE7D" box="[220,384,367,392]" italics="true" pageId="57" pageNumber="58">D. occidentalis</emphasis>
|
||
<bibRefCitation id="32684B41C02CB30FFE27713DFDEEFE7D" box="[393,545,367,392]" pageId="57" pageNumber="58" refId="ref45901" refString="Thorell, T. (1871) Remarks on synonyms of European spiders. Part II. C. J. Lundstrom, Uppsala, 312 pp. [pp. 97 - 228]" type="book chapter">Thorell, 1871</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="91F94D33C02CB30FFD837123FCE2FE7D" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[557,813,367,392]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="typhon">
|
||
<emphasis id="648DEAA2C02CB30FFD837123FD55FE7D" box="[557,666,367,392]" italics="true" pageId="57" pageNumber="58">D. typhon</emphasis>
|
||
<bibRefCitation id="32684B41C02CB30FFD0C713DFCE2FE7D" box="[674,813,367,392]" pageId="57" pageNumber="58" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91F94D33C02CB30FFCC87123FBBCFE7D" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[870,1139,367,392]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="648DEAA2C02CB30FFCC87123FC10FE7D" box="[870,991,369,392]" italics="true" pageId="57" pageNumber="58">D. vicarius</emphasis>
|
||
<bibRefCitation id="32684B41C02CB30FFC46713DFBBCFE7D" box="[1000,1139,367,392]" pageId="57" pageNumber="58" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
all simple variations of
|
||
<taxonomicName id="91F94D33C02CB30FFA2C7123FEC3FE58" baseAuthorityName="Thorell" baseAuthorityYear="1871" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFA2C7123FEC3FE58" italics="true" pageId="57" pageNumber="58">D. macellinus</emphasis>
|
||
</taxonomicName>
|
||
(“des formes très variables”). The descriptions of all these species were carefully analysed by us and if possible, the
|
||
<typeStatus id="89428812C02CB30FFE8A71EAFE9CFE25" box="[292,339,440,464]" pageId="57" pageNumber="58">type</typeStatus>
|
||
material was examined.
|
||
</paragraph>
|
||
<paragraph id="564636B0C02CB30FFF69718EFD0FFD38" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
The
|
||
<typeStatus id="89428812C02CB30FFF55718FFEE4FE00" box="[251,299,477,501]" pageId="57" pageNumber="58">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="91F94D33C02CB30FFE19718FFD9CFE00" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[439,595,476,501]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFE19718FFD9CFE00" box="[439,595,476,501]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
from Nice,
|
||
<collectingCountry id="2EEE7620C02CB30FFD71718FFCE1FE01" box="[735,814,477,500]" name="France" pageId="57" pageNumber="58">France</collectingCountry>
|
||
, was found in the NHRS, which undoubtedly fixes the identity of the species.
|
||
<bibRefCitation id="32684B41C02CB30FFE3471ADFDF2FDED" box="[410,573,511,536]" pageId="57" pageNumber="58" refId="ref45901" refString="Thorell, T. (1871) Remarks on synonyms of European spiders. Part II. C. J. Lundstrom, Uppsala, 312 pp. [pp. 97 - 228]" type="book chapter">Thorell (1871)</bibRefCitation>
|
||
described the male and a juvenile female (“onne omnino adulta?”). The
|
||
<typeStatus id="89428812C02CB30FFAC27252FA53FDED" box="[1388,1436,512,536]" pageId="57" pageNumber="58">type</typeStatus>
|
||
series indeed contains adult males and only subadult females. In the same paper, Thorell described
|
||
<taxonomicName id="91F94D33C02CB30FFAEE7277FEC6FD95" authorityName="Thorell" authorityYear="1871" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nyctalicus">
|
||
<emphasis id="648DEAA2C02CB30FFAEE7277FEC6FD95" italics="true" pageId="57" pageNumber="58">Drassus nyctalicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91F94D33C02CB30FFEED721BFE21FD95" authorityName="Thorell" authorityYear="1871" box="[323,494,583,608]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="648DEAA2C02CB30FFEED721BFE21FD95" box="[323,494,583,608]" italics="true" pageId="57" pageNumber="58">D. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
, of both only females and also from Nice. Examination of the
|
||
<typeStatus id="89428812C02CB30FFB6D721AFB3CFD95" box="[1219,1267,584,608]" pageId="57" pageNumber="58">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="91F94D33C02CB30FFA2F721BFEC6FD70" authorityName="Thorell" authorityYear="1871" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nyctalicus">
|
||
<emphasis id="648DEAA2C02CB30FFA2F721BFEC6FD70" italics="true" pageId="57" pageNumber="58">D. nyctalicus</emphasis>
|
||
</taxonomicName>
|
||
confirms it is the female of
|
||
<taxonomicName id="91F94D33C02CB30FFDE8723FFD2EFD70" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[582,737,620,645]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFDE8723FFD2EFD70" box="[582,737,620,645]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
, as proposed by
|
||
<bibRefCitation id="32684B41C02CB30FFC31723EFBF1FD71" box="[927,1086,620,645]" pageId="57" pageNumber="58" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon (1878)</bibRefCitation>
|
||
. Later,
|
||
<bibRefCitation id="32684B41C02CB30FFB3E723EFAE4FD71" box="[1168,1323,620,645]" pageId="57" pageNumber="58" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon (1914)</bibRefCitation>
|
||
considers
|
||
<taxonomicName id="91F94D33C02CB30FFF3972C3FE8EFD5D" authorityName="Thorell" authorityYear="1871" box="[151,321,655,680]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="648DEAA2C02CB30FFF3972C3FE8EFD5D" box="[151,321,655,680]" italics="true" pageId="57" pageNumber="58">D. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
a simple variation of
|
||
<taxonomicName id="91F94D33C02CB30FFD9572C3FD18FD5D" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[571,727,655,680]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFD9572C3FD18FD5D" box="[571,727,655,680]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
(“forme de
|
||
<taxonomicName id="91F94D33C02CB30FFCCD72C3FC30FD5D" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[867,1023,655,680]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFCCD72C3FC30FD5D" box="[867,1023,655,680]" italics="true" pageId="57" pageNumber="58">D. macellinus</emphasis>
|
||
</taxonomicName>
|
||
”).
|
||
<typeStatus id="89428812C02CB30FFB8A72C2FB93FD5D" box="[1060,1116,656,680]" pageId="57" pageNumber="58">Type</typeStatus>
|
||
material of this species was not available, and Simon’s view is followed here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EE3653BC02CB30FFF69728BFB5FFBAD" pageId="57" pageNumber="58" type="reference_group">
|
||
<paragraph id="564636B0C02CB30FFF69728BFDB6FC19" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
<taxonomicName id="91F94D33C02CB30FFF69728BFEA4FD05" box="[199,363,727,752]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFF69728BFEA4FD05" box="[199,363,727,752]" italics="true" pageId="57" pageNumber="58">Drassus hebes</emphasis>
|
||
</taxonomicName>
|
||
O.P.-
|
||
<bibRefCitation id="32684B41C02CB30FFE0B7285FDBDFD05" box="[421,626,727,752]" pageId="57" pageNumber="58" refId="ref45079" refString="Pickard-Cambridge, O. (1874) On some new species of Drassides. Proceedings of the Zoological Society of London, 42 (3), 370 - 419." type="journal article">Cambridge, 1874</bibRefCitation>
|
||
, described from France, Menton, at
|
||
<quantity id="91019B55C02CB30FFBBF728AFB96FD05" box="[1041,1113,727,752]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="57" pageNumber="58" unit="km" value="20.0">20 km</quantity>
|
||
from the type locality of
|
||
<emphasis id="648DEAA2C02CB30FFA2F728BFED8FCE0" italics="true" pageId="57" pageNumber="58">
|
||
<taxonomicName id="91F94D33C02CB30FFA2F728BFEDCFCE0" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">H. macellinus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
is another problematic species.
|
||
<bibRefCitation id="32684B41C02CB30FFDD372AEFCD7FCE1" box="[637,792,764,789]" pageId="57" pageNumber="58" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon (1878)</bibRefCitation>
|
||
considered the species a synonym of
|
||
<taxonomicName id="91F94D33C02CB30FFB1072AFFA96FCE0" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[1214,1369,764,789]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFB1072AFFA96FCE0" box="[1214,1369,764,789]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
and a subspecies in 1914. Type material was not available, but Cambridge’s figure 15 of the male palpal tibia of
|
||
<taxonomicName id="91F94D33C02CB30FFA947373FA53FCCD" box="[1338,1436,799,824]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFA947373FA53FCCD" box="[1338,1436,799,824]" italics="true" pageId="57" pageNumber="58">D. hebes</emphasis>
|
||
</taxonomicName>
|
||
agrees well the elongated palpal tibia of
|
||
<taxonomicName id="91F94D33C02CB30FFDCA7317FCCFFCA8" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[612,768,836,861]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFDCA7317FCCFFCA8" box="[612,768,836,861]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
. Also the material of
|
||
<taxonomicName id="91F94D33C02CB30FFC547317FB10FCA8" ID-CoL="5HLCR" authorityName=": Denis" authorityYear="1948" box="[1018,1247,836,861]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="macellinus" subSpecies="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFC547317FB10FCA8" box="[1018,1247,836,861]" italics="true" pageId="57" pageNumber="58">D. macellinus hebes</emphasis>
|
||
</taxonomicName>
|
||
examined in the MNHN is identical to
|
||
<taxonomicName id="91F94D33C02CB30FFE35733BFDF6FC75" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[411,569,871,896]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFE35733BFDF6FC75" box="[411,569,871,896]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
. The evident conclusion is
|
||
<taxonomicName id="91F94D33C02CB30FFCD4733BFC11FC75" box="[890,990,871,896]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFCD4733BFC11FC75" box="[890,990,871,896]" italics="true" pageId="57" pageNumber="58">D. hebes</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="91F94D33C02CB30FFB59733BFA5AFC75" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[1271,1429,871,896]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFB59733BFA5AFC75" box="[1271,1429,871,896]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="91F94D33C02CB30FFF3973DFFE26FC51" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[151,489,908,933]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="648DEAA2C02CB30FFF3973DFFE9CFC51" box="[151,339,909,932]" italics="true" pageId="57" pageNumber="58">Drassus vicarius</emphasis>
|
||
<bibRefCitation id="32684B41C02CB30FFEF473DEFE26FC51" box="[346,489,908,933]" pageId="57" pageNumber="58" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
was described on a female from Corse.
|
||
<bibRefCitation id="32684B41C02CB30FFC0473DEFB8AFC51" box="[938,1093,908,933]" pageId="57" pageNumber="58" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon (1914)</bibRefCitation>
|
||
considered it a synonym of
|
||
<taxonomicName id="91F94D33C02CB30FFA2F73DFFE96FC3D" ID-CoL="5HLCR" authorityName=": Denis" authorityYear="1948" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="macellinus" subSpecies="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFA2F73DFFE96FC3D" italics="true" pageId="57" pageNumber="58">D. macellinus hebes</emphasis>
|
||
</taxonomicName>
|
||
. Since the type material is not available, Simon’s vision is followed here and
|
||
<taxonomicName id="91F94D33C02CB30FFB7373E3FA53FC3D" authorityName="Simon" authorityYear="1878" box="[1245,1436,945,968]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vicarius">
|
||
<emphasis id="648DEAA2C02CB30FFB7373E3FA53FC3D" box="[1245,1436,945,968]" italics="true" pageId="57" pageNumber="58">Drassus vicarius</emphasis>
|
||
</taxonomicName>
|
||
enters into the synonymy of
|
||
<taxonomicName id="91F94D33C02CB30FFE787387FDBDFC18" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[470,626,980,1005]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFE787387FDBDFC18" box="[470,626,980,1005]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="564636B0C02CB30FFF6973ABFB5FFBAD" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
<taxonomicName id="91F94D33C02CB30FFF6973ABFDDCFBE5" authority="Simon, 1878" authorityName="Simon" authorityYear="1878" box="[199,531,1015,1040]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="typhon">
|
||
<emphasis id="648DEAA2C02CB30FFF6973ABFEB7FBE5" box="[199,376,1015,1040]" italics="true" pageId="57" pageNumber="58">Drassus typhon</emphasis>
|
||
<bibRefCitation id="32684B41C02CB30FFE2F73A5FDDCFBE5" box="[385,531,1015,1040]" pageId="57" pageNumber="58" refId="ref45521" refString="Simon, E. (1878) Les arachnides de France. Tome IV. Roret, Paris, 334 pp." type="book">Simon, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
was described on the
|
||
<typeStatus id="89428812C02CB30FFCB873AAFCB7FBE5" box="[790,888,1016,1040]" pageId="57" pageNumber="58" type="syntype">syntypes</typeStatus>
|
||
male and female from the Pyrénées-Orientales.
|
||
<bibRefCitation id="32684B41C02CB30FFF39744EFEFCFBC1" box="[151,307,1052,1077]" pageId="57" pageNumber="58" refId="ref45814" refString="Simon, E. (1914) Les arachnides de France. Synopsis general et catalogue des especes francaises de l'ordre des Araneae. Tome VI. 1 re partie. Roret, Paris, 308 pp." type="book">Simon (1914)</bibRefCitation>
|
||
considered it another synonym of
|
||
<taxonomicName id="91F94D33C02CB30FFD13744FFC96FBC0" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[701,857,1052,1077]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFD13744FFC96FBC0" box="[701,857,1052,1077]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
. An adult female identified by Simon shows it is a distinct species and it is taken out of the synonymy of
|
||
<taxonomicName id="91F94D33C02CB30FFD567413FC5CFBAD" ID-CoL="3JMJR" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[760,915,1087,1112]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFD567413FC5CFBAD" box="[760,915,1087,1112]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
. It is described above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EE3653BC02CB30FFF697436FD15FAA1" pageId="57" pageNumber="58" type="materials_examined">
|
||
<paragraph id="564636B0C02CB30FFF697436FD15FAA1" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
The synonymy of
|
||
<taxonomicName id="91F94D33C02CB30FFE387437FDEEFB88" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[406,545,1124,1149]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="invalidus">
|
||
<emphasis id="648DEAA2C02CB30FFE387437FDEEFB88" box="[406,545,1124,1149]" italics="true" pageId="57" pageNumber="58">H. invalidus</emphasis>
|
||
</taxonomicName>
|
||
(O.P.-
|
||
<bibRefCitation id="32684B41C02CB30FFDCC7436FCE1FB89" box="[610,814,1124,1149]" pageId="57" pageNumber="58" refId="ref45031" refString="Pickard-Cambridge, O. (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40 (1), 212 - 354." type="journal article">Cambridge, 1872</bibRefCitation>
|
||
) from
|
||
<collectingCountry id="2EEE7620C02CB30FFCD27436FC72FB88" box="[892,957,1124,1149]" name="Israel" pageId="57" pageNumber="58">Israel</collectingCountry>
|
||
,
|
||
<collectingCountry id="2EEE7620C02CB30FFC647436FBD5FB88" box="[970,1050,1124,1149]" name="Turkey" pageId="57" pageNumber="58">Turkey</collectingCountry>
|
||
and
|
||
<collectingCountry id="2EEE7620C02CB30FFBFC7436FB02FB88" box="[1106,1229,1124,1149]" name="Azerbaijan" pageId="57" pageNumber="58">Azerbaijan</collectingCountry>
|
||
with
|
||
<taxonomicName id="91F94D33C02CB30FFAA17437FEE4FB55" authority="Di Franco, 1996" authorityName="Di Franco" authorityYear="1996" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C02CB30FFAA17437FAB7FB89" box="[1295,1400,1125,1148]" italics="true" pageId="57" pageNumber="58">H. vignai</emphasis>
|
||
<bibRefCitation id="32684B41C02CB30FFA2E7436FEE4FB55" pageId="57" pageNumber="58" refId="ref43938" refString="Di Franco, F. (1996) Gnaphosidae dei Colli Albani (Arachnida, Araneae). Fragmenta Entomologica, 28, 169 - 187." type="journal article">Di Franco, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="2EEE7620C02CB30FFEC174D5FE6CFB55" box="[367,419,1159,1184]" name="Italy" pageId="57" pageNumber="58">Italy</collectingCountry>
|
||
as proposed by
|
||
<bibRefCitation id="32684B41C02CB30FFDF774DBFD2AFB55" box="[601,741,1160,1185]" pageId="57" pageNumber="58" refId="ref44564" refString="Levy, G. (2004) Spiders of the genera Drassodes and Haplodrassus (Araneae, Gnaphosidae) from Israel. Israel Journal of Zoology, 50, 1 - 37. https: // doi. org / 10.1560 / RUQP- 20 ML-VDBA- 3 GKX" type="journal article">Levy (2004)</bibRefCitation>
|
||
is here rejected. Material of recently collected
|
||
<taxonomicName id="91F94D33C02CB30FFB5874DBFAB0FB55" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[1270,1407,1159,1184]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="invalidus">
|
||
<emphasis id="648DEAA2C02CB30FFB5874DBFAB0FB55" box="[1270,1407,1159,1184]" italics="true" pageId="57" pageNumber="58">H. invalidus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="2EEE7620C02CB30FFF3974FEFF27FB31" box="[151,232,1196,1220]" name="Cyprus" pageId="57" pageNumber="58">Cyprus</collectingCountry>
|
||
compared with material collected in
|
||
<collectingCountry id="2EEE7620C02CB30FFD3674FEFD04FB30" box="[664,715,1196,1221]" name="Italy" pageId="57" pageNumber="58">Italy</collectingCountry>
|
||
near the
|
||
<typeStatus id="89428812C02CB30FFC9474FFFCA5FB30" box="[826,874,1197,1221]" pageId="57" pageNumber="58">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="91F94D33C02CB30FFC5D74FFFB92FB31" authorityName="Di Franco" authorityYear="1996" box="[1011,1117,1197,1220]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C02CB30FFC5D74FFFB92FB31" box="[1011,1117,1197,1220]" italics="true" pageId="57" pageNumber="58">H. vignai</emphasis>
|
||
</taxonomicName>
|
||
shows clearly they are two different species. In
|
||
<taxonomicName id="91F94D33C02CB30FFE2E7483FDC4FB1D" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[384,523,1231,1256]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="invalidus">
|
||
<emphasis id="648DEAA2C02CB30FFE2E7483FDC4FB1D" box="[384,523,1231,1256]" italics="true" pageId="57" pageNumber="58">H. invalidus</emphasis>
|
||
</taxonomicName>
|
||
, the tibial apophysis has a claw-like tip, and the embolus has an inner spur-like process, as excellently shown by
|
||
<bibRefCitation id="32684B41C02CB30FFD8E74A6FCFFFAF9" box="[544,816,1268,1293]" pageId="57" pageNumber="58" refId="ref44463" refString="Kovblyuk, M. M., Kastrygina, Z. A. & Omelko, M. M. (2012) A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae). ZooKeys, 205, 59 - 89 https: // doi. org / 10.3897 / zookeys. 205.3491" type="journal article">
|
||
Kovblyuk
|
||
<emphasis id="648DEAA2C02CB30FFD3574A7FD17FAF8" box="[667,728,1268,1293]" italics="true" pageId="57" pageNumber="58">et al.</emphasis>
|
||
(2012)
|
||
</bibRefCitation>
|
||
. These are both absent in
|
||
<taxonomicName id="91F94D33C02CB30FFBC474A7FB18FAF9" authorityName="Di Franco" authorityYear="1996" box="[1130,1239,1269,1292]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C02CB30FFBC474A7FB18FAF9" box="[1130,1239,1269,1292]" italics="true" pageId="57" pageNumber="58">H. vignai</emphasis>
|
||
</taxonomicName>
|
||
, as shown in Di Franco’s figures 1‒4 of
|
||
<taxonomicName id="91F94D33C02CB30FFE17754BFDE8FAC5" authorityName="Di Franco" authorityYear="1996" box="[441,551,1305,1328]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C02CB30FFE17754BFDE8FAC5" box="[441,551,1305,1328]" italics="true" pageId="57" pageNumber="58">H. vignai</emphasis>
|
||
</taxonomicName>
|
||
. Although the
|
||
<typeStatus id="89428812C02CB30FFD4F754AFCDEFAC5" box="[737,785,1304,1328]" pageId="57" pageNumber="58">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="91F94D33C02CB30FFC04754BFBD7FAC5" authorityName="Di Franco" authorityYear="1996" box="[938,1048,1305,1328]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C02CB30FFC04754BFBD7FAC5" box="[938,1048,1305,1328]" italics="true" pageId="57" pageNumber="58">H. vignai</emphasis>
|
||
</taxonomicName>
|
||
was not examined,
|
||
<taxonomicName id="91F94D33C02CB30FFAA0754BFAB3FAC5" authorityName="Di Franco" authorityYear="1996" box="[1294,1404,1305,1328]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="vignai">
|
||
<emphasis id="648DEAA2C02CB30FFAA0754BFAB3FAC5" box="[1294,1404,1305,1328]" italics="true" pageId="57" pageNumber="58">H. vignai</emphasis>
|
||
</taxonomicName>
|
||
is considered here a junior synonym of
|
||
<taxonomicName id="91F94D33C02CB30FFD99756FFD1CFAA0" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[567,723,1340,1365]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFD99756FFD1CFAA0" box="[567,723,1340,1365]" italics="true" pageId="57" pageNumber="58">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EE3653BC02CB30FFF69750DFBE7F948" pageId="57" pageNumber="58" type="description">
|
||
<paragraph id="564636B0C02CB30FFF69750DFCAFFA68" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
<emphasis id="648DEAA2C02CB30FFF69750DFE96FA8D" bold="true" box="[199,345,1375,1400]" pageId="57" pageNumber="58">Description.</emphasis>
|
||
Measurements: Male (n=7): Total length 4.4‒8.2; carapace 2.38‒3.74 long, 1.68‒2.69 wide. Female (n=1): Total length 11.0; carapace 3.71 long, 2.98 wide.
|
||
</paragraph>
|
||
<paragraph id="564636B0C02CB30FFF6975F5FE66F9F2" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">Colour: Carapace brown to dark reddish brown, cephalic part infuscate; chelicerae dark brown; sternum yellowish to reddish brown, darkened to the margins; legs yellowish brown to reddish brown; abdomen grey to brown with paler venter.</paragraph>
|
||
<paragraph id="564636B0C02CB30FFF697646FD38F980" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
Male palp (
|
||
<figureCitation id="CEC22A35C02CB30FFEE27646FE21F9D8" box="[332,494,1556,1581]" captionStart="FIGURES 232–240" captionStartId="56.[151,264,1915,1938]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@56.[214,1372,193,1894]" captionTargetPageId="56" captionText="FIGURES 232–240. Haplodrassus macellinus (France, male from Crozes-Hermitage, female from La Ciotat) 232. Male, dorsal aspect; 233. Female, dorsal aspect; 234. Idem, ventral aspect; 235. Male palp, retrolateral view; 236. Idem, ventral view; 237. Male palpal tibia, dorsolateral view; 238. Epigyne, ventral view; 239. Vulva, ventral view; 240. Idem, dorsal view. LS = Lateral sclerite." httpUri="https://zenodo.org/record/1445132/files/figure.png" pageId="57" pageNumber="58">Figs 235‒237</figureCitation>
|
||
): Tibial apophysis slender, as long as its diameter, constricted in the middle, with broadly truncate tip; embolar apophysis much shorter than embolus, with only slightly bent margins, getting somewhat broader distally, terminally bluntly pointed.
|
||
</paragraph>
|
||
<paragraph id="564636B0C02CB30FFF69762DFBE7F948" blockId="57.[151,1437,151,2013]" pageId="57" pageNumber="58">
|
||
Epigyne (
|
||
<figureCitation id="CEC22A35C02CB30FFE9D762DFE1AF96D" box="[307,469,1663,1688]" captionStart="FIGURES 232–240" captionStartId="56.[151,264,1915,1938]" captionTargetBox="[214,1372,193,1894]" captionTargetId="figure@56.[214,1372,193,1894]" captionTargetPageId="56" captionText="FIGURES 232–240. Haplodrassus macellinus (France, male from Crozes-Hermitage, female from La Ciotat) 232. Male, dorsal aspect; 233. Female, dorsal aspect; 234. Idem, ventral aspect; 235. Male palp, retrolateral view; 236. Idem, ventral view; 237. Male palpal tibia, dorsolateral view; 238. Epigyne, ventral view; 239. Vulva, ventral view; 240. Idem, dorsal view. LS = Lateral sclerite." httpUri="https://zenodo.org/record/1445132/files/figure.png" pageId="57" pageNumber="58">Figs 238‒240</figureCitation>
|
||
): Elongated, 1.4 x longer than wide, with rounded hood, which is much narrower than the total epigyne; pro-fovea somewhat longer than fovea; lateral pockets sac-like.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EE3653BC02CB30CFF697695FD3AFBE8" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="materials_examined">
|
||
<paragraph id="564636B0C02CB30CFF697695FC69FE7D" blockId="57.[151,1437,151,2013]" lastBlockId="58.[151,1437,151,392]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
|
||
<emphasis id="648DEAA2C02CB30FFF697695FDC2F915" bold="true" box="[199,525,1735,1760]" pageId="57" pageNumber="58">Further material examined.</emphasis>
|
||
FRANCE, mixed unknown localities (‘Gallia Meridionalis’), one of them
|
||
<emphasis id="648DEAA2C02CB30FFAFB7695FEC5F8F0" bold="true" italics="true" pageId="57" pageNumber="58">Alpes- Maritimes</emphasis>
|
||
, Menton, 13♂ 16♀, as
|
||
<taxonomicName id="91F94D33C02CB30FFDA576BEFBC4F8F1" authority=", Coll. Simon 2083" authorityName=", Coll. Simon" authorityYear="2083" box="[523,1035,1772,1797]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="macellinus" subSpecies="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFDA576BEFCF2F8F0" box="[523,829,1772,1797]" italics="true" pageId="57" pageNumber="58">Drassodes macellinus hebes</emphasis>
|
||
, Coll. Simon 2083
|
||
</taxonomicName>
|
||
(MNHN AR9228), mixed unknown localities (‘Gallia Meridionalis, Corsica’), 20♂ 7♀, as
|
||
<taxonomicName id="91F94D33C02CB30FFCBD775DFAFDF8DD" authority=", Coll. Simon 2058" authorityName="Coll. Simon" authorityYear="2058" box="[787,1330,1807,1832]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="macellinus" subSpecies="hebes">
|
||
<emphasis id="648DEAA2C02CB30FFCBD775DFB9DF8DD" box="[787,1106,1807,1832]" italics="true" pageId="57" pageNumber="58">Drassodes macellinus hebes</emphasis>
|
||
, Coll. Simon 2058
|
||
</taxonomicName>
|
||
(MNHN AR9229), unknown locality (‘France S.E.’), 2♂, as
|
||
<taxonomicName id="91F94D33C02CB30FFD727766FB58F8B8" authority=", Coll. Dalmas M." authorityName="Coll. Dalmas M." box="[732,1175,1844,1869]" class="Arachnida" family="Gnaphosidae" genus="Drassodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02CB30FFD727766FC02F8B8" box="[732,973,1844,1869]" italics="true" pageId="57" pageNumber="58">Drassodes macellinus</emphasis>
|
||
, Coll. Dalmas M.
|
||
</taxonomicName>
|
||
150 (MNHN AR9230),
|
||
<emphasis id="648DEAA2C02CB30FFF397705FE80F885" bold="true" box="[151,335,1879,1904]" italics="true" pageId="57" pageNumber="58">Alpes -Maritimes</emphasis>
|
||
, Cannes, île de Saint-Honorat,
|
||
<date id="22471070C02CB30FFD0C770AFCC0F885" box="[674,783,1880,1904]" pageId="57" pageNumber="58" value="2013-04-08">8.IV.2013</date>
|
||
, 1♂, P. Ponel leg. (CPO), Menton (N43°46’35” E7°30’15”), 1♀, L. Fage leg. (MNHN AR9223), Nice (43°42’11” E7°15’57”), 7♂, 8 juveniles (NHRS 179a), Nice, 1♀ 1 subadult♀ (as
|
||
<taxonomicName id="91F94D33C02CB30FFE9477F3FDC1F84C" authorityName="Thorell" authorityYear="1871" box="[314,526,1952,1977]" class="Arachnida" family="Gnaphosidae" genus="Drassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nyctalicus">
|
||
<emphasis id="648DEAA2C02CB30FFE9477F3FDC1F84C" box="[314,526,1952,1977]" italics="true" pageId="57" pageNumber="58">Drassus nyctalicus</emphasis>
|
||
</taxonomicName>
|
||
; NHRS 179b),
|
||
<emphasis id="648DEAA2C02CB30FFD1177F2FC50F84C" bold="true" box="[703,927,1952,1977]" italics="true" pageId="57" pageNumber="58">Bouches -du -Rhône</emphasis>
|
||
, La Ciotat, île Verte, 1♀, P. Ponel leg. (CPO), Vauvenargues, La Citadelle (N43°33’14”” E5°36’13”), 1♂♀,
|
||
<date id="22471070C02CB30FFCC97796FC13F829" box="[871,988,1988,2012]" pageId="57" pageNumber="58" value="2013-05-19">19.V.2013</date>
|
||
, P. Ponel leg. (CPO), Drôme, Crozes- Hermitage, Plan d’eau de la Lône, stones in dry grassland,
|
||
<date id="22471070C02FB30CFCDA70CAFC25FF45" box="[884,1002,152,176]" pageId="58" pageNumber="59" value="2013-02-13">13.II.2013</date>
|
||
, P. Dubois leg. (CPO),
|
||
<emphasis id="648DEAA2C02FB30CFABA70CAFAF7FF5A" bold="true" box="[1300,1336,152,175]" italics="true" pageId="58" pageNumber="59">Var</emphasis>
|
||
, Callian (N43°33’0” E6°50’0”), 1♀, Coll. Berland (MNHN AR9227), Ollières, Basses Selves (N43°28’58” E5°49’47”),
|
||
<quantity id="91019B55C02FB30CFF39708DFF16FF0D" box="[151,217,223,248]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="58" pageNumber="59" unit="m" value="540.0">540m</quantity>
|
||
, dry grassland,
|
||
<date id="22471070C02FB30CFE23708DFDC0FF0D" box="[397,527,223,248]" pageId="58" pageNumber="59" value="2012-05-17">17.05.2012</date>
|
||
, P. Ponel leg. (CPP), Pontrève, 1♂, under stones,
|
||
<date id="22471070C02FB30CFBE7708DFB7EFF0D" box="[1097,1201,223,248]" pageId="58" pageNumber="59" value="2015-05-02">2.V.2015</date>
|
||
, P. Oger leg. (CPO), Rians, 1♂, under stones,
|
||
<date id="22471070C02FB30CFE1E7156FDD8FEE8" box="[432,535,260,285]" pageId="58" pageNumber="59" value="2015-05-01">1.V.2015</date>
|
||
, P. Oger leg. (CPO), Roquebrune-sur-Argens, 1♂, stones in maquis,
|
||
<date id="22471070C02FB30CFAB77156FA57FEE8" box="[1305,1432,260,285]" pageId="58" pageNumber="59" value="2016-04-12">12.IV.2016</date>
|
||
, S. Lecigne leg. (CSL); Saint-Cyr-sur-Mer, Calanque de Port d’Alon (N43°8'48” E5°42'36”), 1♀,
|
||
<date id="22471070C02FB30CFAB37175FA53FEB5" box="[1309,1436,295,320]" pageId="58" pageNumber="59" value="1997-08-06">6.VIII.1997</date>
|
||
(NHML), Valcros (43°14’7”6°2’10”), 1♀,
|
||
<date id="22471070C02FB30CFDC5711EFD27FE91" box="[619,744,332,356]" pageId="58" pageNumber="59" value="1989-03-22">22.III.1989</date>
|
||
, P. Poot leg. (CRB). ITALY,
|
||
<emphasis id="648DEAA2C02FB30CFB8D711EFBB2FE96" bold="true" box="[1059,1149,332,355]" italics="true" pageId="58" pageNumber="59">Toscana</emphasis>
|
||
, Livorno, Isola di Capraia (N43°3’1” E9°48’52”), 1♂,
|
||
<date id="22471070C02FB30CFE64713DFD88FE7D" box="[458,583,367,392]" pageId="58" pageNumber="59" value="1992-11-26">26.XI.1992</date>
|
||
, Pantini & Valle leg. (MCSNB).
|
||
</paragraph>
|
||
<paragraph id="564636B0C02FB30CFF397455FD3AFBE8" blockId="58.[151,757,1031,1053]" box="[151,757,1031,1053]" pageId="58" pageNumber="59">
|
||
<emphasis id="648DEAA2C02FB30CFF397455FF2CFBE9" bold="true" box="[151,227,1031,1052]" pageId="58" pageNumber="59">MAP 8</emphasis>
|
||
. Distribution of
|
||
<taxonomicName id="91F94D33C02FB30CFE227455FD3EFBE8" authority="(Thorell)" authorityName="Thorell" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[396,753,1031,1053]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02FB30CFE227455FD42FBE8" box="[396,653,1031,1053]" italics="true" pageId="58" pageNumber="59">Haplodrassus macellinus</emphasis>
|
||
(Thorell)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1EE3653BC02FB30CFF69743FFD03FACF" pageId="58" pageNumber="59" type="distribution">
|
||
<paragraph id="564636B0C02FB30CFF69743FFD03FACF" blockId="58.[151,1436,1133,1338]" pageId="58" pageNumber="59">
|
||
<emphasis id="648DEAA2C02FB30CFF69743FFE96FB73" bold="true" box="[199,345,1133,1158]" pageId="58" pageNumber="59">Distribution</emphasis>
|
||
. According to the literature and the World spider
|
||
<bibRefCitation id="32684B41C02FB30CFC3E743FFBF3FB73" box="[912,1084,1133,1158]" pageId="58" pageNumber="59" refId="ref46054" refString="World Spider Catalog (2017) World Spider Catalog. Version 18.0. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 15 June 2017)" type="url">Catalog (2017)</bibRefCitation>
|
||
, the species occurs in the West European and African Mediterranean region. All material from the Maghreb identified as
|
||
<taxonomicName id="91F94D33C02FB30CFB2774C1FAEAFB5F" baseAuthorityName="Thorell" baseAuthorityYear="1871" box="[1161,1317,1169,1194]" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="macellinus">
|
||
<emphasis id="648DEAA2C02FB30CFB2774C1FAEAFB5F" box="[1161,1317,1169,1194]" italics="true" pageId="58" pageNumber="59">H. macellinus</emphasis>
|
||
</taxonomicName>
|
||
that could be examined appeared to be misidentified, and the species is not present in our abundant material. Only the records from
|
||
<collectingCountry id="2EEE7620C02FB30CFF7A7488FEEFFB04" box="[212,288,1242,1265]" name="France" pageId="58" pageNumber="59">France</collectingCountry>
|
||
including
|
||
<collectingRegion id="943DF852C02FB30CFE36748BFE3FFB07" box="[408,496,1241,1266]" country="France" name="Corse" pageId="58" pageNumber="59">
|
||
<collectingCountry id="2EEE7620C02FB30CFE36748BFE3FFB07" box="[408,496,1241,1266]" name="France" pageId="58" pageNumber="59">Corsica</collectingCountry>
|
||
</collectingRegion>
|
||
, and
|
||
<collectingCountry id="2EEE7620C02FB30CFD85748BFD91FB07" box="[555,606,1241,1266]" name="Italy" pageId="58" pageNumber="59">Italy</collectingCountry>
|
||
including
|
||
<collectingRegion id="943DF852C02FB30CFD78748BFCD1FB07" box="[726,798,1241,1266]" country="Italy" name="Sicilia" pageId="58" pageNumber="59">Sicilia</collectingRegion>
|
||
are confirmed here (map 8). Citations from
|
||
<collectingCountry id="2EEE7620C02FB30CFAA0748BFAA2FB07" box="[1294,1389,1241,1266]" name="Portugal" pageId="58" pageNumber="59">Portugal</collectingCountry>
|
||
and
|
||
<collectingCountry id="2EEE7620C02FB30CFF3974AFFF18FAE3" box="[151,215,1277,1302]" name="Spain" pageId="58" pageNumber="59">Spain</collectingCountry>
|
||
are possibly correct but should be confirmed. Citations from Rhodes by Caporiacco (1948) and from
|
||
<collectingCountry id="2EEE7620C02FB30CFAE274AFFA53FAE3" box="[1356,1436,1277,1302]" name="Turkey" pageId="58" pageNumber="59">Turkey</collectingCountry>
|
||
by
|
||
<bibRefCitation id="32684B41C02FB30CFF147573FE84FACF" box="[186,331,1313,1338]" pageId="58" pageNumber="59" refId="ref44189" refString="Karol, S. (1966) Spiders of Ankara and environs with a description of a new species Xysticus turcicus (Araneae, Thomisidae). Communications de la Faculte des Sciences de l'Universite d'Ankara, 11 (C), 15 - 32. https: // doi. org / 10.1501 / Commuc _ 0000000151" type="journal article">Karol (1966)</bibRefCitation>
|
||
are considered misidentifications.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |