357 lines
54 KiB
XML
357 lines
54 KiB
XML
<document id="06EA1929ACC3A8917470C432853917F2" ID-DOI="10.11646/zootaxa.3994.1.1" ID-GBIF-Dataset="1334b429-62a4-4505-9e05-d68b39058e5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="289137" ID-ZooBank="093B124E-58AE-4303-8C07-2D7B27E6AC38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461271356316" checkinUser="plazi" docAuthor="Nogueira, João Miguel De Matos, Carrerette, Orlemir & Hutchings, Pat" docDate="2015" docId="DD7687BBFF9EFFBFFF66FF62D8CEFD74" docLanguage="en" docName="zt03994p052.pdf" docOrigin="Zootaxa 3994 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amaeana occidentalis Hartman 1944" docType="treatment" docVersion="6" lastPageNumber="40" masterDocId="214FFFC3FFBFFF98FFF1FFF5D92AFF84" masterDocTitle="Review of Amaeana Hartman, 1959 (Annelida, Terebelliformia, Polycirridae), with descriptions of seven new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="34" updateTime="1698611839755" updateUser="plazi">
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<mods:title id="4274F82C4727CFED625E431EDC83A4B2">Review of Amaeana Hartman, 1959 (Annelida, Terebelliformia, Polycirridae), with descriptions of seven new species</mods:title>
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<mods:namePart id="2B88A21C42F8FDD583B380D79A99B9B8">Nogueira, João Miguel De Matos</mods:namePart>
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<mods:namePart id="9819F2BF0B919EDD0A3D9D03FBC51242">Carrerette, Orlemir</mods:namePart>
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<mods:namePart id="B570F5489EE134FA2BF676D967B371E4">Hutchings, Pat</mods:namePart>
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<mods:title id="1F4F399C76FB0A8FD0981B1622E60302">Zootaxa</mods:title>
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<mods:date id="62E2F8E93FF9FF9AA10356ECFF030CA0">2015</mods:date>
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<mods:number id="19D7B8EF6F0A4CE6402410BD49556665">3994</mods:number>
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<treatment id="DD7687BBFF9EFFBFFF66FF62D8CEFD74" ID-DOI="http://doi.org/10.5281/zenodo.6094918" ID-GBIF-Taxon="127692612" ID-Zenodo-Dep="6094918" LSID="urn:lsid:plazi:treatment:DD7687BBFF9EFFBFFF66FF62D8CEFD74" httpUri="http://treatment.plazi.org/id/DD7687BBFF9EFFBFFF66FF62D8CEFD74" lastPageId="39" lastPageNumber="40" pageId="33" pageNumber="34">
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<subSubSection id="1DC56526FF9EFFB9FF66FF62D8F4FF57" pageId="33" pageNumber="34" type="nomenclature">
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<paragraph id="556036ADFF9EFFB9FF66FF62DB52FF36" blockId="33.[151,632,151,211]" box="[151,632,151,178]" pageId="33" pageNumber="34">
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<heading id="0E2881C1FF9EFFB9FF66FF62DB52FF36" bold="true" box="[151,632,151,178]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
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<taxonomicName id="92DF4D2EFF9EFFB9FF66FF62DB52FF36" ID-CoL="CDB5" authority="Hartman, 1944" authorityName="Hartman" authorityYear="1944" box="[151,632,151,178]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="occidentalis">
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<emphasis id="67ABEABFFF9EFFB9FF66FF62DB52FF36" bold="true" box="[151,632,151,178]" pageId="33" pageNumber="34">
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<emphasis id="67ABEABFFF9EFFB9FF66FF62D88BFF35" bold="true" box="[151,417,151,177]" italics="true" pageId="33" pageNumber="34">Amaeana occidentalis</emphasis>
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(
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<bibRefCitation id="314E4B5CFF9EFFB9FE41FF6DDB5AFF36" author="Hartman" box="[432,624,152,178]" pageId="33" pageNumber="34" refString="Hartman, O. (1944) Polychaetous Annelids from California including the descriptions of two new genera and nine new species. Allan Hancock Pacific Expedition, 10, 239 - 290." type="journal article" year="1944">Hartman, 1944</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="556036ADFF9EFFB9FF66FF4FD8F4FF57" blockId="33.[151,632,151,211]" box="[151,478,186,211]" pageId="33" pageNumber="34">
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<figureCitation id="CDE42A28FF9EFFB9FF66FF4FD83AFF57" box="[151,272,186,211]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figures 18</figureCitation>
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,
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<figureCitation id="CDE42A28FF9EFFB9FEEDFF4ED810FF57" box="[284,314,187,211]" captionStart="FIGURE 19" captionStartId="35.[151,250,1886,1908]" captionTargetBox="[209,1375,197,1851]" captionTargetId="figure@35.[204,1380,193,1856]" captionTargetPageId="35" captionText="FIGURE 19. Amaeana occidentalis LACM-AHF Poly 6518. A, entire worm, left ventro-lateral view (anteriorly). C, anterior end, left ventro-lateral view. F, posterior end. Specimen LACM-AHF Poly 6517. B, notochaetae, segment 7; arrow point to notochaetae from anterior row. D, notochaetae from anterior row, segment 7. E, tips of notochaetae from posterior row, segment 7. G, mid-abdominal neurochaetae. H, posterior neurochaetae. Scale bars: A = 1 mm; C, F = 0.6 mm; B = 100 µm; D, G = 20 µm; E = 30 µm; H = 10 µm." httpUri="https://zenodo.org/record/289156/files/figure.png" pageId="33" pageNumber="34">19</figureCitation>
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,
