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<mods:title id="F96218A65F57F4EECB8ACA2DA61D89C9">Review of Amaeana Hartman, 1959 (Annelida, Terebelliformia, Polycirridae), with descriptions of seven new species</mods:title>
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<mods:namePart id="A8B834A2EB560C7F7E7F606946CA666C">Nogueira, João Miguel De Matos</mods:namePart>
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<paragraph id="556036ADFFBDFF9AFF66FAE0DB0CFAAB" blockId="2.[151,550,1301,1327]" box="[151,550,1301,1327]" pageId="2" pageNumber="3">
<heading id="0E2881C1FFBDFF9AFF66FAE0DB0CFAAB" bold="true" box="[151,550,1301,1327]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="67ABEABFFFBDFF9AFF66FAE0DB0CFAAB" bold="true" box="[151,550,1301,1327]" pageId="2" pageNumber="3">
Genus 
<taxonomicName id="92DF4D2EFFBDFF9AFF1FFAE0DB0CFAAB" ID-CoL="84LBD" authority="Hartman, 1959" authorityName="Hartman" authorityYear="1959" box="[238,550,1301,1327]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBDFF9AFF1FFAE0D848FAAB" bold="true" box="[238,354,1301,1327]" italics="true" pageId="2" pageNumber="3">Amaeana</emphasis>
<bibRefCitation id="314E4B5CFFBDFF9AFE99FAE0DB0CFAAB" author="Hartman" box="[360,550,1301,1327]" pageId="2" pageNumber="3" refString="Hartman, O. (1959) Catalogue of the Polychaetous annelids of the world. Part 2. Sedentaria. Allan Hancock Foundation Occasional Paper, 23, 355 - 628." type="journal article" year="1959">Hartman, 1959</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<paragraph id="556036ADFFBDFF9AFF66FAA9DAC5FAF6" blockId="2.[151,1007,1372,1394]" box="[151,1007,1372,1394]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="75CF1183FFBDFF9AFF66FAA9DAC5FAF6" box="[151,1007,1372,1394]" pageId="2" pageNumber="3">
<taxonomicName id="92DF4D2EFFBDFF9AFF66FAA9D8EBFAF6" ID-CoL="84LBD" authority="Hartman 1959: 495" authorityName="Hartman" authorityPageNumber="495" authorityYear="1959" box="[151,449,1372,1394]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="3" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBDFF9AFF66FAA9D9DDFAF5" box="[151,247,1372,1393]" italics="true" pageId="2" pageNumber="3">Amaeana</emphasis>
<treatmentCitation id="D47E10BCFFBDFF9AFF0CFAA8D8EBFAF6" author="Hartman" box="[253,449,1372,1394]" page="495" pageId="2" pageNumber="3" year="1959">
<bibRefCitation id="314E4B5CFFBDFF9AFF0CFAA8D8BBFAF6" author="Hartman" box="[253,401,1372,1394]" pageId="2" pageNumber="3" refString="Hartman, O. (1959) Catalogue of the Polychaetous annelids of the world. Part 2. Sedentaria. Allan Hancock Foundation Occasional Paper, 23, 355 - 628." type="journal article" year="1959">Hartman 1959</bibRefCitation>
: 495
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</taxonomicName>
(replacement for 
<taxonomicName id="92DF4D2EFFBDFF9AFD87FAA9DA45FAF6" ID-CoL="9G2MZ" authority="Malmgren, 1866" authorityName="Malmgren" authorityYear="1866" box="[630,879,1372,1394]" class="Gastropoda" family="Epitoniidae" genus="Amaea" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
<emphasis id="67ABEABFFFBDFF9AFD87FAA9DB97FAF5" box="[630,701,1372,1393]" italics="true" pageId="2" pageNumber="3">Amaea</emphasis>
<bibRefCitation id="314E4B5CFFBDFF9AFD32FAA9DA45FAF6" author="Malmgren" box="[707,879,1372,1394]" pageId="2" pageNumber="3" refString="Malmgren, A. J. (1866) Nordiska Hafs-Annulater Oefv. Forhandlinger Videnskabs Selskabet i Kristiania, 22, 344 - 410." type="journal article" year="1866">Malmgren, 1866</bibRefCitation>
</taxonomicName>
; homonym).
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<paragraph id="556036ADFFBDFF9AFF66FA6ADD2CFA33" blockId="2.[151,1436,1438,2003]" box="[151,1030,1438,1463]" pageId="2" pageNumber="3">
<typeStatus id="8A64880FFFBDFF9AFF66FA6AD9E5FA33" box="[151,207,1439,1463]" pageId="2" pageNumber="3">Type</typeStatus>
species. 
<taxonomicName id="92DF4D2EFFBDFF9AFEC5FA6BDBB4FA33" authority="Sars, 1863" authorityName="Sars" authorityYear="1863" box="[308,670,1438,1463]" class="Polychaeta" family="Terebellidae" genus="Polycirrus" kingdom="Animalia" order="Terebellida" pageId="2" pageNumber="3" phylum="Annelida" rank="species" species="trilobatus">
<emphasis id="67ABEABFFFBDFF9AFEC5FA6BDB34FA32" box="[308,542,1438,1462]" italics="true" pageId="2" pageNumber="3">Polycirrus trilobatus</emphasis>
<bibRefCitation id="314E4B5CFFBDFF9AFDD4FA6ADBB4FA33" author="Sars" box="[549,670,1438,1463]" pageId="2" pageNumber="3" refString="Sars, M. (1863) Geologiske og zoologiske lagttagelser, anstillede paa en Reise i en Deel af Trondhjems stift i Sommeren 1862. Nyt magazin for naturvidenskaberne, 12, 253 - 340." type="journal article" year="1863">Sars, 1863</bibRefCitation>
</taxonomicName>
, designated by 
<bibRefCitation id="314E4B5CFFBDFF9AFCBAFA6ADD2BFA33" author="Hartman" box="[843,1025,1438,1463]" pageId="2" pageNumber="3" refString="Hartman, O. (1959) Catalogue of the Polychaetous annelids of the world. Part 2. Sedentaria. Allan Hancock Foundation Occasional Paper, 23, 355 - 628." type="journal article" year="1959">Hartman (1959)</bibRefCitation>
.
</paragraph>
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<paragraph id="556036ADFFBDFF9AFF36FA37D887F90A" blockId="2.[151,1436,1438,2003]" pageId="2" pageNumber="3">
<emphasis id="67ABEABFFFBDFF9AFF36FA37D86BFA5F" bold="true" box="[199,321,1474,1499]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Transverse prostomium attached to dorsal surface of upper lip; basal part as thick crest, eyespots absent; distal part restricted to base of upper lip, with flaring lobes and frequently also mid-dorsal process. Buccal tentacles of three 
<typeStatus id="8A64880FFFBDFF9AFE80F9FED885F9A7" box="[369,431,1547,1571]" pageId="2" pageNumber="3">types</typeStatus>
, short tentacles thin, uniformly cylindrical, intermediate ones spatulate, long buccal tentacles spatulate or more specialised, with subdistal cylindrical swelling and pointed to blunt tip; apparently, tentacles with longitudinal groove only heavily ciliated at tips, with peduncle smooth or with poorly-developed longitudinal ciliary tract.
</paragraph>
<paragraph id="556036ADFFBDFF9AFF36F96FDA49F953" blockId="2.[151,1436,1438,2003]" pageId="2" pageNumber="3">Peristomium forming lips; upper lip large, frequently circular and convoluted, folded into three lobes, short and swollen lower lip, only mid-ventral and usually button-like.</paragraph>
<paragraph id="556036ADFFBDFF9AFF36F916D86DF857" blockId="2.[151,1436,1438,2003]" pageId="2" pageNumber="3">Segments biannulated throughout, segment 1 reduced, usually visible dorsally and ventrally, laterally covered by expanded prostomium (visibility of segment 1 strongly depends on state of preservation of specimens). Segment 2 distinctly narrower than following segments, separating “head” from remainder of body; segment 2 with rectangular or pentagonal mid-ventral shield at beginning of mid-ventral groove, sometimes extending anteriorly through segment 1 until near posterior margin of lower lip. Body wall papillate throughout, papillae distinctly larger and more abundant on ventro-lateral pads of anterior segments. Pads usually from segment 2 to last segment with notopodia.</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FF62DDC6FF73" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">Notopodia beginning from segment 3, extending for about 10 segments; notopodia elongate and bilobed, lobes about same size; notochaetae throughout usually with distinctly narrow wings, inconspicuous under higher magnifications of light microscopy, clearly visible under SEM; pinnate chaetae sometimes present.</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FEF1D9C8FEE1" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">Neuropodia beginning from midbody, after conspicuous gap of some achaetous segments between termination of notopodia and beginning of neuropodia; neurochaetae as distally tapering acicular spines, sometimes specialised at tips.</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FE9AD888FE29" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">Nephridial and genital papillae present, usually anterior to bases of all notopodia. Pygidium smooth or with rounded ventral papilla.</paragraph>
</subSubSection>
<subSubSection id="1DC56526FFBCFF9BFF36FE42D870FC69" pageId="3" pageNumber="4" type="discussion">
<paragraph id="556036ADFFBCFF9BFF36FE42DA94FE71" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">
<emphasis id="67ABEABFFFBCFF9BFF36FE42D811FE54" bold="true" box="[199,315,439,464]" pageId="3" pageNumber="4">Remarks.</emphasis>
<taxonomicName id="92DF4D2EFFBCFF9BFEB3FE4DD886FE4B" box="[322,428,440,463]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFEB3FE4DD886FE4B" box="[322,428,440,463]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
was originally described as 
<taxonomicName id="92DF4D2EFFBCFF9BFD1AFE4DDAD4FE54" authority="Malmgren, 1866" authorityName="Malmgren" authorityYear="1866" box="[747,1022,439,464]" class="Gastropoda" family="Epitoniidae" genus="Amaea" kingdom="Animalia" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFD1AFE4DDA13FE4B" box="[747,825,440,463]" italics="true" pageId="3" pageNumber="4">Amaea</emphasis>
<bibRefCitation id="314E4B5CFFBCFF9BFCB1FE42DAD4FE54" author="Malmgren" box="[832,1022,439,464]" pageId="3" pageNumber="4" refString="Malmgren, A. J. (1866) Nordiska Hafs-Annulater Oefv. Forhandlinger Videnskabs Selskabet i Kristiania, 22, 344 - 410." type="journal article" year="1866">Malmgren, 1866</bibRefCitation>
</taxonomicName>
, but that name was preoccupied by a mollusk and was subsequently replaced with 
<taxonomicName id="92DF4D2EFFBCFF9BFD63FE29DA93FE71" authority="Hartman 1959" authorityName="Hartman" authorityYear="1959" box="[658,953,476,501]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFD63FE29DBD6FE77" box="[658,764,476,499]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
(
<bibRefCitation id="314E4B5CFFBCFF9BFCFBFE29DA9BFE71" author="Hartman" box="[778,945,476,501]" pageId="3" pageNumber="4" refString="Hartman, O. (1959) Catalogue of the Polychaetous annelids of the world. Part 2. Sedentaria. Allan Hancock Foundation Occasional Paper, 23, 355 - 628." type="journal article" year="1959">Hartman 1959</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FE0AD837FDE4" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">
The most important diagnostic character of 
<taxonomicName id="92DF4D2EFFBCFF9BFD4EFDF5DA03FD93" box="[703,809,512,535]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFD4EFDF5DA03FD93" box="[703,809,512,535]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
is the presence of neuropodia bearing tapering spines, always after a conspicuous gap of several achaetous segments between termination of notopodia and beginning of neuropodia.
</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FD98DDD7FD2C" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">
In a recent phylogenetic study of Polycirridae, 
<bibRefCitation id="314E4B5CFFBCFF9BFD15FD99DAF4FD01" box="[740,990,620,645]" pageId="3" pageNumber="4" refString="Fitzhugh, K., Nogueira, J. M. M., Carrerette, O. &amp; Hutchings, P. (2015) An assessment of the status of Polycirridae genera (Annelida: Terebelliformia) with the evolutionary transformation series of characters within the family. Zoology Journal of Linnean Society, 2015, 1 - 36. [online only]. http: // dx. doi. org / 10.1111 / zoj. 12259" type="journal article">
Fitzhugh 
<emphasis id="67ABEABFFFBCFF9BFCA1FD98DAA1FD00" box="[848,907,620,644]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
(2015)
</bibRefCitation>
found species of 
<taxonomicName id="92DF4D2EFFBCFF9BFB5DFD99DC3CFD07" box="[1196,1302,620,643]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFB5DFD99DC3CFD07" box="[1196,1302,620,643]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
and 
<taxonomicName id="92DF4D2EFFBCFF9BFAA1FD99DCB6FD00" box="[1360,1436,620,644]" class="Polychaeta" family="Terebellidae" genus="Lysilla" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFAA1FD99DCB6FD00" box="[1360,1436,620,644]" italics="true" pageId="3" pageNumber="4">Lysilla</emphasis>
</taxonomicName>
within an apomorphic clade within 
<taxonomicName id="92DF4D2EFFBCFF9BFDD5FD65DBB0FD2C" box="[548,666,656,680]" class="Polychaeta" family="Terebellidae" genus="Polycirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFDD5FD65DBB0FD2C" box="[548,666,656,680]" italics="true" pageId="3" pageNumber="4">Polycirrus</emphasis>
</taxonomicName>
, and therefore none of these genera are monophyletic.
</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FD41DAC2FCD9" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">
Nevertheless, the boundaries between these “genera” are well established morphologically: 
<taxonomicName id="92DF4D2EFFBCFF9BFB04FD41DC41FD48" box="[1269,1387,692,716]" class="Polychaeta" family="Terebellidae" genus="Polycirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFB04FD41DC41FD48" box="[1269,1387,692,716]" italics="true" pageId="3" pageNumber="4">Polycirrus</emphasis>
</taxonomicName>
has notopodia bearing pinnate and/or winged notochaetae, usually broadly-winged 
<emphasis id="67ABEABFFFBCFF9BFBD1FD2CDD74FD74" box="[1056,1118,729,752]" italics="true" pageId="3" pageNumber="4">sensu</emphasis>
<bibRefCitation id="314E4B5CFFBCFF9BFB99FD22DC4CFD74" box="[1128,1382,727,752]" pageId="3" pageNumber="4" refString="Fitzhugh, K., Nogueira, J. M. M., Carrerette, O. &amp; Hutchings, P. (2015) An assessment of the status of Polycirridae genera (Annelida: Terebelliformia) with the evolutionary transformation series of characters within the family. Zoology Journal of Linnean Society, 2015, 1 - 36. [online only]. http: // dx. doi. org / 10.1111 / zoj. 12259" type="journal article">
Fitzhugh 
<emphasis id="67ABEABFFFBCFF9BFB24FD2CDC3AFD74" box="[1237,1296,728,752]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
(2015)
</bibRefCitation>
, and neuropodia with uncini; 
<taxonomicName id="92DF4D2EFFBCFF9BFE38FD09DB3EFC90" box="[457,532,764,788]" class="Polychaeta" family="Terebellidae" genus="Lysilla" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFE38FD09DB3EFC90" box="[457,532,764,788]" italics="true" pageId="3" pageNumber="4">Lysilla</emphasis>
</taxonomicName>
has notopodia only, bearing either pinnate or narrowly-winged, acicular notochaetae; and 
<taxonomicName id="92DF4D2EFFBCFF9BFEACFCD5D8EDFCB3" box="[349,455,800,823]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFEACFCD5D8EDFCB3" box="[349,455,800,823]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
has notopodia usually only bearing acicular notochaetae, although pinnate chaetae are present in two species, and neuropodia with spines, usually distally tapered.
</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FC92D870FC69" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">
Although these differences are conspicuous and easily recognizable in complete specimens, it is sometimes very difficult to distinguish between incomplete members of 
<taxonomicName id="92DF4D2EFFBCFF9BFCA2FC79DAB4FC20" box="[851,926,908,932]" class="Polychaeta" family="Terebellidae" genus="Lysilla" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFCA2FC79DAB4FC20" box="[851,926,908,932]" italics="true" pageId="3" pageNumber="4">Lysilla</emphasis>
</taxonomicName>
and 
<taxonomicName id="92DF4D2EFFBCFF9BFC29FC79DD68FC27" box="[984,1090,908,931]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFC29FC79DD68FC27" box="[984,1090,908,931]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
if neuropodia are not present, and sometimes specimens from museum collections are misidentified, when members of both genera co-occur in the same locality.
</paragraph>
</subSubSection>
<subSubSection id="1DC56526FFBCFF9FFF36FC0DD844FA61" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="556036ADFFBCFF9BFF36FC0DD8D0FB6C" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">
<taxonomicName id="92DF4D2EFFBCFF9BFF36FC0DD81BFB8B" ID-CoL="84LBD" box="[199,305,1016,1039]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFF36FC0DD81BFB8B" box="[199,305,1016,1039]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
currently includes 12 species, considering the seven new ones described herein. Two other species, 
<taxonomicName id="92DF4D2EFFBCFF9BFF66FBE9D896FBB1" authority="McIntosh, 1926" authorityName="McIntosh" authorityYear="1926" box="[151,444,1052,1077]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="colei">
<emphasis id="67ABEABFFFBCFF9BFF66FBE9D9C7FBB0" box="[151,237,1052,1076]" italics="true" pageId="3" pageNumber="4">A. colei</emphasis>
(
<bibRefCitation id="314E4B5CFFBCFF9BFF0FFBE9D89EFBB1" author="McIntosh" box="[254,436,1052,1077]" pageId="3" pageNumber="4" refString="McIntosh, W. C. (1926) Additions to to the marine fauna of St Andrews since 1874. Annals Magazine Natural History, S eries 9, 18, 241 - 266." type="journal article" year="1926">McIntosh, 1926</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="92DF4D2EFFBCFF9BFE04FBE9DA16FBB1" ID-CoL="669LB" authority="Augener, 1926" authorityName="Augener" authorityYear="1926" box="[501,828,1052,1077]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="67ABEABFFFBCFF9BFE04FBE9DB50FBB0" box="[501,634,1052,1076]" italics="true" pageId="3" pageNumber="4">A. antipoda</emphasis>
(
<bibRefCitation id="314E4B5CFFBCFF9BFD7BFBE9DA1EFBB1" author="Augener" box="[650,820,1052,1077]" pageId="3" pageNumber="4" refString="Augener, H. (1926) Papers from Dr. T. H. Mortensen's Pacific Expedition 1914 - 1916. No. 34. Polychaeta Ill. Polychaeten von Neuseeland. 11. Sedentaria. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i Kjobenhavn, 81, 157 - 294." type="journal article" year="1926">Augener, 1926</bibRefCitation>
)
</taxonomicName>
are both treated as 
<emphasis id="67ABEABFFFBCFF9BFBEFFBE8DD85FBB0" box="[1054,1199,1052,1076]" italics="true" pageId="3" pageNumber="4">nomem nuda</emphasis>
. 
<taxonomicName id="92DF4D2EFFBCFF9BFB4EFBE9DC4DFBB0" box="[1215,1383,1052,1076]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="colei">
<emphasis id="67ABEABFFFBCFF9BFB4EFBE9DC4DFBB0" box="[1215,1383,1052,1076]" italics="true" pageId="3" pageNumber="4">Amaeana colei</emphasis>
</taxonomicName>
was described based on an anterior fragment of 5–6 segments (
<bibRefCitation id="314E4B5CFFBCFF9BFCD2FBCADAF9FBDC" author="McIntosh" box="[803,979,1087,1112]" pageId="3" pageNumber="4" refString="McIntosh, W. C. (1926) Additions to to the marine fauna of St Andrews since 1874. Annals Magazine Natural History, S eries 9, 18, 241 - 266." type="journal article" year="1926">McIntosh 1926</bibRefCitation>
); we could not locate that specimen, but, even if we did, it would still be undeterminable unless complete material from the 
<typeStatus id="8A64880FFFBCFF9BFBB8FB90DD53FBF9" box="[1097,1145,1125,1149]" pageId="3" pageNumber="4">type</typeStatus>
locality is found, as with only 5–6 segments it is not possible to determine if the specimen belongs to 
<taxonomicName id="92DF4D2EFFBCFF9BFC09FB7DDD48FB1B" ID-CoL="84LBD" box="[1016,1122,1160,1183]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFC09FB7DDD48FB1B" box="[1016,1122,1160,1183]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
, 
<taxonomicName id="92DF4D2EFFBCFF9BFB81FB7DDD91FB24" ID-CoL="84PVR" box="[1136,1211,1160,1184]" class="Polychaeta" family="Terebellidae" genus="Lysilla" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFB81FB7DDD91FB24" box="[1136,1211,1160,1184]" italics="true" pageId="3" pageNumber="4">Lysilla</emphasis>
</taxonomicName>
, 
<taxonomicName id="92DF4D2EFFBCFF9BFB3BFB7DDC6AFB24" ID-CoL="84SJF" box="[1226,1344,1160,1184]" class="Polychaeta" family="Terebellidae" genus="Polycirrus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFB3BFB7DDC6AFB24" box="[1226,1344,1160,1184]" italics="true" pageId="3" pageNumber="4">Polycirrus</emphasis>
</taxonomicName>
or even 
<taxonomicName id="92DF4D2EFFBCFF9BFF66FB59D87AFB40" ID-CoL="84NFZ" box="[151,336,1196,1220]" class="Polychaeta" family="Terebellidae" genus="Enoplobranchus" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFF66FB59D87AFB40" box="[151,336,1196,1220]" italics="true" pageId="3" pageNumber="4">Enoplobranchus</emphasis>
</taxonomicName>
, since the differences between those genera are only visible from midbody. Therefore, 
<taxonomicName id="92DF4D2EFFBCFF9BFADAFB59DCABFB40" box="[1323,1409,1196,1220]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="colei">
<emphasis id="67ABEABFFFBCFF9BFADAFB59DCABFB40" box="[1323,1409,1196,1220]" italics="true" pageId="3" pageNumber="4">A. colei</emphasis>
</taxonomicName>
is treated herein as 
<emphasis id="67ABEABFFFBCFF9BFEA4FB24D8D9FB6C" box="[341,499,1232,1256]" italics="true" pageId="3" pageNumber="4">nomen nudum</emphasis>
.
</paragraph>
<paragraph id="556036ADFFBCFF9BFF36FB00DD0AFA45" blockId="3.[151,1437,151,1473]" pageId="3" pageNumber="4">
In the case of 
<taxonomicName id="92DF4D2EFFBCFF9BFE92FB01D8CCFA88" ID-CoL="669LB" box="[355,486,1268,1292]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="67ABEABFFFBCFF9BFE92FB01D8CCFA88" box="[355,486,1268,1292]" italics="true" pageId="3" pageNumber="4">A. antipoda</emphasis>
</taxonomicName>
, material from Western 
<collectingCountry id="2DC8763DFFBCFF9BFD05FB01DA71FA89" box="[756,859,1268,1293]" name="Australia" pageId="3" pageNumber="4">Australia</collectingCountry>
previously identified as belonging to this species is described herein as a new species, because the original description does not mention several characters important to distinguish this species from the remaining within the genus, and no material of this species from the 
<typeStatus id="8A64880FFFBCFF9BFAE0FAC8DC6BFAD1" box="[1297,1345,1341,1365]" pageId="3" pageNumber="4">type</typeStatus>
locality or nearby regions is currently available for study; a modern description for specimens of 
<taxonomicName id="92DF4D2EFFBCFF9BFB71FA95DC29FAFC" ID-CoL="669LB" box="[1152,1283,1376,1400]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="67ABEABFFFBCFF9BFB71FA95DC29FAFC" box="[1152,1283,1376,1400]" italics="true" pageId="3" pageNumber="4">A. antipoda</emphasis>
</taxonomicName>
from another region of 
<collectingCountry id="2DC8763DFFBCFF9BFEF5FA70D8B3FA19" box="[260,409,1412,1437]" name="New Zealand" pageId="3" pageNumber="4">New Zealand</collectingCountry>
will be provided elsewhere (C. Glasby personal communication), but until that is available, it is not possible to compare 
<taxonomicName id="92DF4D2EFFBCFF9BFE28FA5DDB76FA44" ID-CoL="669LB" box="[473,604,1448,1472]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="antipoda">
<emphasis id="67ABEABFFFBCFF9BFE28FA5DDB76FA44" box="[473,604,1448,1472]" italics="true" pageId="3" pageNumber="4">A. antipoda</emphasis>
</taxonomicName>
with several other species of this genus.
</paragraph>
<paragraph id="556036ADFFBCFF9BFF66FA1ADD58F98C" blockId="3.[151,1138,1519,1545]" box="[151,1138,1519,1545]" pageId="3" pageNumber="4">
The distinction between the currently known species of this genus is based on (
<tableCitation id="185D0316FFBCFF9BFBFDFA1ADD4BF98C" box="[1036,1121,1519,1544]" captionStart="TABLE 1" captionStartId="4.[119,199,561,582]" captionText="TABLE 1. Comparison of selected morphological characters between species of Amaeana." pageId="3" pageNumber="4">Table 1</tableCitation>
):
</paragraph>
<paragraph id="556036ADFFBCFF9BFF66F9CDD9F4F8AC" blockId="3.[151,1436,1591,2013]" pageId="3" pageNumber="4">
(1) the presence or absence of a mid-dorsal process originating from the distal part of prostomium, which also occurs in some species of 
<taxonomicName id="92DF4D2EFFBCFF9BFE4BF9A9DB2FF9F0" ID-CoL="84PVR" box="[442,517,1628,1652]" class="Polychaeta" family="Terebellidae" genus="Lysilla" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFE4BF9A9DB2FF9F0" box="[442,517,1628,1652]" italics="true" pageId="3" pageNumber="4">Lysilla</emphasis>
</taxonomicName>
. Some species of 
<taxonomicName id="92DF4D2EFFBCFF9BFD3FF9A9DA12F9F7" ID-CoL="84LBD" box="[718,824,1628,1651]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBCFF9BFD3FF9A9DA12F9F7" box="[718,824,1628,1651]" italics="true" pageId="3" pageNumber="4">Amaeana</emphasis>
</taxonomicName>
have a clearly marked mid-dorsal process on the distal part of the prostomium (for instance, see 
<figureCitation id="CDE42A28FFBCFF9BFD80F98ADB92F91C" box="[625,696,1663,1688]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="3" pageNumber="4">Figs 9</figureCitation>
A, G; 14B, G), in others it is not as clearly marked, but it is still conspicuous (
<figureCitation id="CDE42A28FFBCFF9BFEC3F951D8ACF938" box="[306,390,1700,1725]" captionStart="FIGURE 28" captionStartId="47.[151,250,1914,1936]" captionTargetBox="[155,1429,197,1891]" captionTargetId="figure@47.[151,1436,193,1892]" captionTargetPageId="47" captionText="FIGURE 28. Amaeana brasiliensis sp. nov. holotype MZUSP 2349. A – C, thorax, ventral, dorsal and left lateral views, respectively. D – E, close ups of the anterior end, ventral and dorsal views, respectively. F, anterior end, right lateral view. G, buccal tentacles. H, posterior end, right dorso-lateral view. Numbers refer to segments; li = lower lip; P = basal part of prostomium; PP = prostomial process; ls = upper lip. Scale bars: A – C, F, H = 1 mm; D – E, G = 0.5 mm." httpUri="https://zenodo.org/record/289165/files/figure.png" pageId="3" pageNumber="4">Fig. 28</figureCitation>
B, E), while in still others such a prostomial process is apparently absent (
<figureCitation id="CDE42A28FFBCFF9BFB25F951DC35F938" box="[1236,1311,1700,1725]" captionStart="FIGURE 4" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1431,229,1310]" captionTargetId="figure@13.[152,1435,225,1319]" captionTargetPageId="13" captionText="FIGURE 4. Amaeana breviachaeta sp. nov., holotype AM W. 46526. A – B, incomplete worm, right ventro-lateral and left dorso-lateral views, respectively. C – D, anterior end, right ventro-lateral and left dorso-lateral views, respectively; arrows point to last notopodium. E – F, close ups of the anterior end, left dorso-lateral and ventral views, respectively. G, transition between thorax and abdomen; upper arrow points to last notopodium, lower arrow points to first neuropodium. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.4 mm; C – D, G = 0.3 mm; E – F = 0.2 mm." httpUri="https://zenodo.org/record/289141/files/figure.png" pageId="3" pageNumber="4">Figs 4</figureCitation>
B, E; 25F). However, although this character is easily visible in specimens missing buccal tentacles, it is frequently difficult to ascertain the presence of such processes in specimens with large numbers of buccal tentacles remaining (
<figureCitation id="CDE42A28FFBCFF9BFAC3F919DCA9F880" box="[1330,1411,1772,1797]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="3" pageNumber="4">Fig. 18</figureCitation>
B, D, G).
</paragraph>
<paragraph id="556036ADFFBCFF9BFF36F8C1DD45F859" blockId="3.[151,1436,1591,2013]" pageId="3" pageNumber="4">
<bibRefCitation id="314E4B5CFFBCFF9BFF36F8C1DB3BF8C9" author="Hutchings" box="[199,529,1844,1869]" pageId="3" pageNumber="4" refString="Hutchings, P. A. &amp; Glasby, C. J. (1986) The Polycirrinae (F. Terebellidae) of Australia. Records of the Australian Museum, 38, 319 - 350. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.38.1986.185" type="journal article" year="1986">Hutchings &amp; Glasby (1986)</bibRefCitation>
reported cocoon-like structures, possibly egg tubes, among the material the authors identified as 
<taxonomicName id="92DF4D2EFFBCFF9BFE74F8ADDB2DF8F4" ID-CoL="CDB6" box="[389,519,1880,1904]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="trilobata">
<emphasis id="67ABEABFFFBCFF9BFE74F8ADDB2DF8F4" box="[389,519,1880,1904]" italics="true" pageId="3" pageNumber="4">A. trilobata</emphasis>
</taxonomicName>
(in part herein described as 
<taxonomicName id="92DF4D2EFFBCFF9BFCBCF8ADDADDF8F4" ID-CoL="CDB3" box="[845,1015,1880,1904]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="3" pageNumber="4" phylum="Annelida" rank="species" species="ellobophora" status="sp. nov.">
<emphasis id="67ABEABFFFBCFF9BFCBCF8ADDADDF8F4" box="[845,1015,1880,1904]" italics="true" pageId="3" pageNumber="4">A. ellobophora</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFBCFF9BFC0EF8A2DD70F8F4" bold="true" box="[1023,1114,1879,1904]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="7C9857C4FFBCFF9BFC0EF8A2DD70F8F4" box="[1023,1114,1879,1904]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, see below); such structures are attached to the distal part of prostomium, in a position corresponding to the mid-dorsal prostomial process (
<figureCitation id="CDE42A28FFBCFF9BFA9EF889D98BF83C" captionStart="FIGURE 1" captionStartId="8.[151,250,1538,1560]" captionTargetBox="[157,1431,805,1488]" captionTargetId="figure@8.[151,1435,801,1491]" captionTargetPageId="8" captionText="FIGURE 1. Amaeana ellobophora, paratype 1 AM W. 10341, with egg cocoons: A, entire worm, dorsal view. B, anterior end, dorsal view. Scale bars: A = 0.5 mm; B = 0.3 mm." httpUri="https://zenodo.org/record/289138/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
A–B). It seems likely that the presence of a mid-dorsal prostomial process may suggest they brood their embryos, although egg cocoons have not been observed in any other species of this genus to date.
</paragraph>
<caption id="01A06625FFBBFF9CFF86FDC4DAFBFDC2" box="[119,977,561,583]" pageId="4" pageNumber="5">
<paragraph id="556036ADFFBBFF9CFF86FDC4DAFBFDC2" blockId="4.[119,977,561,583]" box="[119,977,561,583]" pageId="4" pageNumber="5">
<emphasis id="67ABEABFFFBBFF9CFF86FDC4D9F4FDC2" bold="true" box="[119,222,561,582]" pageId="4" pageNumber="5">TABLE 1.</emphasis>
Comparison of selected morphological characters between species of 
<taxonomicName id="92DF4D2EFFBBFF9CFC83FDC4DAE1FDC3" ID-CoL="84LBD" box="[882,971,561,583]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBBFF9CFC83FDC4DAE1FDC3" box="[882,971,561,583]" italics="true" pageId="4" pageNumber="5">Amaeana</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="556036ADFFBBFF9CFEEAFD9DDE7EFDFF" blockId="4.[119,1904,616,1627]" box="[283,1876,616,635]" pageId="4" pageNumber="5">Body Prostomial Modified Upper lip Lower ip Ventro- Number of Beginning of Number of Nephridial/ Pygidium</paragraph>
<paragraph id="556036ADFFBBFF9CFEEAFD71DF98FD13" blockId="4.[119,1904,616,1627]" box="[283,1714,644,663]" pageId="4" pageNumber="5">measurement (H process buccal lateral pads pairs of neuropodia achaetous Genital</paragraph>
<paragraph id="556036ADFFBBFF9CFEEAFD54DF92FD30" blockId="4.[119,1904,616,1627]" box="[283,1720,673,692]" pageId="4" pageNumber="5">
= 
<typeStatus id="8A64880FFFBBFF9CFEDCFD54D850FD30" box="[301,378,673,692]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
; L = tentacles (segs); notopodia (seg.); number segments; papillae
</paragraph>
<paragraph id="556036ADFFBBFF9CFEEAFD48DF8CFD54" blockId="4.[119,1904,616,1627]" box="[283,1702,701,720]" pageId="4" pageNumber="5">
<typeStatus id="8A64880FFFBBFF9CFEEAFD48D846FD54" box="[283,364,701,720]" pageId="4" pageNumber="5" type="lectotype">lectotype</typeStatus>
; P = mid-ventral (segs); 
<typeStatus id="8A64880FFFBBFF9CFB11FD48DC22FD54" box="[1248,1288,701,720]" pageId="4" pageNumber="5">type</typeStatus>
(s) of spines and length (segs)
</paragraph>
<paragraph id="556036ADFFBBFF9CFEEAFD2CDF63FD68" blockId="4.[119,1904,616,1627]" box="[283,1609,729,748]" pageId="4" pageNumber="5">
<typeStatus id="8A64880FFFBBFF9CFEEAFD2CD84CFD68" box="[283,358,729,748]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
) shield of of notochaetae morphology proportional
</paragraph>
<paragraph id="556036ADFFBBFF9CFBE2FD00DF07FC8C" blockId="4.[119,1904,616,1627]" box="[1043,1581,757,776]" pageId="4" pageNumber="5">segment 2 to thorax</paragraph>
<paragraph id="556036ADFFBBFF9CFF86FCE7DFFBFCDA" blockId="4.[119,1904,616,1627]" pageId="4" pageNumber="5">
<taxonomicName id="92DF4D2EFFBBFF9CFF86FCE7D9E2FCA1" ID-CoL="84LBD" box="[119,200,786,805]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBBFF9CFF86FCE7D9E2FCA1" box="[119,200,786,805]" italics="true" pageId="4" pageNumber="5">Amaeana</emphasis>
</taxonomicName>
H: ~ 
<quantity id="92279B48FFBBFF9CFEB0FCE6D8A7FCA2" box="[321,397,787,806]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="5" unit="mm" value="14.5">14.5 mm</quantity>
Rectangular to Apparently Circular Relatively 3–15; small, 12–13 (
<date id="2161106DFFBBFF9CFB13FCE6DC7AFCA2" box="[1250,1360,786,806]" pageId="4" pageNumber="5" value="1921-03-14">3–14 21</date>
; 8 slightly 5–6 (15–16 to 3 to 14–15 Unknown 
<taxonomicName id="92DF4D2EFFBBFF9CFF86FCDAD9F8FCC5" ID-CoL="5TXLB" box="[119,210,815,833]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="accraensis">
<emphasis id="67ABEABFFFBBFF9CFF86FCDAD9F8FCC5" box="[119,210,815,833]" italics="true" pageId="4" pageNumber="5">accraensis</emphasis>
</taxonomicName>
long, 
<quantity id="92279B48FFBBFF9CFEBDFCDAD8A7FCC6" box="[332,397,815,834]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="5" unit="mm" value="2.1">2.1 mm</quantity>
crescent-like absent, few large, rounded rectangular to (left side) or hooked, distally 20); shorter (all 
<bibRefCitation id="314E4B5CFFBBFF9CFF8CFCBED9D4FCDA" author="Augener" box="[125,254,843,862]" pageId="4" pageNumber="5" refString="Augener, H. (1918) Polychaeta Beitr a ge zur Kenntnis des Meeresfauna West-Afrikas. Herausgegeben von W. Michaelsen Hamburg, 2, 67 - 625." type="journal article" year="1918">Augener, 1918</bibRefCitation>
) wide (incomplete) tentacles to oblong pentagonal 15 (right side); blunt notopodia)
</paragraph>
<paragraph id="556036ADFFBBFF9CFD94FC92DC22FCFE" blockId="4.[119,1904,616,1627]" box="[613,1288,871,890]" pageId="4" pageNumber="5">remaining acicular and</paragraph>
<paragraph id="556036ADFFBBFF9CFD94FC76DDF5FC12" blockId="4.[119,1904,616,1627]" box="[613,1247,899,918]" pageId="4" pageNumber="5">on specimen pinnate</paragraph>
<paragraph id="556036ADFFBBFF9CFF86FC6ADE11FC6F" blockId="4.[119,1904,616,1627]" pageId="4" pageNumber="5">
<taxonomicName id="92DF4D2EFFBBFF9CFF86FC6AD9E2FC36" ID-CoL="84LBD" box="[119,200,927,946]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBBFF9CFF86FC6AD9E2FC36" box="[119,200,927,946]" italics="true" pageId="4" pageNumber="5">Amaeana</emphasis>
</taxonomicName>
H: 
<quantity id="92279B48FFBBFF9CFEC7FC55D858FC37" box="[310,370,928,947]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="mm" value="20.0">20 mm</quantity>
long, 3 Present, Present Circular Rounded to 2–15; large, 10 (3–12); 21; 4–5 thin, 8 (13–20); 3-12 (all Smooth, 
<taxonomicName id="92DF4D2EFFBBFF9CFF86FC4ED991FC4A" ID-CoL="65XLB" box="[119,187,955,974]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="angulus">
<emphasis id="67ABEABFFFBBFF9CFF86FC4ED991FC4A" box="[119,187,955,974]" italics="true" pageId="4" pageNumber="5">angulus</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFBBFF9CFF31FC49D9F0FC4B" bold="true" box="[192,218,956,975]" pageId="4" pageNumber="5">sp.</emphasis>
mm wide rectangular crescent-like pentagonal acicular distally tapered twice as long notopodia); ventral. spines, with enlarged on papilla
</paragraph>
<paragraph id="556036ADFFBBFF9CFAC8FC01DE1DFB83" blockId="4.[119,1904,616,1627]" box="[1337,1847,1012,1031]" pageId="4" pageNumber="5">thinner knob- segs 6–10 absent</paragraph>
<paragraph id="556036ADFFBBFF9CFAC8FBE4DC5FFBA0" blockId="4.[119,1904,616,1627]" box="[1337,1397,1041,1060]" pageId="4" pageNumber="5">like tip</paragraph>
<paragraph id="556036ADFFBBFF9CFF86FBD9DE11FB10" blockId="4.[119,1904,616,1627]" pageId="4" pageNumber="5">
<taxonomicName id="92DF4D2EFFBBFF9CFF86FBD9D9E2FBBB" ID-CoL="84LBD" box="[119,200,1068,1087]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBBFF9CFF86FBD9D9E2FBBB" box="[119,200,1068,1087]" italics="true" pageId="4" pageNumber="5">Amaeana</emphasis>
</taxonomicName>
H: 
<quantity id="92279B48FFBBFF9CFEC7FBD8D858FBC4" box="[310,370,1069,1088]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="5" unit="mm" value="12.0">12 mm</quantity>
long, Rounded to Present Circular Rounded to 3–12; small, 9–10 (3–11 or 22–23; 
<date id="2161106DFFBBFF9CFA8BFBD8DF20FBC4" box="[1402,1546,1068,1088]" pageId="4" pageNumber="5" value="2010-09-01" valueMax="2010-09-02" valueMin="2010-09-01">1–2 9–10</date>
(
<date id="2161106DFFBBFF9CF9E6FBD8DFB4FBC4" box="[1559,1694,1068,1088]" pageId="4" pageNumber="5" value="2012-03-13" valueMax="2012-11-13" valueMin="2012-03-13">12–13 3–11</date>
or 12 With 
<taxonomicName id="92DF4D2EFFBBFF9CFF86FBBDD992FBDF" ID-CoL="5TXMF" box="[119,184,1096,1115]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="apheles">
<emphasis id="67ABEABFFFBBFF9CFF86FBBDD992FBDF" box="[119,184,1096,1115]" italics="true" pageId="4" pageNumber="5">apheles</emphasis>
</taxonomicName>
~ 
<quantity id="92279B48FFBBFF9CFED7FBBCD842FBD8" box="[294,360,1097,1116]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="5" unit="mm" value="2.5">2.5 mm</quantity>
long squared rectangular, pentagonal 12); acicular thick, sharp to 21–22); (all rounded Hutchings, (incomplete). button-like (pinnate under spines twice as long notopodia); ventral) Longest specimen SEM) enlarged on papilla
</paragraph>
<paragraph id="556036ADFFBBFF9CFEEAFB6BDFE3FB35" blockId="4.[119,1904,616,1627]" box="[283,1737,1181,1201]" pageId="4" pageNumber="5">examined ~15 segs 5–10</paragraph>
<paragraph id="556036ADFFBBFF9CFEEAFB4FD885FB49" blockId="4.[119,1904,616,1627]" box="[283,431,1210,1229]" pageId="4" pageNumber="5">
mm long, ~ 
<quantity id="92279B48FFBBFF9CFE8CFB4FD885FB49" box="[381,431,1210,1229]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="mm" value="2.0">2 mm</quantity>
</paragraph>
<paragraph id="556036ADFFBBFF9CFEEAFB23D86FFB6D" blockId="4.[119,1904,616,1627]" box="[283,325,1238,1257]" pageId="4" pageNumber="5">wide</paragraph>
<paragraph id="556036ADFFBBFF9CFF86FB04DE16FABA" blockId="4.[119,1904,616,1627]" pageId="4" pageNumber="5">
<taxonomicName id="92DF4D2EFFBBFF9CFF86FB04D9E2FA80" ID-CoL="84LBD" box="[119,200,1265,1284]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBBFF9CFF86FB04D9E2FA80" box="[119,200,1265,1284]" italics="true" pageId="4" pageNumber="5">Amaeana</emphasis>
</taxonomicName>
H: ~ 
<quantity id="92279B48FFBBFF9CFEB0FB07D859FA81" box="[321,371,1266,1285]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="5" unit="mm" value="7.0">7 mm</quantity>
long, Rounded, Present Elliptical, Rounded, 2–13; small, 11 (3–13); 21; 2–3 spines, 7 (14–20); 3–13; With 
<taxonomicName id="92DF4D2EFFBBFF9CFF86FAFBD9F3FAA5" ID-CoL="65XMF" box="[119,217,1294,1313]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="brasiliensis">
<emphasis id="67ABEABFFFBBFF9CFF86FAFBD9F3FAA5" box="[119,217,1294,1313]" italics="true" pageId="4" pageNumber="5">brasiliensis</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFBBFF9CFF2EFAFAD9D2FAA6" bold="true" box="[223,248,1295,1314]" pageId="4" pageNumber="5">sp.</emphasis>
<quantity id="92279B48FFBBFF9CFEEAFAFAD876FAA6" box="[283,348,1295,1314]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="mm" value="1.5">1.5 mm</quantity>
wide raised higher than button-like pentagonal acicular distally blunt, or slightly enlarged on rounded. broad with flattened, shorter to segs 6–10 ventral
</paragraph>
<paragraph id="556036ADFFBBFF9CFAC8FAB2DC95FA2B" blockId="4.[119,1904,616,1627]" pageId="4" pageNumber="5">
oblique tips same length papilla 
<taxonomicName id="92DF4D2EFFBBFF9CFF86FA97D9E2FAF1" ID-CoL="84LBD" box="[119,200,1378,1397]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBBFF9CFF86FA97D9E2FAF1" box="[119,200,1378,1397]" italics="true" pageId="4" pageNumber="5">Amaeana</emphasis>
</taxonomicName>
H: ~ 
<quantity id="92279B48FFBBFF9CFEB3FA96D859FAF2" box="[322,371,1379,1398]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="mm" value="5.0">5 mm</quantity>
long, Absent Present Circular, Rounded, 2–11; small 9 (3–11); 15; 5–6 thin, 3 (12–14); 3–11 (all Unknown 
<taxonomicName id="92DF4D2EFFBBFF9CFF86FA8BD9CCFA15" ID-CoL="669MF" box="[119,230,1406,1425]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="breviachaeta">
<emphasis id="67ABEABFFFBBFF9CFF86FA8BD9CCFA15" box="[119,230,1406,1425]" italics="true" pageId="4" pageNumber="5">breviachaeta</emphasis>
</taxonomicName>
<quantity id="92279B48FFBBFF9CFEEAFA8AD876FA16" box="[283,348,1407,1426]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="5" unit="mm" value="1.1">1.1 mm</quantity>
wide slightly button-like acicular tapering spines, shorter notopodia) 
<emphasis id="67ABEABFFFBBFF9CFF67FA69D996FA2B" bold="true" box="[150,188,1436,1455]" pageId="4" pageNumber="5">nov.</emphasis>
(incomplete) convoluted with hooked tip
</paragraph>
<paragraph id="556036ADFFBBFF9CFF86FA42DE11F987" blockId="4.[119,1904,616,1627]" pageId="4" pageNumber="5">
<taxonomicName id="92DF4D2EFFBBFF9CFF86FA42D9E2FA4E" ID-CoL="84LBD" box="[119,200,1463,1482]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBBFF9CFF86FA42D9E2FA4E" box="[119,200,1463,1482]" italics="true" pageId="4" pageNumber="5">Amaeana</emphasis>
</taxonomicName>
H: 
<quantity id="92279B48FFBBFF9CFEC7FA4DD84DFA4F" box="[310,359,1464,1483]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="5" unit="mm" value="7.0">7 mm</quantity>
long, 1 Absent Tentacles Higher than Rounded, 2–12; large, 10 (3–12); 21; single spine 8 (13–20); 3–10, middle Crenulate, 
<taxonomicName id="92DF4D2EFFBBFF9CFF86FA26D9D3FA62" ID-CoL="5TXMG" box="[119,249,1491,1510]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="4" pageNumber="5" phylum="Annelida" rank="species" species="crassispinulata">
<emphasis id="67ABEABFFFBBFF9CFF86FA26D9D3FA62" box="[119,249,1491,1510]" italics="true" pageId="4" pageNumber="5">crassispinulata</emphasis>
</taxonomicName>
mm wide missing on broad, button-like pentagonal acicular per slightly longer ones slightly ventral 
<emphasis id="67ABEABFFFBBFF9CFF67FA04D996F980" bold="true" box="[150,188,1521,1540]" pageId="4" pageNumber="5">nov.</emphasis>
<typeStatus id="8A64880FFFBBFF9CFD94FA05DBA0F987" box="[613,650,1520,1539]" pageId="4" pageNumber="5">type</typeStatus>
elliptical neuropodium, enlarged papilla
</paragraph>
<paragraph id="556036ADFFBBFF9CFD94F9F9DE71F99B" blockId="4.[119,1904,616,1627]" box="[613,1883,1548,1567]" pageId="4" pageNumber="5">specimen basally broader, apparently</paragraph>
<paragraph id="556036ADFFBBFF9CFAC8F9DCDE12F9B8" blockId="4.[119,1904,616,1627]" box="[1337,1848,1577,1596]" pageId="4" pageNumber="5">distally truncate absent</paragraph>
<paragraph id="556036ADFFBBFF9DF9B8F9B0DF18FD70" blockId="4.[119,1904,616,1627]" lastBlockId="5.[125,1901,564,1632]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="67ABEABFFFBBFF9CF9B8F9B0DE5AF9DF" box="[1609,1904,1605,1627]" italics="true" pageId="4" pageNumber="5">……continued on the next page</emphasis>
Body Prostomial Modified Upper lip Lower ip Ventro- Number of Beginning of Number of Nephridial/ Pygidium measurement (H process buccal lateral pads pairs of neuropodia achaetous Genital = 
<typeStatus id="8A64880FFFBAFF9DFEC3FD78D855FD24" box="[306,383,653,672]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
; L = tentacles (segs); notopodia (seg.); number segments; papillae 
<typeStatus id="8A64880FFFBAFF9DFED0FD5CD858FD38" box="[289,370,681,700]" pageId="5" pageNumber="6" type="lectotype">lectotype</typeStatus>
; P = mid-ventral (segs); 
<typeStatus id="8A64880FFFBAFF9DFB15FD5CDC26FD38" box="[1252,1292,681,700]" pageId="5" pageNumber="6">type</typeStatus>
(s) of spines and length (segs) 
<typeStatus id="8A64880FFFBAFF9DFED0FD30D846FD5C" box="[289,364,709,728]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
) shield of of notochaetae morphology proportional segment 2 to thorax
</paragraph>
<caption id="01A06625FFBAFF9DFF8CFDC1D871FDCD" box="[125,347,564,585]" pageId="5" pageNumber="6">
<paragraph id="556036ADFFBAFF9DFF8CFDC1D871FDCD" blockId="5.[125,1901,564,1632]" box="[125,347,564,585]" pageId="5" pageNumber="6">
<emphasis id="67ABEABFFFBAFF9DFF8CFDC1D9C9FDCD" bold="true" box="[125,227,564,585]" pageId="5" pageNumber="6">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="556036ADFFBAFF9DFF8CFD0BDC7FFCE2" blockId="5.[125,1901,564,1632]" pageId="5" pageNumber="6">
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFD0BD9E4FC95" ID-CoL="84LBD" box="[125,206,766,785]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBAFF9DFF8CFD0BD9E4FC95" box="[125,206,766,785]" italics="true" pageId="5" pageNumber="6">Amaeana</emphasis>
</taxonomicName>
H: 
<quantity id="92279B48FFBAFF9DFECAFD0BD847FC95" box="[315,365,766,785]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="mm" value="7.0">7 mm</quantity>
long, 2 Absent Apparently Elliptical, Rounded to 2–13; large, 11 (3–13); 26–27; 
<date id="2161106DFFBAFF9DFA8EFD0BDF31FC95" box="[1407,1563,766,785]" pageId="5" pageNumber="6" value="2012-04-13" valueMax="2012-05-13" valueMin="2012-04-13">4-5 12–13</date>
(14 to 3–10 (all Unknown 
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFCEFD9C5FCA9" ID-CoL="CDB2" box="[125,239,794,813]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="dampierensis">
<emphasis id="67ABEABFFFBAFF9DFF8CFCEFD9C5FCA9" box="[125,239,794,813]" italics="true" pageId="5" pageNumber="6">dampierensis</emphasis>
</taxonomicName>
mm wide absent, few higher than rectangular, pentagonal acicular sharp spines, 25–26); twice notopodia);. 
<emphasis id="67ABEABFFFBAFF9DFF6AFCC2D9E8FCCE" bold="true" box="[155,194,823,842]" pageId="5" pageNumber="6">nov.</emphasis>
(incomplete) tentacles broad button-like with flattened as long all small remaining tip
</paragraph>
<paragraph id="556036ADFFBAFF9DFD9AFC9ADBFDFC06" blockId="5.[125,1901,564,1632]" box="[619,727,879,898]" pageId="5" pageNumber="6">on specimen</paragraph>
<paragraph id="556036ADFFBAFF9DFF8CFC7EDC50FB8B" blockId="5.[125,1901,564,1632]" pageId="5" pageNumber="6">
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFC7ED9E4FC1A" ID-CoL="84LBD" box="[125,206,907,926]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBAFF9DFF8CFC7ED9E4FC1A" box="[125,206,907,926]" italics="true" pageId="5" pageNumber="6">Amaeana</emphasis>
</taxonomicName>
H: 
<quantity id="92279B48FFBAFF9DFECAFC7ED85DFC1A" box="[315,375,907,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="6" unit="mm" value="16.0">16 mm</quantity>
long, 3 Rounded, Apparently Elliptical, Relatively 2–12; large, 10 (3–12); 18; 8–11 thin, 5 (13– 17); 3–12 (all With 
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFC52D9CFFC3E" ID-CoL="CDB3" box="[125,229,935,954]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="ellobophora">
<emphasis id="67ABEABFFFBAFF9DFF8CFC52D9CFFC3E" box="[125,229,935,954]" italics="true" pageId="5" pageNumber="6">ellobophora</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFBAFF9DFF1BFC5DD82EFC3F" bold="true" box="[234,260,936,955]" pageId="5" pageNumber="6">sp.</emphasis>
mm wide raised absent, few higher than large, rounded, pentagonal acicular distally tapered shorter, ½–2/3 notopodia) rounded. (incomplete). tentacles broad button-like spines, with as long ventral Longest specimen remaining thinner knob- papilla examined ~15 on specimen like tip
</paragraph>
<paragraph id="556036ADFFBAFF9DFED0FBEDD89EFBAF" blockId="5.[125,1901,564,1632]" box="[289,436,1048,1067]" pageId="5" pageNumber="6">
mm long, ~ 
<quantity id="92279B48FFBAFF9DFE72FBEDD89EFBAF" box="[387,436,1048,1067]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="2.0">2 mm</quantity>
</paragraph>
<paragraph id="556036ADFFBAFF9DFED0FBC0D897FBCC" blockId="5.[125,1901,564,1632]" box="[289,445,1077,1096]" pageId="5" pageNumber="6">wide (incomplete)</paragraph>
<paragraph id="556036ADFFBAFF9DFF8CFBA5DFE4FB75" blockId="5.[125,1901,564,1632]" pageId="5" pageNumber="6">
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFBA5D9E4FBE7" ID-CoL="84LBD" box="[125,206,1104,1123]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBAFF9DFF8CFBA5D9E4FBE7" box="[125,206,1104,1123]" italics="true" pageId="5" pageNumber="6">Amaeana</emphasis>
</taxonomicName>
H: 
<quantity id="92279B48FFBAFF9DFECAFBA4D85DFBE0" box="[315,375,1105,1124]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="6" unit="mm" value="17.0">17 mm</quantity>
long, 1 Absent Present Higher than Rounded to 3–12; large, 10 (3–12); 20; 2–3 thin, 7 (13–19); 3–12 or 13 With 
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFB99D994FBFB" ID-CoL="CDB4" box="[125,190,1132,1151]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
<emphasis id="67ABEABFFFBAFF9DFF8CFB99D994FBFB" box="[125,190,1132,1151]" italics="true" pageId="5" pageNumber="6">hsiehae</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFBAFF9DFF32FB98D823FB04" bold="true" box="[195,265,1133,1152]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="7C9857C4FFBAFF9DFF32FB98D823FB04" box="[195,265,1133,1152]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
mm wide; longest broad, rectangular, pentagonal pinnate tapering spines, same length (all rounded 
<typeStatus id="8A64880FFFBAFF9DFED0FB7CD840FB18" box="[289,362,1161,1180]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
<quantity id="92279B48FFBAFF9DFE9EFB7CD881FB18" box="[367,427,1161,1180]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="6" unit="mm" value="34.0">34 mm</quantity>
almost button-like slightly or slightly notopodia in ventral long, ~ 
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rectangular expanded longer most specs), papilla wide subdistally, with enlarged on short oblique tip segs 6–10
</paragraph>
<paragraph id="556036ADFFBAFF9DFF8CFB0FDCE6FAFA" blockId="5.[125,1901,564,1632]" pageId="5" pageNumber="6">
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFB0FD9E4FA89" ID-CoL="84LBD" box="[125,206,1274,1293]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBAFF9DFF8CFB0FD9E4FA89" box="[125,206,1274,1293]" italics="true" pageId="5" pageNumber="6">Amaeana</emphasis>
</taxonomicName>
H: ~ 
<quantity id="92279B48FFBAFF9DFEB6FB0FD8A9FA89" box="[327,387,1274,1293]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="mm" value="65.0">65 mm</quantity>
long, Unknown, Present Circular Rounded, 2–14; small, 12 (3–14); 38; 
<date id="2161106DFFBAFF9DFAAEFB0FDCD1FA89" box="[1375,1531,1274,1293]" pageId="5" pageNumber="6" value="1923-09-08" valueMax="1923-12-08" valueMin="1923-09-08">8–9 to 12– 23</date>
(15–37); 3–14 (all Crenulate, 
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFAE3D9CEFAAD" ID-CoL="CDB5" box="[125,228,1302,1321]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="occidentalis">
<emphasis id="67ABEABFFFBAFF9DFF8CFAE3D9CEFAAD" box="[125,228,1302,1321]" italics="true" pageId="5" pageNumber="6">occidentalis</emphasis>
</taxonomicName>
~ 
<quantity id="92279B48FFBAFF9DFEDDFAE3D877FAAD" box="[300,349,1302,1321]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="6" unit="mm" value="4.0">4 mm</quantity>
wide prostomium button-like pentagonal acicular 13 spines, much longer notopodia); with ventral Hartman, (incomplete) hidden by slightly enlarged on papilla 1944) buccal expanded segments 6–9 tentacles subdistally, with
</paragraph>
<paragraph id="556036ADFFBAFF9DFACEFA72DCB1FA1E" blockId="5.[125,1901,564,1632]" box="[1343,1435,1415,1434]" pageId="5" pageNumber="6">oblique tip</paragraph>
<paragraph id="556036ADFFBAFF9DFF8CFA56DCEFFA6B" blockId="5.[125,1901,564,1632]" pageId="5" pageNumber="6">
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFA56D9E4FA32" ID-CoL="84LBD" box="[125,206,1443,1462]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBAFF9DFF8CFA56D9E4FA32" box="[125,206,1443,1462]" italics="true" pageId="5" pageNumber="6">Amaeana</emphasis>
</taxonomicName>
L: ~ 
<quantity id="92279B48FFBAFF9DFEB5FA51D8AAFA33" box="[324,384,1444,1463]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="6" unit="mm" value="31.0">31 mm</quantity>
long, Rounded to Absent Circular, Rectangular, 3–12; large, 10 (3–12); 16; 7–8 thin, 3 (13–15); 3–12 (all Unknown 
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFA4AD9E2FA56" ID-CoL="CDB6" box="[125,200,1471,1490]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="trilobata">
<emphasis id="67ABEABFFFBAFF9DFF8CFA4AD9E2FA56" box="[125,200,1471,1490]" italics="true" pageId="5" pageNumber="6">trilobata</emphasis>
</taxonomicName>
(Sars, ~ 
<quantity id="92279B48FFBAFF9DFEDDFA35D847FA57" box="[300,365,1472,1491]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="6" unit="mm" value="1.9">1.9 mm</quantity>
wide squared folded into short and broad pentagonal acicular tapering spines, shorter notopodia) 1863) (incomplete) three lobes with hooked tip
</paragraph>
<paragraph id="556036ADFFBAFF9DFF8CFA0DDF0CF9C0" blockId="5.[125,1901,564,1632]" pageId="5" pageNumber="6">
<taxonomicName id="92DF4D2EFFBAFF9DFF8CFA0DD9E4F98F" ID-CoL="84LBD" box="[125,206,1528,1547]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFBAFF9DFF8CFA0DD9E4F98F" box="[125,206,1528,1547]" italics="true" pageId="5" pageNumber="6">Amaeana</emphasis>
</taxonomicName>
P: 
<quantity id="92279B48FFBAFF9DFEC9FA0DD843F98F" box="[312,361,1528,1547]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="mm" value="7.0">7 mm</quantity>
long, 1 Rounded to Present Relatively Rounded to 3–14; small, 12 (3–14); 22–24; 1–2 7–9 (15 to 3–14 (all Unknown 
<taxonomicName id="92DF4D2EFFBAFF9DFF8CF9E0D990F9A3" ID-CoL="CDB7" box="[125,186,1557,1575]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="5" pageNumber="6" phylum="Annelida" rank="species" species="yirrarn">
<emphasis id="67ABEABFFFBAFF9DFF8CF9E0D990F9A3" box="[125,186,1557,1575]" italics="true" pageId="5" pageNumber="6">yirrarn</emphasis>
</taxonomicName>
mm wide squared short, rectangular, rectangular to acicular tapered spines 21–23); notopodia) 
<bibRefCitation id="314E4B5CFFBAFF9DFF8CF9C4D820F9C0" author="Hutchings" box="[125,266,1585,1604]" pageId="5" pageNumber="6" refString="Hutchings, P. A. (1997) New species of the family Terebellidae (Polychaeta) from Darwin Harbour Australia. In: Hanley, R. (Ed.), ' Proceedings of the 6 th International Marine Biological Workshop. The Marine Flora and Fauna of Darwin Harbour, Northern Territory'. pp. 133 - 161." type="book chapter" year="1997">Hutchings, 1997</bibRefCitation>
(incomplete) circular button-like pentagonal slightly
</paragraph>
<paragraph id="556036ADFFBAFF9DFA17F9B8DF0BF9E4" blockId="5.[125,1901,564,1632]" box="[1510,1569,1613,1632]" pageId="5" pageNumber="6">shorter</paragraph>
<paragraph id="556036ADFFB9FF9EFF36FF6DDBD1FE98" blockId="6.[151,1436,151,285]" pageId="6" pageNumber="7">
On the other hand, the specialised 
<typeStatus id="8A64880FFFB9FF9EFD91FF6DDBBAFF34" box="[608,656,152,176]" pageId="6" pageNumber="7">type</typeStatus>
of buccal tentacles apparently originates exclusively in mid-dorsal position, from the mid-dorsal process when this is present, or an equivalent position in species in which such a prostomial process is absent (
<figureCitation id="CDE42A28FFB9FF9EFE17FF2ADB01FF7C" box="[486,555,223,248]" captionStart="FIGURE 4" captionStartId="13.[151,250,1348,1370]" captionTargetBox="[158,1431,229,1310]" captionTargetId="figure@13.[152,1435,225,1319]" captionTargetPageId="13" captionText="FIGURE 4. Amaeana breviachaeta sp. nov., holotype AM W. 46526. A – B, incomplete worm, right ventro-lateral and left dorso-lateral views, respectively. C – D, anterior end, right ventro-lateral and left dorso-lateral views, respectively; arrows point to last notopodium. E – F, close ups of the anterior end, left dorso-lateral and ventral views, respectively. G, transition between thorax and abdomen; upper arrow points to last notopodium, lower arrow points to first neuropodium. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.4 mm; C – D, G = 0.3 mm; E – F = 0.2 mm." httpUri="https://zenodo.org/record/289141/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
B, D–F). However, other 
<typeStatus id="8A64880FFFB9FF9EFCBDFF15DAADFF7C" box="[844,903,224,248]" pageId="6" pageNumber="7">types</typeStatus>
of tentacles also originate from the mid-dorsal process, not only the specialised 
<typeStatus id="8A64880FFFB9FF9EFDF6FEF0DB1DFE99" box="[519,567,261,285]" pageId="6" pageNumber="7">type</typeStatus>
(see 
<figureCitation id="CDE42A28FFB9FF9EFD82FEF1DB9EFE98" box="[627,692,260,285]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
A, G).
</paragraph>
<paragraph id="556036ADFFB9FF9EFF66FEB9DDBEFD9C" blockId="6.[151,1437,332,861]" pageId="6" pageNumber="7">
(2) the morphology of the long buccal tentacles varies: they may be with highly specialised tips, or only distally expanded, spatulate, without further modifications, similar to those of other polycirrids. However, buccal tentacles are frequently missing in 
<typeStatus id="8A64880FFFB9FF9EFE3BFE60D8D0FE29" box="[458,506,405,429]" pageId="6" pageNumber="7">type</typeStatus>
material and the specialised tentacles are usually much fewer in number than remaining ones. For that reason, the absence of this 
<typeStatus id="8A64880FFFB9FF9EFCF6FE4DDA1DFE54" box="[775,823,440,464]" pageId="6" pageNumber="7">type</typeStatus>
of tentacle in some specimens does not necessarily indicate that they were not originally present; however, in the cases in which many buccal tentacles are present with no specialised ones (
<figureCitation id="CDE42A28FFB9FF9EFE46FE0AD8D1FD9C" box="[439,507,511,536]" captionStart="FIGURE 2" captionStartId="10.[151,250,1856,1878]" captionTargetBox="[212,1379,181,1835]" captionTargetId="figure@10.[206,1382,177,1840]" captionTargetPageId="10" captionText="FIGURE 2. Amaeana trilobata, NMO C 3208: A – C, anterior end, dorsal, right lateral and ventral views, respectively; arrow in C points to first neuropodium. D, mid-abdominal segments, left lateral view. E, buccal tentacles. F, anterior end, right lateral view; arrows point to nephridial / genital papillae. G – I, close ups of the anterior end, ventral, right lateral and dorsal views, respectively. J, mid-abdominal segments, right lateral view; arrows point to neuropodia with protruding spines. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.8 mm; B, E – F = 0.5 mm; C = 0.7 mm; D, H, J = 0.4 mm; G = 0.3 mm; I = 0.2 mm." httpUri="https://zenodo.org/record/289139/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
E), we have assumed this 
<typeStatus id="8A64880FFFB9FF9EFCEDFDF5DA66FD9C" box="[796,844,512,536]" pageId="6" pageNumber="7">type</typeStatus>
of buccal tentacles is absent.
</paragraph>
<paragraph id="556036ADFFB9FF9EFF36FDD1DAEAFCD9" blockId="6.[151,1437,332,861]" pageId="6" pageNumber="7">
The specialised buccal tentacles of some species of 
<taxonomicName id="92DF4D2EFFB9FF9EFCEEFDD1DAA3FDBF" ID-CoL="84LBD" box="[799,905,548,571]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFB9FF9EFCEEFDD1DAA3FDBF" box="[799,905,548,571]" italics="true" pageId="6" pageNumber="7">Amaeana</emphasis>
</taxonomicName>
are quite remarkable. Although we have only examined the buccal tentacles of 
<taxonomicName id="92DF4D2EFFB9FF9EFDFFFDBDDB9FFDE4" ID-CoL="CDB5" box="[526,693,584,608]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="occidentalis">
<emphasis id="67ABEABFFFB9FF9EFDFFFDBDDB9FFDE4" box="[526,693,584,608]" italics="true" pageId="6" pageNumber="7">A. occidentalis</emphasis>
</taxonomicName>
and 
<taxonomicName id="92DF4D2EFFB9FF9EFD1CFDBDDA48FDE4" ID-CoL="CDB4" box="[749,866,584,608]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
<emphasis id="67ABEABFFFB9FF9EFD1CFDBDDA48FDE4" box="[749,866,584,608]" italics="true" pageId="6" pageNumber="7">A. hsiehae</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFB9FF9EFC9AFDB2DAEFFDE4" bold="true" box="[875,965,583,608]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="7C9857C4FFB9FF9EFC9AFDB2DAEFFDE4" box="[875,965,583,608]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
under the SEM, the tentacles of the other species in this genus look similar to those, at least under stereomicroscope. In 
<taxonomicName id="92DF4D2EFFB9FF9EFBF6FD99DD84FD00" ID-CoL="CDB5" box="[1031,1198,620,644]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="occidentalis">
<emphasis id="67ABEABFFFB9FF9EFBF6FD99DD84FD00" box="[1031,1198,620,644]" italics="true" pageId="6" pageNumber="7">A. occidentalis</emphasis>
</taxonomicName>
all 
<typeStatus id="8A64880FFFB9FF9EFB28FD98DC3EFD01" box="[1241,1300,621,645]" pageId="6" pageNumber="7">types</typeStatus>
of tentacles are thin, with an uniformly wide peduncle and specialised tip, from elliptical or spherical, to spatulate or more elaborate, with large subdistal expansion and tapered tip; a ciliated groove is only present distally, the peduncle being smooth or nearly so (
<figureCitation id="CDE42A28FFB9FF9EFE3BFD22DB0BFD74" box="[458,545,727,752]" captionStart="FIGURE 20" captionStartId="36.[151,250,1899,1921]" captionTargetBox="[199,1394,178,1870]" captionTargetId="figure@36.[193,1399,173,1878]" captionTargetPageId="36" captionText="FIGURE 20. Amaeana occidentalis LACM-AHF Poly 6518. A – B, thorax, right lateral and left dorso-lateral views. C, anterior end, left dorso-lateral view. D – E, progressively closer ventral views of the thorax, ventral views. F – G, thorax, dorsal and left lateral views. H, segments 10 – 15, left lateral view; arrows point to genital papillae. I, posterior end, left ventro-lateral view. Scale bars: A – D, F = 0.7 mm; E, G = 0.4 mm; H = 0.3 mm; I = 0.15 mm." httpUri="https://zenodo.org/record/289157/files/figure.png" pageId="6" pageNumber="7">Figs 20</figureCitation>
A–G; 21A–G); the specimen of 
<taxonomicName id="92DF4D2EFFB9FF9EFC7BFD2DDAD4FD74" ID-CoL="CDB4" box="[906,1022,728,752]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
<emphasis id="67ABEABFFFB9FF9EFC7BFD2DDAD4FD74" box="[906,1022,728,752]" italics="true" pageId="6" pageNumber="7">A. hsiehae</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFB9FF9EFBF7FD22DD4AFD74" bold="true" box="[1030,1120,727,752]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="7C9857C4FFB9FF9EFBF7FD22DD4AFD74" box="[1030,1120,727,752]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
examined has few tentacles remaining, but exhibiting a similar pattern. This means that these worms most likely insert the tips of the tentacles inside the mouth, or brush them against the highly ciliated upper lip to remove food particles gathered from the substrate, rather than conveying those particles along the entire tentacle.
</paragraph>
<paragraph id="556036ADFFB9FF9EFF66FC79DD35FB40" blockId="6.[151,1436,908,1221]" pageId="6" pageNumber="7">
(3) the number of pairs of nephridial/genital papillae and notopodia: nephridial/genital papillae occur at the anterior bases of all notopodia, from segment 3 to last one, nephridial papillae on segments 3–5, genital papillae from segment 6 to last thoracic chaetiger (
<bibRefCitation id="314E4B5CFFB9FF9EFDC0FC21DA36FC68" author="Nogueira" box="[561,796,980,1005]" pageId="6" pageNumber="7" refString="Nogueira, J. M. M., Hutchings, P. A. &amp; Fukuda, M. V. (2010) Morphology of terebelliform polychaetes (Annelida: Polychaeta: Terebelliformia), with a focus on Terebellidae. Zootaxa, 2460, 1 - 185." type="journal article" year="2010">
Nogueira 
<emphasis id="67ABEABFFFB9FF9EFD53FC20DBF9FC68" box="[674,723,980,1004]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2010
</bibRefCitation>
, 
<bibRefCitation id="314E4B5CFFB9FF9EFCD6FC21DA4EFC68" author="Nogueira" box="[807,868,980,1004]" pageId="6" pageNumber="7" refString="Nogueira, J. M. M., Fitzhugh, K. &amp; Hutchings, P. (2013) The continuing challenge of phylogenetic relationships in Terebelliformia (Annelida: Polychaeta). Invertebrate Systematics, 27, 186 - 238. http: // dx. doi. org / 10.1071 / IS 12062" type="journal article" year="2013">2013</bibRefCitation>
). There is a strong correlation between the number of pairs of papillae and the pairs of notopodia, so the variation found in the number of pairs of notopodia present is also observed in regards to the number of pairs of nephridial/genital papillae. However, sometimes papillae of anterior- and/or posterior-most thoracic chaetigers are distinctly shorter than remaining ones (
<figureCitation id="CDE42A28FFB9FF9EFB35FBCADC36FBDC" box="[1220,1308,1087,1112]" captionStart="FIGURE 20" captionStartId="36.[151,250,1899,1921]" captionTargetBox="[199,1394,178,1870]" captionTargetId="figure@36.[193,1399,173,1878]" captionTargetPageId="36" captionText="FIGURE 20. Amaeana occidentalis LACM-AHF Poly 6518. A – B, thorax, right lateral and left dorso-lateral views. C, anterior end, left dorso-lateral view. D – E, progressively closer ventral views of the thorax, ventral views. F – G, thorax, dorsal and left lateral views. H, segments 10 – 15, left lateral view; arrows point to genital papillae. I, posterior end, left ventro-lateral view. Scale bars: A – D, F = 0.7 mm; E, G = 0.4 mm; H = 0.3 mm; I = 0.15 mm." httpUri="https://zenodo.org/record/289157/files/figure.png" pageId="6" pageNumber="7">Figs 20</figureCitation>
G–H; 22G) and they may be difficult to distinguish from the numerous glandular papillae which cover the ventral surface of ventro-lateral pads. On the other hand, in some species, genital papillae of some mid-thoracic segments are distinctly larger than those of remaining segments (
<figureCitation id="CDE42A28FFB9FF9EFD24FB59DA31FB41" box="[725,795,1196,1221]" captionStart="FIGURE 5" captionStartId="15.[151,250,1791,1813]" captionTargetBox="[236,1361,191,1760]" captionTargetId="figure@15.[230,1364,187,1773]" captionTargetPageId="15" captionText="FIGURE 5. Amaeana hsiehae sp. nov., ASIZW 0000935. A – D, entire worm, right ventro-lateral, right lateral, left lateral and left dorso-lateral views, respectively; arrows point to beginning of neuropodia; E – F, progressively closer views of the anterior end, ventral view; G – H, thorax, right and left lateral views, respectively; I – K, close ups of the anterior end, left dorso-lateral, ventral and right lateral views, respectively; L, posterior end, left dorso-lateral view; M, close up of the anterior end, left dorsolateral view; N, abdominal segments; arrows point to neuropodia; O, posterior end, right ventro-lateral view. Paratype AM W. 47363: P, entire worm, left lateral view (anteriorly); Q, thorax, left lateral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip. Scale bars: A = 0.6 mm; B – D, H = 0.5 mm; E, G = 0.4 mm; F, I – L, N = 0.2 mm; M, O = 0.15 mm; P = 1.5 mm; Q = 1 mm." httpUri="https://zenodo.org/record/289142/files/figure.png" pageId="6" pageNumber="7">Figs 5</figureCitation>
P–Q; 19A, C; 20G–H).
</paragraph>
<paragraph id="556036ADFFB9FF9EFF66FB01DA48FA44" blockId="6.[151,1436,1268,1472]" pageId="6" pageNumber="7">
(4) the number of segments with ventro-lateral glandular pads: this character is also strongly correlated with the number of pairs of notopodia, as the paired glandular ventro-lateral pads usually extend from segment 2 to the last with notopodia, frequently with fewer papillae, restricted to the posterior annulation of each segment, on last chaetigers with notopodia. The only species of 
<taxonomicName id="92DF4D2EFFB9FF9EFD54FA95DA25FAF3" ID-CoL="84LBD" box="[677,783,1376,1399]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFB9FF9EFD54FA95DA25FAF3" box="[677,783,1376,1399]" italics="true" pageId="6" pageNumber="7">Amaeana</emphasis>
</taxonomicName>
with ventral pads extending beyond the termination of the notopodia is 
<taxonomicName id="92DF4D2EFFB9FF9EFEC3FA71D885FA18" ID-CoL="65XLB" box="[306,431,1412,1436]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="angulus" status="sp. nov.">
<emphasis id="67ABEABFFFB9FF9EFEC3FA71D885FA18" box="[306,431,1412,1436]" italics="true" pageId="6" pageNumber="7">A. angulus</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFB9FF9EFE4DFA71DB30FA19" bold="true" box="[444,538,1412,1437]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="7C9857C4FFB9FF9EFE4DFA71DB30FA19" box="[444,538,1412,1437]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which has notopodia until segment 12 and pads extending to segment 15, although the last three pairs are distinctly smaller (
<figureCitation id="CDE42A28FFB9FF9EFD3FFA52DA34FA44" box="[718,798,1447,1472]" captionStart="FIGURE 14" captionStartId="27.[151,250,1842,1864]" captionTargetBox="[181,1407,185,1808]" captionTargetId="figure@27.[174,1411,181,1815]" captionTargetPageId="27" captionText="FIGURE 14. Amaeana angulus sp. nov., holotype AM W. 13640. A, entire worm, left lateral view (anteriorly). B – C, anterior end, dorsal and ventral views, respectively. D, buccal tentacles. E – F, anterior end, left and right lateral views, respectively. G – I, close ups of the anterior end, dorsal, left ventro-lateral and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A = 1.5 mm; B – C, F = 0.7 mm; D = 0.5 mm; E = 1 mm; G – I = 0.5 mm." httpUri="https://zenodo.org/record/289151/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
A, E).
</paragraph>
<paragraph id="556036ADFFB9FF9EFF66FA1AD9F2F8AC" blockId="6.[151,1436,1519,1832]" pageId="6" pageNumber="7">
(5) the 
<typeStatus id="8A64880FFFB9FF9EFF18FA04D80EF98D" box="[233,292,1521,1545]" pageId="6" pageNumber="7">types</typeStatus>
of notochaetae present: most species of 
<taxonomicName id="92DF4D2EFFB9FF9EFD1FFA05DA72F983" ID-CoL="84LBD" box="[750,856,1520,1543]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFB9FF9EFD1FFA05DA72F983" box="[750,856,1520,1543]" italics="true" pageId="6" pageNumber="7">Amaeana</emphasis>
</taxonomicName>
have narrowly-winged notochaetae 
<emphasis id="67ABEABFFFB9FF9EFB03FA04DC1AF98C" box="[1266,1328,1521,1544]" italics="true" pageId="6" pageNumber="7">sensu</emphasis>
<bibRefCitation id="314E4B5CFFB9FF9EFAC9FA1AD80CF9A9" pageId="6" pageNumber="7" refString="Fitzhugh, K., Nogueira, J. M. M., Carrerette, O. &amp; Hutchings, P. (2015) An assessment of the status of Polycirridae genera (Annelida: Terebelliformia) with the evolutionary transformation series of characters within the family. Zoology Journal of Linnean Society, 2015, 1 - 36. [online only]. http: // dx. doi. org / 10.1111 / zoj. 12259" type="journal article">
Fitzhugh 
<emphasis id="67ABEABFFFB9FF9EFF66F9E0D9F8F9A8" box="[151,210,1556,1580]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(2015)
</bibRefCitation>
in both rows; those chaetae are acicular, with inconspicuous wings under higher magnifications of light microscopy, but visible as fine hairs under SEM. Among members of two species, however, pinnate chaetae are present, in the posterior row of notochaetae in 
<taxonomicName id="92DF4D2EFFB9FF9EFD23F9A9DA40F9F0" ID-CoL="5TXLB" box="[722,874,1628,1652]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="accraensis">
<emphasis id="67ABEABFFFB9FF9EFD23F9A9DA40F9F0" box="[722,874,1628,1652]" italics="true" pageId="6" pageNumber="7">A. accraensis</emphasis>
</taxonomicName>
(
<figureCitation id="CDE42A28FFB9FF9EFC8BF9A9DAE6F9F0" box="[890,972,1628,1653]" captionStart="FIGURE 27" captionStartId="45.[151,250,757,779]" captionTargetBox="[158,1431,197,730]" captionTargetId="figure@45.[152,1435,193,736]" captionTargetPageId="45" captionText="FIGURE 27. Amaeana accraensis holotype ZMH V- 1710. A, notochaetae, segment 8; B, notochaetae from anterior row, segment 8; C, tips of notochaetae from posterior row, segment 8; D, neuropodial spines, segment 42. Scale bars: A = 30 µm; B – D = 10 µm." httpUri="https://zenodo.org/record/289164/files/figure.png" pageId="6" pageNumber="7">Fig. 27</figureCitation>
A–C), and in both rows in 
<taxonomicName id="92DF4D2EFFB9FF9EFB0CF9A9DC59F9F0" ID-CoL="CDB4" box="[1277,1395,1628,1652]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
<emphasis id="67ABEABFFFB9FF9EFB0CF9A9DC59F9F0" box="[1277,1395,1628,1652]" italics="true" pageId="6" pageNumber="7">A. hsiehae</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFB9FF9EFA8BF9A9D9E2F91C" bold="true" pageId="6" pageNumber="7">
<taxonomicNameLabel id="7C9857C4FFB9FF9EFA8BF9A9D9E2F91C" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="CDE42A28FFB9FF9EFF2DF98AD80EF91C" box="[220,292,1663,1688]" captionStart="FIGURE 6" captionStartId="16.[151,250,1269,1291]" captionTargetBox="[158,1430,197,1238]" captionTargetId="figure@16.[152,1435,193,1247]" captionTargetPageId="16" captionText="FIGURE 6. Amaeana hsiehae sp. nov., holotype ASIZW 0000935. A – B, progressively closer views of the notochaetae of segment 6; C – E, neurochaetae, segments 21 and two consecutive posterior ones, respectively. Scale bars: A = 30 µm; B = 15 µm; C – E = 7 µm." httpUri="https://zenodo.org/record/289143/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
A–B; 8B–F). Some species, such as 
<taxonomicName id="92DF4D2EFFB9FF9EFD2BF975DA7EF91C" ID-CoL="5TXMF" box="[730,852,1664,1688]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="apheles">
<emphasis id="67ABEABFFFB9FF9EFD2BF975DA7EF91C" box="[730,852,1664,1688]" italics="true" pageId="6" pageNumber="7">A. apheles</emphasis>
</taxonomicName>
, have notochaetae that strongly resemble pinnate chaetae under SEM, but this is not visible under higher magnifications of light microscopy (compare 
<figureCitation id="CDE42A28FFB9FF9EFADAF951DC59F938" box="[1323,1395,1700,1725]" captionStart="FIGURE 9" captionStartId="20.[151,250,1851,1873]" captionTargetBox="[193,1404,197,1818]" captionTargetId="figure@20.[187,1408,193,1826]" captionTargetPageId="20" captionText="FIGURE 9. Amaeana apheles, holotype AM W. 5239. A – C, anterior end, dorsal, right ventro-lateral, and ventral views, respectively. D – F, closer views of the anterior end, left and right ventro-lateral, and ventral views, respectively; arrows point to nephridial / genital papillae. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Specimen AM W. 29203: I – J, notochaetae, segment 9. K, neurochaetae, posterior segment. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; P * = distal part of prostomium. Scale bars: A = 0.7 mm; B – C = 1 mm; D – F = 0.5 mm; G – H = 0.3 mm; I – K = 20 µm." httpUri="https://zenodo.org/record/289146/files/figure.png" pageId="6" pageNumber="7">
Figs 
<date id="2161106DFFB9FF9EFA94F951DC59F938" box="[1381,1395,1700,1724]" pageId="6" pageNumber="7">9</date>
</figureCitation>
I–J, 11A–E, to see those chaetae under light microscopy and SEM, respectively), and so these chaetae are also treated here as acicular notochaetae, because it is not practical to diagnose a species by characters only visible under the SEM.
</paragraph>
<paragraph id="556036ADFFB9FF9EFF66F8A2DC75F83D" blockId="6.[151,1437,1879,1977]" pageId="6" pageNumber="7">(6) the extension of the achaetous gap between termination of notopodia and beginning of neuropodia: this region is often swollen, fragile, with segments distinctly longer than those with notopodia and with thinner body wall. Segmentation is poorly marked in this area, frequently rendering it difficult to count the number of segments.</paragraph>
<paragraph id="556036ADFFB8FF9FFF36FF6DD825FE29" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="8">However, the whole concept of segmentation is challenged here, because terebelliforms have no internal septa other than the gular membrane, between segments 4 and 5, and if segmentation is also not clear externally, then the counting of “segments” between termination of notopodia and beginning of neuropodia may be a misconception. It seems more reasonable to measure this region in relation to the length of the region with notopodia, and frequently it is at least as long as that part of the body, but this proportion is also strongly dependent on the state of preservation of specimens, together with muscular contraction at the time of fixation. Both measurements are provided herein for each species, the number of achaetous “segments” and the length relative to the region with notopodia.</paragraph>
<paragraph id="556036ADFFB8FF9FFF36FE42D9C5FD9C" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="8">Posteriorly, this region usually tapers abruptly to an uniformly cylindrical abdomen, and neuropodia first occur somewhere among these tapering segments and the first ones of the uniformly cylindrical region, depending on the species.</paragraph>
<paragraph id="556036ADFFB8FF9FFF66FDBDDD44FC21" blockId="7.[151,1437,583,1509]" pageId="7" pageNumber="8">
(7) the number of spines per neuropodium and their morphology: considering we have worked with 
<typeStatus id="8A64880FFFB8FF9FFAF9FDBDDC12FDE4" box="[1288,1336,584,608]" pageId="7" pageNumber="8">type</typeStatus>
material for the present study, we could not prepare slides of several neuropodia of our specimens, to determine the variation in the number of neuropodial spines per neuropodium. Instead, except for some of the species herein described, we mounted two slides per specimen, one from an anterior neuropodium, a few segments after the beginning of neuropodia, and another from a posterior segment; in other cases we mounted a single slide, usually from a mid-abdominal segment. However, only 
<taxonomicName id="92DF4D2EFFB8FF9FFD3EFD09DA51FC90" ID-CoL="CDB5" box="[719,891,764,788]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="occidentalis">
<emphasis id="67ABEABFFFB8FF9FFD3EFD09DA51FC90" box="[719,891,764,788]" italics="true" pageId="7" pageNumber="8">A. occidentalis</emphasis>
</taxonomicName>
and 
<taxonomicName id="92DF4D2EFFB8FF9FFC4AFD09DD42FC90" ID-CoL="CDB3" box="[955,1128,764,788]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="ellobophora" status="sp. nov.">
<emphasis id="67ABEABFFFB8FF9FFC4AFD09DD42FC90" box="[955,1128,764,788]" italics="true" pageId="7" pageNumber="8">A. ellobophora</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFB8FF9FFB85FD09DDF9FC91" bold="true" box="[1140,1235,764,789]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="7C9857C4FFB8FF9FFB85FD09DDF9FC91" box="[1140,1235,764,789]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
exhibit variation greater than 1–2 spines per neuropodium between anterior and posterior abdomen. There was little to no variation between members of the same species in cases where we were able to study more than one specimen of each taxon. Therefore, we believe a great variation on the number of spines per neuropodium, as reported sometimes in the literature (see below), is possibly due to the presence of several species being confused.
</paragraph>
<paragraph id="556036ADFFB8FF9FFF36FC5ADB82FA88" blockId="7.[151,1437,583,1509]" pageId="7" pageNumber="8">
When more than 3 spines are present per neuropodium, these usually have uneven lengths and only 1–2 spines protrude from each neuropodium (for instance, compare 
<figureCitation id="CDE42A28FFB8FF9FFCC4FC21DAA0FC69" box="[821,906,980,1005]" captionStart="FIGURE 15" captionStartId="29.[151,250,1777,1799]" captionTargetBox="[204,1385,197,1750]" captionTargetId="figure@29.[198,1388,193,1756]" captionTargetPageId="29" captionText="FIGURE 15. Amaeana ellobophora sp. nov., holotype AM W. 7055. A – B, entire worm, ventral and right lateral views (anteriorly), respectively; arrows point to first neuropodium. C – E, anterior end, dorsal, ventral and right dorso-lateral views, respectively. F, posterior end of fragment. G – J, close ups of the anterior end, dorsal, left lateral, ventral and right lateral views, respectively; arrows point to nephridial / genital papillae. K, close up of the posterior end of fragment. L, notochaetae, segment 10; M, anterior neurochaetae. N – O, progressively closer views of the neurochaetae of a posterior neuropodium. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = mid-dorsal prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 1 mm; C – E = 0.4 mm; F = 0.5 mm, G – I, K = 0.2 mm; J = 0.3 mm; L = 50 µm; M = 40 µm; N = 30 µm; O = 10 µm." httpUri="https://zenodo.org/record/289152/files/figure.png" pageId="7" pageNumber="8">Figs 15</figureCitation>
M–O and 17C–D, showing neuropodia of 
<taxonomicName id="92DF4D2EFFB8FF9FFA75FC21D80AFB94" ID-CoL="CDB3" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="ellobophora" status="sp. nov.">
<emphasis id="67ABEABFFFB8FF9FFA75FC21D80AFB94" italics="true" pageId="7" pageNumber="8">A. ellobophora</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7C9857C4FFB8FF9FFED6FC02D8ABFB8B" box="[295,385,1015,1040]" pageId="7" pageNumber="8" rank="species">
<emphasis id="67ABEABFFFB8FF9FFED6FC02D850FB94" bold="true" box="[295,378,1015,1040]" pageId="7" pageNumber="8">sp. nov</emphasis>
.
</taxonomicNameLabel>
under light microscopy and SEM, respectively). With the exception of 
<taxonomicName id="92DF4D2EFFB8FF9FFB59FC0DDC59FB94" ID-CoL="5TXMG" box="[1192,1395,1016,1040]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="crassispinulata" status="sp. nov.">
<emphasis id="67ABEABFFFB8FF9FFB59FC0DDC59FB94" box="[1192,1395,1016,1040]" italics="true" pageId="7" pageNumber="8">A. crassispinulata</emphasis>
</taxonomicName>
<emphasis id="67ABEABFFFB8FF9FFA8BFC02D9E2FBB1" bold="true" pageId="7" pageNumber="8">
<taxonomicNameLabel id="7C9857C4FFB8FF9FFA8BFC02D9E2FBB1" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the neuropodial spines are straight and distally tapered, but frequently the spines are further modified to a narrower knob-like or slightly hooked tip, sometimes flattened and concave, spoon-shaped (
<figureCitation id="CDE42A28FFB8FF9FFB50FBCADDC2FBDC" box="[1185,1256,1087,1112]" captionStart="FIGURE 3" captionStartId="11.[151,250,1763,1785]" captionTargetBox="[158,1431,197,1727]" captionTargetId="figure@11.[152,1435,193,1741]" captionTargetPageId="11" captionText="FIGURE 3. Chaetae. Amaeana trilobata NMO C 3208: A, notochaetae, segment 5. B, neurochaetae, segment 45. Amaeana breviachaeta sp. nov., holotype AM W. 46526: C, notochaetae, segment 8. D – E, neurochaetae, segments 21 and 27, respectively. Amaeana angulus sp. nov., holotype AM W. 13640: F – G, notochaetae, segment 3. H, neurochaetae, segment 25. Scale bars: A, G – H = 80 µm; B, D – E = 10 µm; C = 20 µm; F = 150 µm." httpUri="https://zenodo.org/record/289140/files/figure.png" pageId="7" pageNumber="8">Figs 3</figureCitation>
B, D–E, H; 6C– E; 
<date id="2161106DFFB8FF9FFF47FB91D9E7FBF8" box="[182,205,1124,1148]" pageId="7" pageNumber="8">8I</date>
–L; 11H–I; 19G–H; 24C, E, H–I; 27D). This is difficult to clearly visualize because it depends critically on the alignment of the spines on slides; further, in compound microscopes the highest magnification does not allow for a proper analysis of this character, while under the SEM the tips of the spines are often not visible, as these structures do not always protrude from the neuropodial pinnules, and when they do, frequently only a short part of the tip is exposed (
<figureCitation id="CDE42A28FFB8FF9FFEF3FB01D87EFA88" box="[258,340,1268,1293]" captionStart="FIGURE 11" captionStartId="22.[151,250,1897,1919]" captionTargetBox="[171,1413,196,1851]" captionTargetId="figure@22.[164,1417,192,1859]" captionTargetPageId="22" captionText="FIGURE 11. Amaeana apheles, photos A – E from specimen AM W. 19333, photos F – I from specimen AM W. 10864. A, notopodia, segments 9 – 10; B – E, notochaetae, segment 10; F – G, posterior neuropodia; H – I, close up of posterior neuropodia with protruding spines. Scale bars: A = 70 µm; B, I = 10 µm; C – D = 5 µm; E = 4 µm; F = 200 µm; G = 8 µm; H = 20 µm." httpUri="https://zenodo.org/record/289148/files/figure.png" pageId="7" pageNumber="8">Figs 11</figureCitation>
F–I; 17C–D; 22A–F; 29 H–J).
</paragraph>
<paragraph id="556036ADFFB8FF9FFF36FAE2D844FA61" blockId="7.[151,1437,583,1509]" pageId="7" pageNumber="8">
Other characters also useful to distinguish between the species of 
<taxonomicName id="92DF4D2EFFB8FF9FFC5FFAEDDD32FAAB" ID-CoL="84LBD" box="[942,1048,1304,1327]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="genus">
<emphasis id="67ABEABFFFB8FF9FFC5FFAEDDD32FAAB" box="[942,1048,1304,1327]" italics="true" pageId="7" pageNumber="8">Amaeana</emphasis>
</taxonomicName>
are the shapes of upper and lower lips, and of the mid-ventral shield of segment 2. The upper lip is usually circular, but it may be elliptical, higher than wide, as in 
<taxonomicName id="92DF4D2EFFB8FF9FFEBDFA95D8C6FAFC" ID-CoL="65XMF" box="[332,492,1376,1400]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="67ABEABFFFB8FF9FFEBDFA95D8C6FAFC" box="[332,492,1376,1400]" italics="true" pageId="7" pageNumber="8">A. brasiliensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7C9857C4FFB8FF9FFE03FAAADB61FAFC" box="[498,587,1375,1400]" pageId="7" pageNumber="8" rank="species">
<emphasis id="67ABEABFFFB8FF9FFE03FAAADB6FFAFC" bold="true" box="[498,581,1375,1400]" pageId="7" pageNumber="8">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="CDE42A28FFB8FF9FFDA8FAAADB85FAFC" box="[601,687,1375,1400]" captionStart="FIGURE 28" captionStartId="47.[151,250,1914,1936]" captionTargetBox="[155,1429,197,1891]" captionTargetId="figure@47.[151,1436,193,1892]" captionTargetPageId="47" captionText="FIGURE 28. Amaeana brasiliensis sp. nov. holotype MZUSP 2349. A – C, thorax, ventral, dorsal and left lateral views, respectively. D – E, close ups of the anterior end, ventral and dorsal views, respectively. F, anterior end, right lateral view. G, buccal tentacles. H, posterior end, right dorso-lateral view. Numbers refer to segments; li = lower lip; P = basal part of prostomium; PP = prostomial process; ls = upper lip. Scale bars: A – C, F, H = 1 mm; D – E, G = 0.5 mm." httpUri="https://zenodo.org/record/289165/files/figure.png" pageId="7" pageNumber="8">Figs 28</figureCitation>
A, C–D, F; 29B–C, E–F) or almost rectangular, as in 
<taxonomicName id="92DF4D2EFFB8FF9FFAF1FA95DC59FAFC" ID-CoL="CDB4" box="[1280,1395,1376,1400]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
<emphasis id="67ABEABFFFB8FF9FFAF1FA95DC59FAFC" box="[1280,1395,1376,1400]" italics="true" pageId="7" pageNumber="8">A. hsiehae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7C9857C4FFB8FF9FFA8BFAAAD9EDFA18" pageId="7" pageNumber="8" rank="species">
<emphasis id="67ABEABFFFB8FF9FFA8BFAAAD9EBFA19" bold="true" pageId="7" pageNumber="8">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="CDE42A28FFB8FF9FFF27FA71D836FA19" box="[214,284,1412,1437]" captionStart="FIGURE 5" captionStartId="15.[151,250,1791,1813]" captionTargetBox="[236,1361,191,1760]" captionTargetId="figure@15.[230,1364,187,1773]" captionTargetPageId="15" captionText="FIGURE 5. Amaeana hsiehae sp. nov., ASIZW 0000935. A – D, entire worm, right ventro-lateral, right lateral, left lateral and left dorso-lateral views, respectively; arrows point to beginning of neuropodia; E – F, progressively closer views of the anterior end, ventral view; G – H, thorax, right and left lateral views, respectively; I – K, close ups of the anterior end, left dorso-lateral, ventral and right lateral views, respectively; L, posterior end, left dorso-lateral view; M, close up of the anterior end, left dorsolateral view; N, abdominal segments; arrows point to neuropodia; O, posterior end, right ventro-lateral view. Paratype AM W. 47363: P, entire worm, left lateral view (anteriorly); Q, thorax, left lateral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip. Scale bars: A = 0.6 mm; B – D, H = 0.5 mm; E, G = 0.4 mm; F, I – L, N = 0.2 mm; M, O = 0.15 mm; P = 1.5 mm; Q = 1 mm." httpUri="https://zenodo.org/record/289142/files/figure.png" pageId="7" pageNumber="8">Figs 5</figureCitation>
A, E–F, I–K, M; 7C, G, I). The lower lip and the mid-ventral shield of segment 
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be small or large, swollen, but this is also strongly dependent on the state of preservation of the specimens, with some intraspecific variation observed.
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