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<figureCitation id="CDE42A28FF9EFFB9FEB7FF4ED84EFF57" box="[326,356,187,211]" captionStart="FIGURE 20" captionStartId="36.[151,250,1899,1921]" captionTargetBox="[199,1394,178,1870]" captionTargetId="figure@36.[193,1399,173,1878]" captionTargetPageId="36" captionText="FIGURE 20. Amaeana occidentalis LACM-AHF Poly 6518. A – B, thorax, right lateral and left dorso-lateral views. C, anterior end, left dorso-lateral view. D – E, progressively closer ventral views of the thorax, ventral views. F – G, thorax, dorsal and left lateral views. H, segments 10 – 15, left lateral view; arrows point to genital papillae. I, posterior end, left ventro-lateral view. Scale bars: A – D, F = 0.7 mm; E, G = 0.4 mm; H = 0.3 mm; I = 0.15 mm." httpUri="https://zenodo.org/record/289157/files/figure.png" pageId="33" pageNumber="34">20</figureCitation>
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,
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<figureCitation id="CDE42A28FF9EFFB9FE81FF4ED8A6FF57" box="[368,396,187,211]" captionStart="FIGURE 21" captionStartId="37.[151,250,1950,1972]" captionTargetBox="[181,1407,197,1921]" captionTargetId="figure@37.[174,1411,193,1928]" captionTargetPageId="37" captionText="FIGURE 21. Amaeana occidentalis LACM-AHF Poly 6518. A – F, tips of buccal tentacles. G, close up view of the anterior end, ventral view. H, close up of the posterior end, frontal view. Numbers refer to segments; ll = lower lip; P = basal part of prostomium; ul = upper lip. Scale bars: A, G = 0.2 mm; B, D, F = 0.1 mm; C = 0.15 mm; E = 40 µm; H = 50 µm." httpUri="https://zenodo.org/record/289158/files/figure.png" pageId="33" pageNumber="34">21</figureCitation>
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and
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<figureCitation id="CDE42A28FF9EFFB9FE33FF4ED8F4FF57" box="[450,478,187,211]" captionStart="FIGURE 22" captionStartId="38.[151,250,1914,1936]" captionTargetBox="[180,1422,197,1876]" captionTargetId="figure@38.[174,1427,193,1888]" captionTargetPageId="38" captionText="FIGURE 22. Amaeana occidentalis LACM-AHF Poly 6518. A, posterior end, ventral view (anteriorly). B, abdominal segments. C – F, close ups of abdominal neuropodia, with protruding spines. G, notopodia, segments 3 – 5; arrows point to nephridial papillae. H – I, progressively closer views of the notochaetae of segment 5. Scale bars: A = 0.5 mm; B = 0.15 mm; C = 10 µm; D = 3 µm; E = 5 µm; F = 2 µm; G = 70 µm; H – I = 20 µm." httpUri="https://zenodo.org/record/289159/files/figure.png" pageId="33" pageNumber="34">22</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="1DC56526FF9EFFB9FF66FEF6DA8DFEB3" pageId="33" pageNumber="34" type="reference_group">
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<paragraph id="556036ADFF9EFFB9FF66FEF6DA08FE9C" blockId="33.[151,935,258,311]" box="[151,802,258,280]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="75CF1183FF9EFFB9FF66FEF6DA08FE9C" box="[151,802,258,280]" pageId="33" pageNumber="34">
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<taxonomicName id="92DF4D2EFF9EFFB9FF66FEF6DB00FE9C" authority="Hartman 1944: 277" authorityName="Hartman" authorityPageNumber="277" authorityYear="1944" box="[151,554,258,280]" class="Gastropoda" family="Epitoniidae" genus="Amaea" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="occidentalis">
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<emphasis id="67ABEABFFF9EFFB9FF66FEF6D874FE9C" box="[151,350,258,280]" italics="true" pageId="33" pageNumber="34">Amaea occidentalis</emphasis>
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<treatmentCitation id="D47E10BCFF9EFFB9FE95FEF6DB00FE9C" author="Hartman" box="[356,554,258,280]" page="277" pageId="33" pageNumber="34" year="1944">
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<bibRefCitation id="314E4B5CFF9EFFB9FE95FEF6D8D3FE9C" author="Hartman" box="[356,505,258,280]" pageId="33" pageNumber="34" refString="Hartman, O. (1944) Polychaetous Annelids from California including the descriptions of two new genera and nine new species. Allan Hancock Pacific Expedition, 10, 239 - 290." type="journal article" year="1944">Hartman 1944</bibRefCitation>
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: 277
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</treatmentCitation>
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</taxonomicName>
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–278, Pl. 26, figs 68, 69.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="556036ADFF9EFFB9FF66FED7DA8DFEB3" blockId="33.[151,935,258,311]" box="[151,935,289,311]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="75CF1183FF9EFFB9FF66FED7DA8DFEB3" box="[151,935,289,311]" pageId="33" pageNumber="34">
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<taxonomicName id="92DF4D2EFF9EFFB9FF66FED7D85DFEB3" ID-CoL="CDB5" box="[151,375,289,311]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="occidentalis">
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<emphasis id="67ABEABFFF9EFFB9FF66FED7D85DFEB3" box="[151,375,289,311]" italics="true" pageId="33" pageNumber="34">Amaeana occidentalis</emphasis>
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</taxonomicName>
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.
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<treatmentCitation id="D47E10BCFF9EFFB9FE75FED7DB61FEB3" author="Hartman" box="[388,587,289,311]" page="495" pageId="33" pageNumber="34" year="1959">
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||
<bibRefCitation id="314E4B5CFF9EFFB9FE75FED7DB32FEB3" author="Hartman" box="[388,536,289,311]" pageId="33" pageNumber="34" refString="Hartman, O. (1959) Catalogue of the Polychaetous annelids of the world. Part 2. Sedentaria. Allan Hancock Foundation Occasional Paper, 23, 355 - 628." type="journal article" year="1959">Hartman 1959</bibRefCitation>
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||
: 495
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||
</treatmentCitation>
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||
.
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<emphasis id="67ABEABFFF9EFFB9FDBEFED7DB4FFEB3" box="[591,613,290,311]" italics="true" pageId="33" pageNumber="34">—</emphasis>
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<treatmentCitation id="D47E10BCFF9EFFB9FD94FED4DA38FEB3" author="Hilbig" box="[613,786,289,311]" page="238" pageId="33" pageNumber="34" year="2000">
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<bibRefCitation id="314E4B5CFF9EFFB9FD94FED4DBC9FEB3" author="Hilbig" box="[613,739,289,311]" pageId="33" pageNumber="34" refString="Hilbig, B. (2000) Family Terebellidae Grube, 1851. In: Blake, J., Hilbig, B. & Scott, P. V. (Eds.), Taxonomic Atlas of the Benthic Fauna of the Santa Barbara Basin and the Western Santa Barbara Channel. Vol. 7. The Annelida Part 4: Polychaeta (Flabelligeridae to Sternaspidae). Santa Barbara Museum of Natural History, Santa Barbara, California, pp. 230 - 293." type="book chapter" year="2000">Hilbig 2000</bibRefCitation>
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: 238
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</treatmentCitation>
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–240, Fig. 9.1.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="1DC56526FF9EFFB9FF66FE91DDDAFCA8" pageId="33" pageNumber="34" type="materials_examined">
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<paragraph id="556036ADFF9EFFB9FF66FE91DBE0FC8C" blockId="33.[151,1437,356,2001]" pageId="33" pageNumber="34">
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<emphasis id="67ABEABFFF9EFFB9FF66FE91D8A2FEF9" bold="true" box="[151,392,356,381]" pageId="33" pageNumber="34">Material examined.</emphasis>
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<typeStatus id="8A64880FFF9EFFB9FE67FE91D8D4FEF9" box="[406,510,356,381]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
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LACM-AHF Poly 230 (coll.
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<collectingCountry id="2DC8763DFF9EFFB9FC81FE91DA81FEF8" box="[880,939,356,380]" name="United States of America" pageId="33" pageNumber="34">USA</collectingCountry>
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, California, Marin County, Tomales Bay, Inverness, just north of village,
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<geoCoordinate id="30EB506AFF9EFFB9FE06FE72DB54FE24" box="[503,638,391,416]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="15" value="38.09833">38°05'54"N</geoCoordinate>
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,
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<geoCoordinate id="30EB506AFF9EFFB9FD78FE72DA0BFE24" box="[649,801,391,416]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="15" value="-122.851944">122°51'07"W</geoCoordinate>
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, mud flat, soft black mud mixed with shell fragments, by O. Hartman,
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<date id="2161106DFF9EFFB9FED9FE59D8A6FE41" box="[296,396,428,453]" pageId="33" pageNumber="34" value="1935-07">Jul.1935</date>
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): incomplete, in good state of preservation, ~
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<quantity id="92279B48FF9EFFB9FC73FE59DAFBFE40" box="[898,977,428,453]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="33" pageNumber="34" unit="mm" value="65.0">65 mm</quantity>
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||
long, ~
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<quantity id="92279B48FF9EFFB9FBD4FE59DD4CFE40" box="[1061,1126,428,452]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="33" pageNumber="34" unit="mm" value="4.0">4 mm</quantity>
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wide. Non-type specimens: LACM-AHF Poly 6517 [coll.
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<collectingCountry id="2DC8763DFF9EFFB9FDFBFE25DB6FFE6C" box="[522,581,464,488]" name="United States of America" pageId="33" pageNumber="34">USA</collectingCountry>
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||
, California, Monterey County, Moss Landing, Elkhorn Slough, several hundreds north of the current mouth (south mouth before mouth shifted),
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<geoCoordinate id="30EB506AFF9EFFB9FC27FE01DD76FD89" box="[982,1116,500,525]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="15" value="36.641945">36°38'31"N</geoCoordinate>
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||
,
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||
<geoCoordinate id="30EB506AFF9EFFB9FB99FE01DC2AFD88" box="[1128,1280,500,524]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="15" value="-121.78611">121°47'10"W</geoCoordinate>
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, mudflat, Sta. McGinitie 5, coll. T. H. Bullock,
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<date id="2161106DFF9EFFB9FDE8FDEDDB82FDB4" box="[537,680,536,560]" pageId="33" pageNumber="34" value="1939-06-19">19.Jun.1939</date>
|
||
]:
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||
<specimenCount id="43D9FD24FF9EFFB9FD4DFDE2DA61FDB4" box="[700,843,535,560]" pageId="33" pageNumber="34" type="generic">6 specimens</specimenCount>
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||
, some complete, although several in more than one piece, most of them in good state of preservation, dark brown to purple; slides (from complete specimen, in two pieces, LACM-AHF 6517): notochaetae segment 7, neurochaetae mid-abdominal segment, neurochaetae posterior segment. LACM-AHF Poly 6518 (coll.
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||
<collectingCountry id="2DC8763DFF9EFFB9FDA7FD71DBBBFD18" box="[598,657,644,668]" name="United States of America" pageId="33" pageNumber="34">USA</collectingCountry>
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||
, California, Humboldt County, off Eureka, Eel River South, MBARI- Seep,
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<geoCoordinate id="30EB506AFF9EFFB9FF2DFD5DD848FD44" box="[220,354,680,704]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="15" value="40.703888">40°42'14"N</geoCoordinate>
|
||
,
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||
<geoCoordinate id="30EB506AFF9EFFB9FE9CFD5DDB2CFD44" box="[365,518,680,704]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="15" value="-124.37278">124°22'22"W</geoCoordinate>
|
||
,
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||
<quantity id="92279B48FF9EFFB9FDE0FD52DB5AFD3B" box="[529,624,679,704]" metricMagnitude="1" metricUnit="m" metricValue="3.9619999999999997" pageId="33" pageNumber="34" unit="m" value="39.62">39.62 m</quantity>
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, sediment near cold seep, Shipek grab, R/V McGAW Sta. Shipek 65, by Lisa Levin, Leslie H. Harris et al.,
|
||
<date id="2161106DFF9EFFB9FDE9FD39DB8FFD60" box="[536,677,716,740]" pageId="33" pageNumber="34" value="1998-10-21">21.Oct.1998</date>
|
||
, id L. Harris): complete specimen, in excellent state of preservation, ~
|
||
<quantity id="92279B48FF9EFFB9FF57FD05D9DFFC83" box="[166,245,752,776]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="33" pageNumber="34" unit="mm" value="31.0">31 mm</quantity>
|
||
long,
|
||
<quantity id="92279B48FF9EFFB9FECAFD05D856FC83" box="[315,380,752,776]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="33" pageNumber="34" unit="mm" value="2.0">2 mm</quantity>
|
||
wide, mounted on SEM stub.
|
||
</paragraph>
|
||
<paragraph id="556036ADFF9EFFB9FF36FCE1DDDAFCA8" blockId="33.[151,1437,356,2001]" box="[199,1264,788,813]" pageId="33" pageNumber="34">
|
||
<emphasis id="67ABEABFFF9EFFB9FF36FCE1D84FFCA9" bold="true" box="[199,357,788,813]" pageId="33" pageNumber="34">
|
||
<typeStatus id="8A64880FFF9EFFB9FF36FCE1D828FCA9" box="[199,258,788,813]" pageId="33" pageNumber="34">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="2DC8763DFF9EFFB9FE9DFCE1D88DFCA8" box="[364,423,788,812]" name="United States of America" pageId="33" pageNumber="34">USA</collectingCountry>
|
||
, California, Marin County, Tomales Bay,
|
||
<geoCoordinate id="30EB506AFF9EFFB9FC8BFCE1DD2BFCA9" box="[890,1025,788,813]" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="15" value="38.09833">38°05'54"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30EB506AFF9EFFB9FBFDFCE1DD8FFCA9" box="[1036,1189,788,813]" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="15" value="-122.851944">122°51'07"W</geoCoordinate>
|
||
,
|
||
<specimenCount id="43D9FD24FF9EFFB9FB41FCE1DDDAFCA8" box="[1200,1264,788,813]" pageId="33" pageNumber="34" type="generic">
|
||
<quantity id="92279B48FF9EFFB9FB41FCE1DDC6FCA8" box="[1200,1260,788,813]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="33" pageNumber="34" unit="m" value="58.0">58 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1DC56526FF9EFFBFFF36FCC2DC0EFF50" lastPageId="39" lastPageNumber="40" pageId="33" pageNumber="34" type="description">
|
||
<paragraph id="556036ADFF9EFFB9FF36FCC2DC41FCF1" blockId="33.[151,1437,356,2001]" pageId="33" pageNumber="34">
|
||
<emphasis id="67ABEABFFF9EFFB9FF36FCC2D873FCD4" bold="true" box="[199,345,823,848]" pageId="33" pageNumber="34">Description.</emphasis>
|
||
<typeStatus id="8A64880FFF9EFFB9FE93FCC2D8E0FCD4" box="[354,458,823,848]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
|
||
incomplete specimen, ~
|
||
<quantity id="92279B48FF9EFFB9FD12FCC2DA1FFCCB" box="[739,821,823,848]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="33" pageNumber="34" unit="mm" value="65.0">65 mm</quantity>
|
||
long, ~
|
||
<quantity id="92279B48FF9EFFB9FC7EFCCDDAF9FCCB" box="[911,979,824,848]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="33" pageNumber="34" unit="mm" value="4.0">4 mm</quantity>
|
||
wide at segment 8, maximum width of body. Life specimens yellow sulphur, becoming purple after preservation (L. Harris personal communication).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF9EFFB9FF36FC8AD9EDFB11" blockId="33.[151,1437,356,2001]" pageId="33" pageNumber="34">
|
||
Prostomium at base of upper lip, both basal and distal parts developed, basal part as thickened crest, distal part with large, rounded flaring lobes, mid-dorsal process not visible, if present, hidden by large number of buccal tentacles; prostomium covering segment 1 laterally and terminating laterally to lower lip, near mouth (
|
||
<figureCitation id="CDE42A28FF9EFFB9FAFFFC32DC4AFC64" box="[1294,1376,967,992]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A–E, G–H; 19A, C; 20B–E, G; 21G). Buccal tentacles of three
|
||
<typeStatus id="8A64880FFF9EFFB9FCDBFC18DA42FB81" box="[810,872,1005,1029]" pageId="33" pageNumber="34">types</typeStatus>
|
||
, short ones thin, uniformly cylindrical, except for expanded sphaerical to elliptical tips; intermediate tentacles thin, uniformly cylindrical, also expanded at tip; long buccal tentacles progressively widening at tip, towards clearly marked subdistal cylindrical inflation, with short and pointed tip (
|
||
<figureCitation id="CDE42A28FF9EFFB9FEA2FBA2D88DFBF4" box="[339,423,1111,1136]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A–E, G–H; 19A, C; 20A–G; 21A–G); all
|
||
<typeStatus id="8A64880FFF9EFFB9FC7BFBACDAEFFBF5" box="[906,965,1113,1137]" pageId="33" pageNumber="34">types</typeStatus>
|
||
with ciliated groove only conspicuous at tips.
|
||
</paragraph>
|
||
<paragraph id="556036ADFF9EFFB9FF36FB6ADB13FA14" blockId="33.[151,1437,356,2001]" pageId="33" pageNumber="34">
|
||
Peristomium restricted to lips, upper lip almost circular; lower lip short, rounded (
|
||
<figureCitation id="CDE42A28FF9EFFB9FB94FB6ADD91FB3C" box="[1125,1211,1183,1208]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A, C, E, G–H; 19A, C; 20B, D–E, G; 21G). Body abruptly broader on segments 3–5, of relatively uniform width until segment 10, slightly tapered on segments 11–14, then more markedly tapering to narrower uniformly cylindrical posterior body, beginning from segment 17–18 and tapering near pygidium (
|
||
<figureCitation id="CDE42A28FF9EFFB9FCB2FAF9DAB5FAA0" box="[835,927,1292,1317]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A–H; 19A, C, F; 20A–I); long achaetous gap between termination of notopodia and beginning of neuropodia, corresponding to segments 15–37, with poorly marked segmentation and fragile, with thin body wall dorsally, distinctly longer than region with notopodia (
|
||
<figureCitation id="CDE42A28FF9EFFB9FA9BFAA1D999FA14" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A, B–D, F; 19A, F; 20A–B, G–H).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF9EFFB9FF21FA69DC7EF908" blockId="33.[151,1437,356,2001]" pageId="33" pageNumber="34">
|
||
Segments biannulated, segment 1 short, visible dorsally and ventrally; segment 2 narrower and shorter than following segments, with relatively small, pentagonal mid-ventral shield at beginning of mid-ventral groove, extending anteriorly through segment 1 until near ventral edge of lower lip, shield about same length as that of segment 3 and slightly broader (
|
||
<figureCitation id="CDE42A28FF9EFFB9FE0EF9F2DB7BF9A4" box="[511,593,1543,1568]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A–E, G–H; 19A, C; 20D–E; 21G). Ventrum highly glandular, covered with small papillae, arranged in paired ventro-lateral pads on segments 2–14; papillae about same size throughout and slightly more numerous on anterior segments; from after termination of notopodia, paired longitudinal crests bordering mid-ventral groove through posterior body (
|
||
<figureCitation id="CDE42A28FF9EFFB9FD08F981DA67F908" box="[761,845,1652,1677]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A, C, E, H; 19A, C, F; 20A–E, G–H; 22A–B).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF9EFFB9FF36F962D844F8C4" blockId="33.[151,1437,356,2001]" pageId="33" pageNumber="34">
|
||
Notopodia extending through 12 segments, until segment 14; elongate, cylindrical notopodia, with equal sized lobes and elongate and distally blunt tips (
|
||
<figureCitation id="CDE42A28FF9EFFB9FD88F949DBE7F950" box="[633,717,1724,1749]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A–E, G–H; 19A, C; 20A–H; 21G; 22G–I). Acicular, narrowlywinged notochaetae in both rows, wings inconspicuous under higher magnifications of light microscopy, those from anterior row ¼ length of chaetae from posterior row, or less, barely protruding from parapodial lobes (
|
||
<figureCitation id="CDE42A28FF9EFFB9FA9BF8F1D99EF8C4" captionStart="FIGURE 21" captionStartId="37.[151,250,1950,1972]" captionTargetBox="[181,1407,197,1921]" captionTargetId="figure@37.[174,1411,193,1928]" captionTargetPageId="37" captionText="FIGURE 21. Amaeana occidentalis LACM-AHF Poly 6518. A – F, tips of buccal tentacles. G, close up view of the anterior end, ventral view. H, close up of the posterior end, frontal view. Numbers refer to segments; ll = lower lip; P = basal part of prostomium; ul = upper lip. Scale bars: A, G = 0.2 mm; B, D, F = 0.1 mm; C = 0.15 mm; E = 40 µm; H = 50 µm." httpUri="https://zenodo.org/record/289158/files/figure.png" pageId="33" pageNumber="34">Figs 21</figureCitation>
|
||
B, D–E; 22G–I).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF9EFFB9FF36F8B9DAF8F854" blockId="33.[151,1437,356,2001]" pageId="33" pageNumber="34">
|
||
Neuropodia present from segment 38, laterally to mid-ventral groove, on outer margins of longitudinal crests. On anterior neuropodia, neurochaetae as up to 12–13 relatively stout, distally tapered spines, slightly expanded subdistally, with oblique tip (
|
||
<figureCitation id="CDE42A28FF9EFFB9FE1AF861DB6AF828" box="[491,576,1940,1965]" captionStart="FIGURE 19" captionStartId="35.[151,250,1886,1908]" captionTargetBox="[209,1375,197,1851]" captionTargetId="figure@35.[204,1380,193,1856]" captionTargetPageId="35" captionText="FIGURE 19. Amaeana occidentalis LACM-AHF Poly 6518. A, entire worm, left ventro-lateral view (anteriorly). C, anterior end, left ventro-lateral view. F, posterior end. Specimen LACM-AHF Poly 6517. B, notochaetae, segment 7; arrow point to notochaetae from anterior row. D, notochaetae from anterior row, segment 7. E, tips of notochaetae from posterior row, segment 7. G, mid-abdominal neurochaetae. H, posterior neurochaetae. Scale bars: A = 1 mm; C, F = 0.6 mm; B = 100 µm; D, G = 20 µm; E = 30 µm; H = 10 µm." httpUri="https://zenodo.org/record/289156/files/figure.png" pageId="33" pageNumber="34">Fig. 19</figureCitation>
|
||
G); posterior neuropodia with fewer spines, 8–9 (
|
||
<figureCitation id="CDE42A28FF9EFFB9FB70F861DDFCF828" box="[1153,1238,1940,1965]" captionStart="FIGURE 19" captionStartId="35.[151,250,1886,1908]" captionTargetBox="[209,1375,197,1851]" captionTargetId="figure@35.[204,1380,193,1856]" captionTargetPageId="35" captionText="FIGURE 19. Amaeana occidentalis LACM-AHF Poly 6518. A, entire worm, left ventro-lateral view (anteriorly). C, anterior end, left ventro-lateral view. F, posterior end. Specimen LACM-AHF Poly 6517. B, notochaetae, segment 7; arrow point to notochaetae from anterior row. D, notochaetae from anterior row, segment 7. E, tips of notochaetae from posterior row, segment 7. G, mid-abdominal neurochaetae. H, posterior neurochaetae. Scale bars: A = 1 mm; C, F = 0.6 mm; B = 100 µm; D, G = 20 µm; E = 30 µm; H = 10 µm." httpUri="https://zenodo.org/record/289156/files/figure.png" pageId="33" pageNumber="34">Fig. 19</figureCitation>
|
||
H); at least some spines protruding from neuropodia (
|
||
<figureCitation id="CDE42A28FF9EFFB9FDDAF84DDB55F854" box="[555,639,1976,2001]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="33" pageNumber="34">Figs 18</figureCitation>
|
||
A–B, F; 19A, F;
|
||
<date id="2161106DFF9EFFB9FCCFF84DDA4DF854" box="[830,871,1976,2000]" pageId="33" pageNumber="34" value="1920-01">20I</date>
|
||
; 22A–E).
|
||
</paragraph>
|
||
<caption id="01A06625FF9DFFBAFF66F8AFD8D0F86F" httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="34" pageNumber="35" targetBox="[207,1387,197,1851]" targetPageId="34">
|
||
<paragraph id="556036ADFF9DFFBAFF66F8AFD8D0F86F" blockId="34.[151,1436,1882,2027]" pageId="34" pageNumber="35">
|
||
<emphasis id="67ABEABFFF9DFFBAFF66F8AFD835F8EB" bold="true" box="[151,287,1882,1904]" pageId="34" pageNumber="35">FIGURE 18.</emphasis>
|
||
<taxonomicName id="92DF4D2EFF9DFFBAFED7F8AEDB2DF8F4" box="[294,519,1882,1904]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="34" pageNumber="35" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF9DFFBAFED7F8AEDB2DF8F4" box="[294,519,1882,1904]" italics="true" pageId="34" pageNumber="35">Amaeana occidentalis</emphasis>
|
||
</taxonomicName>
|
||
holotype LACM-AHF Poly 230. A–B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D–E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G–H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A–B, E = 1.5 mm; C–D = 1 mm; F–H = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="01A06625FF9CFFBBFF66F8ABD89CF86A" httpUri="https://zenodo.org/record/289156/files/figure.png" pageId="35" pageNumber="36" targetBox="[209,1375,197,1851]" targetPageId="35">
|
||
<paragraph id="556036ADFF9CFFBBFF66F8ABD89CF86A" blockId="35.[151,1436,1886,2030]" pageId="35" pageNumber="36">
|
||
<emphasis id="67ABEABFFF9CFFBBFF66F8ABD80AF8F7" bold="true" box="[151,288,1886,1908]" pageId="35" pageNumber="36">FIGURE 19.</emphasis>
|
||
<taxonomicName id="92DF4D2EFF9CFFBBFED9F8AADB23F8F0" box="[296,521,1886,1908]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="35" pageNumber="36" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF9CFFBBFED9F8AADB23F8F0" box="[296,521,1886,1908]" italics="true" pageId="35" pageNumber="36">Amaeana occidentalis</emphasis>
|
||
</taxonomicName>
|
||
LACM-AHF Poly 6518. A, entire worm, left ventro-lateral view (anteriorly). C, anterior end, left ventro-lateral view. F, posterior end. Specimen LACM-AHF Poly 6517. B, notochaetae, segment 7; arrow point to notochaetae from anterior row. D, notochaetae from anterior row, segment 7. E, tips of notochaetae from posterior row, segment 7. G, mid-abdominal neurochaetae. H, posterior neurochaetae. Scale bars: A = 1 mm; C, F = 0.6 mm; B = 100 µm; D, G = 20 µm; E = 30 µm; H = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="01A06625FF9BFFBCFF66F89EDA57F859" httpUri="https://zenodo.org/record/289157/files/figure.png" pageId="36" pageNumber="37" targetBox="[199,1394,178,1870]" targetPageId="36">
|
||
<paragraph id="556036ADFF9BFFBCFF66F89EDA57F859" blockId="36.[151,1437,1899,2013]" pageId="36" pageNumber="37">
|
||
<emphasis id="67ABEABFFF9BFFBCFF66F89ED835F804" bold="true" box="[151,287,1899,1921]" pageId="36" pageNumber="37">FIGURE 20.</emphasis>
|
||
<taxonomicName id="92DF4D2EFF9BFFBCFED4F899DB2FF805" box="[293,517,1899,1921]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF9BFFBCFED4F899DB2FF805" box="[293,517,1899,1921]" italics="true" pageId="36" pageNumber="37">Amaeana occidentalis</emphasis>
|
||
</taxonomicName>
|
||
LACM-AHF Poly 6518. A–B, thorax, right lateral and left dorso-lateral views. C, anterior end, left dorso-lateral view. D–E, progressively closer ventral views of the thorax, ventral views. F–G, thorax, dorsal and left lateral views. H, segments 10–15, left lateral view; arrows point to genital papillae. I, posterior end, left ventro-lateral view. Scale bars: A–D, F = 0.7 mm; E, G = 0.4 mm; H = 0.3 mm; I = 0.15 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="01A06625FF9AFFBDFF66F86BDC39F875" httpUri="https://zenodo.org/record/289158/files/figure.png" pageId="37" pageNumber="38" targetBox="[181,1407,197,1921]" targetPageId="37">
|
||
<paragraph id="556036ADFF9AFFBDFF66F86BDC39F875" blockId="37.[151,1436,1950,2033]" pageId="37" pageNumber="38">
|
||
<emphasis id="67ABEABFFF9AFFBDFF66F86BD835F837" bold="true" box="[151,287,1950,1972]" pageId="37" pageNumber="38">FIGURE 21.</emphasis>
|
||
<taxonomicName id="92DF4D2EFF9AFFBDFED5F86ADB2EF830" box="[292,516,1950,1972]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF9AFFBDFED5F86ADB2EF830" box="[292,516,1950,1972]" italics="true" pageId="37" pageNumber="38">Amaeana occidentalis</emphasis>
|
||
</taxonomicName>
|
||
LACM-AHF Poly 6518. A–F, tips of buccal tentacles. G, close up view of the anterior end, ventral view. H, close up of the posterior end, frontal view. Numbers refer to segments; ll = lower lip; P = basal part of prostomium; ul = upper lip. Scale bars: A, G = 0.2 mm; B, D, F = 0.1 mm; C = 0.15 mm; E = 40 µm; H = 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="01A06625FF99FFBEFF66F88FDA61F86F" httpUri="https://zenodo.org/record/289159/files/figure.png" pageId="38" pageNumber="39" targetBox="[180,1422,197,1876]" targetPageId="38">
|
||
<paragraph id="556036ADFF99FFBEFF66F88FDA61F86F" blockId="38.[151,1436,1914,2027]" pageId="38" pageNumber="39">
|
||
<emphasis id="67ABEABFFF99FFBEFF66F88FD80FF80B" bold="true" box="[151,293,1914,1936]" pageId="38" pageNumber="39">FIGURE 22.</emphasis>
|
||
<taxonomicName id="92DF4D2EFF99FFBEFEC3F88FDB32F814" box="[306,536,1914,1936]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="38" pageNumber="39" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF99FFBEFEC3F88FDB32F814" box="[306,536,1914,1936]" italics="true" pageId="38" pageNumber="39">Amaeana occidentalis</emphasis>
|
||
</taxonomicName>
|
||
LACM-AHF Poly 6518. A, posterior end, ventral view (anteriorly). B, abdominal segments. C–F, close ups of abdominal neuropodia, with protruding spines. G, notopodia, segments 3–5; arrows point to nephridial papillae. H–I, progressively closer views of the notochaetae of segment 5. Scale bars: A = 0.5 mm; B = 0.15 mm; C = 10 µm; D = 3 µm; E = 5 µm; F = 2 µm; G = 70 µm; H–I = 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="556036ADFF98FFBFFF36FF62DC0EFF50" blockId="39.[151,1437,151,752]" pageId="39" pageNumber="40">
|
||
Nephridial and genital papillae anterior to bases of all notopodia, those of segments 6–9 largest (
|
||
<figureCitation id="CDE42A28FF98FFBFFAFCFF62DC48FF34" box="[1293,1378,151,176]" captionStart="FIGURE 18" captionStartId="34.[151,250,1882,1904]" captionTargetBox="[207,1387,197,1851]" captionTargetId="figure@34.[201,1391,193,1862]" captionTargetPageId="34" captionText="FIGURE 18. Amaeana occidentalis holotype LACM-AHF Poly 230. A – B, entire worm, ventral and dorsal views (anteriorly), respectively. C, anterior end, ventral view. D – E, thorax dorsal and ventral views, respectively. F, abdominal segments; arrow points to first neuropodium. G – H, close ups of the anterior end, dorsal and ventral views, respectively. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; ul = upper lip; * = distal part of prostomium. Scale bars: A – B, E = 1.5 mm; C – D = 1 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/289155/files/figure.png" pageId="39" pageNumber="40">Figs 18</figureCitation>
|
||
B–E, G–H; 19A, C; 20A–H). Pygidium crenulate, with slightly larger ventral papilla (
|
||
<figureCitation id="CDE42A28FF98FFBFFBE8FF49DD44FF50" box="[1049,1134,188,213]" captionStart="FIGURE 19" captionStartId="35.[151,250,1886,1908]" captionTargetBox="[209,1375,197,1851]" captionTargetId="figure@35.[204,1380,193,1856]" captionTargetPageId="35" captionText="FIGURE 19. Amaeana occidentalis LACM-AHF Poly 6518. A, entire worm, left ventro-lateral view (anteriorly). C, anterior end, left ventro-lateral view. F, posterior end. Specimen LACM-AHF Poly 6517. B, notochaetae, segment 7; arrow point to notochaetae from anterior row. D, notochaetae from anterior row, segment 7. E, tips of notochaetae from posterior row, segment 7. G, mid-abdominal neurochaetae. H, posterior neurochaetae. Scale bars: A = 1 mm; C, F = 0.6 mm; B = 100 µm; D, G = 20 µm; E = 30 µm; H = 10 µm." httpUri="https://zenodo.org/record/289156/files/figure.png" pageId="39" pageNumber="40">Figs 19</figureCitation>
|
||
A, F;
|
||
<date id="2161106DFF98FFBFFB5EFF49DDF2FF50" box="[1199,1240,188,212]" pageId="39" pageNumber="40" value="1920-01">20I</date>
|
||
; 21H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1DC56526FF98FFBFFF36FF2AD8CEFD74" pageId="39" pageNumber="40" type="discussion">
|
||
<paragraph id="556036ADFF98FFBFFF36FF2ADB6DFE0C" blockId="39.[151,1437,151,752]" pageId="39" pageNumber="40">
|
||
<emphasis id="67ABEABFFF98FFBFFF36FF2AD811FF7C" bold="true" box="[199,315,223,248]" pageId="39" pageNumber="40">Remarks.</emphasis>
|
||
<taxonomicName id="92DF4D2EFF98FFBFFEB0FF15DB13FF7C" box="[321,569,224,248]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF98FFBFFEB0FF15DB13FF7C" box="[321,569,224,248]" italics="true" pageId="39" pageNumber="40">Amaeana occidentalis</emphasis>
|
||
</taxonomicName>
|
||
is characterised by the great length of the achaetous gap between termination of notopodia and beginning of neuropodia, consisting of more than 20 segments. Also, members of
|
||
<taxonomicName id="92DF4D2EFF98FFBFFB04FEF1DCB1FE98" box="[1269,1435,260,284]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF98FFBFFB04FEF1DCB1FE98" box="[1269,1435,260,284]" italics="true" pageId="39" pageNumber="40">A. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
present the highest numbers of spines per neuropodium, 12–13 on anterior neuropodia, 8–9 on posterior ones, and numbers of pairs of notopodia among all species of
|
||
<taxonomicName id="92DF4D2EFF98FFBFFD2AFEB9DA6FFEE7" box="[731,837,332,355]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="genus">
|
||
<emphasis id="67ABEABFFF98FFBFFD2AFEB9DA6FFEE7" box="[731,837,332,355]" italics="true" pageId="39" pageNumber="40">Amaeana</emphasis>
|
||
</taxonomicName>
|
||
, 12 pairs, although this latter character is shared with
|
||
<taxonomicName id="92DF4D2EFF98FFBFFF66FE85D804FE03" box="[151,302,368,391]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="accraensis">
|
||
<emphasis id="67ABEABFFF98FFBFFF66FE85D804FE03" box="[151,302,368,391]" italics="true" pageId="39" pageNumber="40">A. accraensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="92DF4D2EFF98FFBFFE95FE85D8FEFE03" box="[356,468,368,391]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="yirrarn">
|
||
<emphasis id="67ABEABFFF98FFBFFE95FE85D8FEFE03" box="[356,468,368,391]" italics="true" pageId="39" pageNumber="40">A. yirrarn</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="185D0316FF98FFBFFE13FE9ADB1CFE0C" box="[482,566,367,392]" captionStart="TABLE 1" captionStartId="4.[119,199,561,582]" captionText="TABLE 1. Comparison of selected morphological characters between species of Amaeana." pageId="39" pageNumber="40">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF98FFBFFF36FE61DD6DFD9C" blockId="39.[151,1437,151,752]" pageId="39" pageNumber="40">
|
||
The only other species with more segments in the achaetous gap is
|
||
<taxonomicName id="92DF4D2EFF98FFBFFC33FE61DC29FE29" authority="Augener 1926" authorityName="Augener" authorityYear="1926" box="[962,1283,404,429]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="antipoda">
|
||
<emphasis id="67ABEABFFF98FFBFFC33FE61DD6CFE28" box="[962,1094,404,428]" italics="true" pageId="39" pageNumber="40">A. antipoda</emphasis>
|
||
(
|
||
<bibRefCitation id="314E4B5CFF98FFBFFBA7FE61DDD6FE29" author="Augener" box="[1110,1276,404,429]" pageId="39" pageNumber="40" refString="Augener, H. (1926) Papers from Dr. T. H. Mortensen's Pacific Expedition 1914 - 1916. No. 34. Polychaeta Ill. Polychaeten von Neuseeland. 11. Sedentaria. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening i Kjobenhavn, 81, 157 - 294." type="journal article" year="1926">Augener 1926</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, according to the original description, but members of this species only have 13–15 achaetous segments and have fewer neuropodial spines per neuropodium, about 4–5 spines, and only 11 pairs of notopodia (
|
||
<tableCitation id="185D0316FF98FFBFFB79FE29DDF7FE70" box="[1160,1245,476,501]" captionStart="TABLE 1" captionStartId="4.[119,199,561,582]" captionText="TABLE 1. Comparison of selected morphological characters between species of Amaeana." pageId="39" pageNumber="40">Table 1</tableCitation>
|
||
). In addition, as said above, we consider this species as
|
||
<emphasis id="67ABEABFFF98FFBFFDBEFDF4DBD9FD9C" box="[591,755,512,536]" italics="true" pageId="39" pageNumber="40">nomem nudum</emphasis>
|
||
(see
|
||
<emphasis id="67ABEABFFF98FFBFFCDFFE0ADD1CFD9C" bold="true" box="[814,1078,511,536]" pageId="39" pageNumber="40">Material and methods</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF98FFBFFF36FDD1D8CEFD74" blockId="39.[151,1437,151,752]" pageId="39" pageNumber="40">
|
||
The other species of
|
||
<taxonomicName id="92DF4D2EFF98FFBFFE48FDD1DB09FDBF" box="[441,547,548,571]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="genus">
|
||
<emphasis id="67ABEABFFF98FFBFFE48FDD1DB09FDBF" box="[441,547,548,571]" italics="true" pageId="39" pageNumber="40">Amaeana</emphasis>
|
||
</taxonomicName>
|
||
with 12 pairs of notopodia are both easily distinguished from
|
||
<taxonomicName id="92DF4D2EFF98FFBFFB02FDD1D867FDE4" authority="Hartman 1944" authorityName="Hartman" authorityYear="1944" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF98FFBFFB02FDD1DCB1FDB8" box="[1267,1435,548,572]" italics="true" pageId="39" pageNumber="40">A. occidentalis</emphasis>
|
||
(
|
||
<bibRefCitation id="314E4B5CFF98FFBFFF6FFDBDD86CFDE4" author="Hartman" box="[158,326,584,608]" pageId="39" pageNumber="40" refString="Hartman, O. (1944) Polychaetous Annelids from California including the descriptions of two new genera and nine new species. Allan Hancock Pacific Expedition, 10, 239 - 290." type="journal article" year="1944">Hartman 1944</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, by the number of spines per neuropodium, in addition to the extension of the achaetous gap. The
|
||
<typeStatus id="8A64880FFF98FFBFFF66FD98D9EDFD01" box="[151,199,621,645]" pageId="39" pageNumber="40">type</typeStatus>
|
||
of
|
||
<taxonomicName id="92DF4D2EFF98FFBFFF00FD99D8A1FD00" box="[241,395,620,644]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="accraensis">
|
||
<emphasis id="67ABEABFFF98FFBFFF00FD99D8A1FD00" box="[241,395,620,644]" italics="true" pageId="39" pageNumber="40">A. accraensis</emphasis>
|
||
</taxonomicName>
|
||
has an achaetous gap of 5–6 segments and 8 spines per neuropodium, while in
|
||
<taxonomicName id="92DF4D2EFF98FFBFFAD8FD99D872FD2C" authority="Hutchings, 1997" authorityName="Hutchings" authorityYear="1997" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="yirrarn">
|
||
<emphasis id="67ABEABFFF98FFBFFAD8FD99DCB6FD00" box="[1321,1436,620,644]" italics="true" pageId="39" pageNumber="40">A. yirrarn</emphasis>
|
||
<bibRefCitation id="314E4B5CFF98FFBFFF66FD7AD872FD2C" author="Hutchings" box="[151,344,655,680]" pageId="39" pageNumber="40" refString="Hutchings, P. A. (1997) New species of the family Terebellidae (Polychaeta) from Darwin Harbour Australia. In: Hanley, R. (Ed.), ' Proceedings of the 6 th International Marine Biological Workshop. The Marine Flora and Fauna of Darwin Harbour, Northern Territory'. pp. 133 - 161." type="book chapter" year="1997">Hutchings, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
, the gap extends for 7–9 segments and there are 1–2 spines per neuropodium (
|
||
<tableCitation id="185D0316FF98FFBFFAFFFD7ADC4CFD2C" box="[1294,1382,655,680]" captionStart="TABLE 1" captionStartId="4.[119,199,561,582]" captionText="TABLE 1. Comparison of selected morphological characters between species of Amaeana." pageId="39" pageNumber="40">Table 1</tableCitation>
|
||
). In addition, notochaetae in the posterior row are distally pinnate among specimens of
|
||
<taxonomicName id="92DF4D2EFF98FFBFFB9DFD41DC2DFD48" box="[1132,1287,692,716]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="accraensis">
|
||
<emphasis id="67ABEABFFF98FFBFFB9DFD41DC2DFD48" box="[1132,1287,692,716]" italics="true" pageId="39" pageNumber="40">A. accraensis</emphasis>
|
||
</taxonomicName>
|
||
, while in
|
||
<taxonomicName id="92DF4D2EFF98FFBFFA75FD41D834FD74" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="39" pageNumber="40" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF98FFBFFA75FD41D834FD74" italics="true" pageId="39" pageNumber="40">A. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
they are acicular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |