227 lines
32 KiB
XML
227 lines
32 KiB
XML
<document id="DF8675CC65CEE0261ED1BDF4B3DFE8BE" ID-DOI="10.11646/zootaxa.3994.1.1" ID-GBIF-Dataset="1334b429-62a4-4505-9e05-d68b39058e5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="289137" ID-ZooBank="093B124E-58AE-4303-8C07-2D7B27E6AC38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461271356316" checkinUser="plazi" docAuthor="Nogueira, João Miguel De Matos, Carrerette, Orlemir & Hutchings, Pat" docDate="2015" docId="DD7687BBFF92FFB6FF66FC63D9CBFD42" docLanguage="en" docName="zt03994p052.pdf" docOrigin="Zootaxa 3994 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amaeana accraensis Augener 1918" docType="treatment" docVersion="6" lastPageNumber="47" masterDocId="214FFFC3FFBFFF98FFF1FFF5D92AFF84" masterDocTitle="Review of Amaeana Hartman, 1959 (Annelida, Terebelliformia, Polycirridae), with descriptions of seven new species" masterLastPageNumber="52" masterPageNumber="1" pageNumber="46" updateTime="1698611839755" updateUser="plazi">
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<mods:mods id="5F3BAB83EB3E246B44D4A5DA67EC99C1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="972FD7C9BEFFA2E533739E2828B26B15">Review of Amaeana Hartman, 1959 (Annelida, Terebelliformia, Polycirridae), with descriptions of seven new species</mods:title>
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<mods:name id="9CC674EDA72BD0B3AD778FDFE464641A" type="personal">
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<mods:namePart id="781B71D63F04A15A223C02FC8C77AD3B">Nogueira, João Miguel De Matos</mods:namePart>
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<mods:roleTerm id="3BE110EE9CC745549F06BCD924AE08A8">Author</mods:roleTerm>
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<mods:namePart id="4B515B4B0D3C75C7AD16C912A545869A">Carrerette, Orlemir</mods:namePart>
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<mods:name id="8BB8AB998378B89FBB3ED80C3ABA79D8" type="personal">
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<mods:roleTerm id="F444172031FC520FD010301EEBC632E5">Author</mods:roleTerm>
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<mods:namePart id="197F943C690CA969CC3BA0275AB35332">Hutchings, Pat</mods:namePart>
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<mods:typeOfResource id="4C896EC82A7225B8D9AA20579544B2F9">text</mods:typeOfResource>
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<mods:title id="83BD2987D04369D0AE5E44BF3F84011D">Zootaxa</mods:title>
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<mods:date id="AD8B9373DA2F2E53AC9DDF9DD7CF86B2">2015</mods:date>
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<mods:number id="91BAB8B33251748CFF7B20277E7FC55E">3994</mods:number>
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<mods:classification id="5467686619698DB3469BEDD4E0CBE858">journal article</mods:classification>
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<mods:identifier id="FFFAD47D23C07118EBD488FB69BFF607" type="DOI">10.11646/zootaxa.3994.1.1</mods:identifier>
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<mods:identifier id="E1D1CEFCE509E8374D1D1A3A83BC12E3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="DD7687BBFF92FFB6FF66FC63D9CBFD42" ID-DOI="http://doi.org/10.5281/zenodo.6094926" ID-GBIF-Taxon="127692615" ID-Zenodo-Dep="6094926" LSID="urn:lsid:plazi:treatment:DD7687BBFF92FFB6FF66FC63D9CBFD42" httpUri="http://treatment.plazi.org/id/DD7687BBFF92FFB6FF66FC63D9CBFD42" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<subSubSection id="1DC56526FF92FFB5FF66FC63D84AFC56" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="556036ADFF92FFB5FF66FC63DB77FC35" blockId="45.[151,605,918,978]" box="[151,605,918,945]" pageId="45" pageNumber="46">
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<heading id="0E2881C1FF92FFB5FF66FC63DB77FC35" bold="true" box="[151,605,918,945]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
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<taxonomicName id="92DF4D2EFF92FFB5FF66FC63DB77FC35" ID-CoL="5TXLB" authority="Augener, 1918" authorityName="Augener" authorityYear="1918" box="[151,605,918,945]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="45" pageNumber="46" phylum="Annelida" rank="species" species="accraensis">
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<emphasis id="67ABEABFFF92FFB5FF66FC63DB77FC35" bold="true" box="[151,605,918,945]" pageId="45" pageNumber="46">
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<emphasis id="67ABEABFFF92FFB5FF66FC63D8BBFC34" bold="true" box="[151,401,918,944]" italics="true" pageId="45" pageNumber="46">Amaeana accraensis</emphasis>
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(
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<bibRefCitation id="314E4B5CFF92FFB5FE51FC62DB7FFC35" author="Augener" box="[416,597,919,945]" pageId="45" pageNumber="46" refString="Augener, H. (1918) Polychaeta Beitr a ge zur Kenntnis des Meeresfauna West-Afrikas. Herausgegeben von W. Michaelsen Hamburg, 2, 67 - 625." type="journal article" year="1918">Augener, 1918</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="556036ADFF92FFB5FF66FC4CD84AFC56" blockId="45.[151,605,918,978]" box="[151,352,953,978]" pageId="45" pageNumber="46">
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<figureCitation id="CDE42A28FF92FFB5FF66FC4CD824FC56" box="[151,270,953,978]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="45" pageNumber="46">Figures 26</figureCitation>
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and
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<figureCitation id="CDE42A28FF92FFB5FEB5FC4FD84AFC56" box="[324,352,954,978]" captionStart="FIGURE 27" captionStartId="45.[151,250,757,779]" captionTargetBox="[158,1431,197,730]" captionTargetId="figure@45.[152,1435,193,736]" captionTargetPageId="45" captionText="FIGURE 27. Amaeana accraensis holotype ZMH V- 1710. A, notochaetae, segment 8; B, notochaetae from anterior row, segment 8; C, tips of notochaetae from posterior row, segment 8; D, neuropodial spines, segment 42. Scale bars: A = 30 µm; B – D = 10 µm." httpUri="https://zenodo.org/record/289164/files/figure.png" pageId="45" pageNumber="46">27</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1DC56526FF92FFB5FF66FBF7DB7FFBB2" pageId="45" pageNumber="46" type="reference_group">
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<paragraph id="556036ADFF92FFB5FF66FBF7DB7FFBB2" blockId="45.[151,954,1025,1078]" pageId="45" pageNumber="46">
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<treatmentCitationGroup id="75CF1183FF92FFB5FF66FBF7DB7FFBB2" pageId="45" pageNumber="46">
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<taxonomicName id="92DF4D2EFF92FFB5FF66FBF7DB32FB93" authority="Augener 1918: 561" authorityName="Augener" authorityPageNumber="561" authorityYear="1918" box="[151,536,1025,1047]" class="Gastropoda" family="Epitoniidae" genus="Amaea" kingdom="Animalia" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="accraensis">
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<emphasis id="67ABEABFFF92FFB5FF66FBF7D87AFB93" box="[151,336,1026,1047]" italics="true" pageId="45" pageNumber="46">Amaea accraensis</emphasis>
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<treatmentCitation id="D47E10BCFF92FFB5FEA7FBF7DB32FB93" author="Augener" box="[342,536,1025,1047]" page="561" pageId="45" pageNumber="46" year="1918">
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<bibRefCitation id="314E4B5CFF92FFB5FEA7FBF7D8C3FB93" author="Augener" box="[342,489,1025,1047]" pageId="45" pageNumber="46" refString="Augener, H. (1918) Polychaeta Beitr a ge zur Kenntnis des Meeresfauna West-Afrikas. Herausgegeben von W. Michaelsen Hamburg, 2, 67 - 625." type="journal article" year="1918">Augener 1918</bibRefCitation>
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: 561
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</treatmentCitation>
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</taxonomicName>
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–562, Pl. VII, Fig. 246, Text fig. ZCVIII.
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<taxonomicName id="92DF4D2EFF92FFB5FF66FBD4D843FBB2" ID-CoL="5TXLB" box="[151,361,1057,1078]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="45" pageNumber="46" phylum="Annelida" rank="species" species="accraensis">
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<emphasis id="67ABEABFFF92FFB5FF66FBD4D843FBB2" box="[151,361,1057,1078]" italics="true" pageId="45" pageNumber="46">Amaeana accraensis</emphasis>
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</taxonomicName>
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.—
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<treatmentCitation id="D47E10BCFF92FFB5FE77FBD4DB7AFBB2" author="Hartman" box="[390,592,1056,1078]" page="495" pageId="45" pageNumber="46" year="1959">
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<bibRefCitation id="314E4B5CFF92FFB5FE77FBD4DB34FBB2" author="Hartman" box="[390,542,1056,1078]" pageId="45" pageNumber="46" refString="Hartman, O. (1959) Catalogue of the Polychaetous annelids of the world. Part 2. Sedentaria. Allan Hancock Foundation Occasional Paper, 23, 355 - 628." type="journal article" year="1959">Hartman 1959</bibRefCitation>
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: 495
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="1DC56526FF92FFB5FF66FB97DA33FB47" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="556036ADFF92FFB5FF66FB97DD4AFB1B" blockId="45.[151,1437,1122,1976]" pageId="45" pageNumber="46">
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<emphasis id="67ABEABFFF92FFB5FF66FB97D8AFFBFF" bold="true" box="[151,389,1122,1147]" pageId="45" pageNumber="46">Material examined.</emphasis>
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<typeStatus id="8A64880FFF92FFB5FE60FB97D8D3FBFF" box="[401,505,1122,1147]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="33CEAE68FF92FFB5FDFDFB96DB66FBFF" box="[524,588,1123,1147]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="45" pageNumber="46">ZMH</collectionCode>
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<date id="2161106DFF92FFB5FDA9FB96DB9BFBFF" box="[600,689,1123,1147]" pageId="45" pageNumber="46" value="1710-05">
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<collectionCode id="33CEAE68FF92FFB5FDA9FB96DB5CFBFF" box="[600,630,1123,1147]" pageId="45" pageNumber="46">V-</collectionCode>
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1710
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</date>
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): incomplete specimen, in relatively good state of preservation, missing most buccal tentacles; slides: notochaetae, segment 8; neurochaetae, segment 42.
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</paragraph>
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<paragraph id="556036ADFF92FFB5FF36FB5FDA33FB47" blockId="45.[151,1437,1122,1976]" box="[199,793,1194,1219]" pageId="45" pageNumber="46">
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<emphasis id="67ABEABFFF92FFB5FF36FB5FD84FFB47" bold="true" box="[199,357,1194,1219]" pageId="45" pageNumber="46">
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<typeStatus id="8A64880FFF92FFB5FF36FB5FD828FB47" box="[199,258,1194,1219]" pageId="45" pageNumber="46">Type</typeStatus>
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locality.
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</emphasis>
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Accra,
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<collectingCountry id="2DC8763DFF92FFB5FE4CFB5FDB23FB47" box="[445,521,1194,1219]" name="Ghana" pageId="45" pageNumber="46">Ghana</collectingCountry>
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,
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<geoCoordinate id="30EB506AFF92FFB5FDE5FB5FDBA6FB47" box="[532,652,1194,1219]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="15" value="5.5333333">5°32'00"N</geoCoordinate>
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,
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<geoCoordinate id="30EB506AFF92FFB5FD69FB5EDA3FFB47" box="[664,789,1195,1219]" direction="west" orientation="longitude" pageId="45" pageNumber="46" precision="15" value="-0.2">0°12'00"W</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="1DC56526FF92FFB6FF36FB3BD851FE3F" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
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<paragraph id="556036ADFF92FFB5FF36FB3BD9FFFA8F" blockId="45.[151,1437,1122,1976]" pageId="45" pageNumber="46">
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<emphasis id="67ABEABFFF92FFB5FF36FB3BD873FB63" bold="true" box="[199,345,1230,1255]" pageId="45" pageNumber="46">Description.</emphasis>
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<typeStatus id="8A64880FFF92FFB5FE91FB3BD8E2FB63" box="[352,456,1230,1255]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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incomplete specimen, ~
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<quantity id="92279B48FF92FFB5FD2AFB3BDA6BFB63" box="[731,833,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="45" pageNumber="46" unit="mm" value="14.5">14.5 mm</quantity>
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long,
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<quantity id="92279B48FF92FFB5FC78FB3ADACAFB63" box="[905,992,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="45" pageNumber="46" unit="mm" value="2.1">2.1 mm</quantity>
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wide at segment 8, maximum width of body.
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</paragraph>
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<paragraph id="556036ADFF92FFB5FF36FAE3D9DBFA3B" blockId="45.[151,1437,1122,1976]" pageId="45" pageNumber="46">
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Prostomium at base of upper lip, both basal and distal parts developed, basal part as thickened crest, distal part with large, rounded and flaring lobes, and also rectangular to crescent-like mid-dorsal process, rounded at inferior corners; prostomium covering segment 1 laterally and terminating laterally to lower lip, near mouth (
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<figureCitation id="CDE42A28FF92FFB5FAFFFAABDC4AFAF3" box="[1294,1376,1374,1399]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
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A–D, F–I). Most buccal tentacles missing, only few remaining, relatively short and thin, uniformly cylindrical (
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<figureCitation id="CDE42A28FF92FFB5FADBFA77DC53FA1F" box="[1322,1401,1410,1435]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
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A– D, F–I).
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</paragraph>
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<paragraph id="556036ADFF92FFB5FF36FA3FDB6AF983" blockId="45.[151,1437,1122,1976]" pageId="45" pageNumber="46">
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||
Peristomium restricted to lips, upper lip almost circular, folded into three lobes; lower lip relatively large, rounded to oblong (
|
||
<figureCitation id="CDE42A28FF92FFB5FE85FA1BD8EFF983" box="[372,453,1518,1543]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
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A–D, F–I).
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</paragraph>
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<paragraph id="556036ADFF92FFB5FF36F9E7DD26F93F" blockId="45.[151,1437,1122,1976]" pageId="45" pageNumber="46">
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Body progressively broader until segments 7–8, slightly tapered on segments 9–11, then of uniform width through segment 15, more markedly tapering to narrower uniformly cylindrical posterior body, beginning from segment 20; achaetous gap between termination of notopodia and beginning of neuropodia, corresponding to segments 15 (left side of body)–16 (right side) through 20, with poorly marked segmentation and fragile, with thin body wall dorsally, distinctly shorter than region with notopodia (
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<figureCitation id="CDE42A28FF92FFB5FC84F957DAE2F93F" box="[885,968,1698,1723]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
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A–B).
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||
</paragraph>
|
||
<paragraph id="556036ADFF92FFB5FF36F932DAADF833" blockId="45.[151,1437,1122,1976]" pageId="45" pageNumber="46">
|
||
Segments biannulated, segment 1 short, visible dorsal and ventrally, laterally covered by expanded prostomium; segment 2 narrower and shorter than following segments, with relatively small, rectangular to pentagonal mid-ventral shield at beginning of mid-ventral groove, about same length as that of segment 3 and slightly narrower (
|
||
<figureCitation id="CDE42A28FF92FFB5FE9AF8C7D895F8CF" box="[363,447,1842,1867]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
|
||
A–D, F–I). Ventrum highly glandular, covered with small papillae, arranged on paired ventro-lateral pads on segments 2–15; papillae about same size throughout and slightly more numerous on anterior segments, progressively less abundant on segments 12–15, then smooth body wall, with paired longitudinal crests bordering mid-ventral groove through posterior body (
|
||
<figureCitation id="CDE42A28FF92FFB5FD08F86BDA61F833" box="[761,843,1950,1975]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
|
||
A–J).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF91FFB6FF36FF62DAD2FE90" blockId="46.[151,1437,151,710]" pageId="46" pageNumber="47">
|
||
Notopodia extending through 12 (left side of body)–13 (right side) segments, until segments 14 and 15, respectively; elongate, cylindrical notopodia, with equal sized lobes and elongate and distally blunt tips (
|
||
<figureCitation id="CDE42A28FF91FFB6FADBFF4CDC53FF56" box="[1322,1401,185,210]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="46" pageNumber="47">Fig. 26</figureCitation>
|
||
A– D, F–I). Acicular, narrowly-winged notochaetae in anterior row, wings not visible under higher magnifications of light microscopy, distally pinnate notochaetae in posterior row (
|
||
<figureCitation id="CDE42A28FF91FFB6FC93FF09DA99FE90" box="[866,947,252,277]" captionStart="FIGURE 27" captionStartId="45.[151,250,757,779]" captionTargetBox="[158,1431,197,730]" captionTargetId="figure@45.[152,1435,193,736]" captionTargetPageId="45" captionText="FIGURE 27. Amaeana accraensis holotype ZMH V- 1710. A, notochaetae, segment 8; B, notochaetae from anterior row, segment 8; C, tips of notochaetae from posterior row, segment 8; D, neuropodial spines, segment 42. Scale bars: A = 30 µm; B – D = 10 µm." httpUri="https://zenodo.org/record/289164/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
|
||
A–C).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF91FFB6FF36FEE8D8AEFEFC" blockId="46.[151,1437,151,710]" pageId="46" pageNumber="47">
|
||
Neuropodia present from segment 21, laterally to mid-ventral groove, on outer margins of longitudinal crests (
|
||
<figureCitation id="CDE42A28FF91FFB6FF6EFECBD9DBFED3" box="[159,241,318,343]" captionStart="FIGURE 26" captionStartId="44.[151,250,1830,1852]" captionTargetBox="[158,1431,197,1795]" captionTargetId="figure@44.[152,1435,193,1809]" captionTargetPageId="44" captionText="FIGURE 26. Amaeana accraensis holotype ZMH V – 1710. A – B, entire worm, right and left lateral views, respectively; arrows point to last notopodia (upper) and first neuropodia (lower). C – D, anterior end, dorsal and ventral views. E, abdominal segments. F – I, close ups of the anterior end, left dorso-lateral, right ventro-lateral, dorsal and ventral views, respectively. J, posterior end, ventral view. Numbers refer to segments; ll = lower lip, P = basal part of prostomium; PP = prostomial process; ul = upper lip; * = distal part of prostomium. Scale bars: A – B = 0.6 mm; C – E, J = 0.3 mm; F – I = 0.2 mm." httpUri="https://zenodo.org/record/289163/files/figure.png" pageId="46" pageNumber="47">Fig. 26</figureCitation>
|
||
A–B, E, J). Neurochaetae up to
|
||
<emphasis id="67ABEABFFF91FFB6FDA5FECBDB4DFED2" bold="true" box="[596,615,318,342]" pageId="46" pageNumber="47">~</emphasis>
|
||
8 relatively thin, distally tapered spines, with slightly hooked and distally blunt tips (
|
||
<figureCitation id="CDE42A28FF91FFB6FEFEFEAAD848FEFC" box="[271,354,351,376]" captionStart="FIGURE 27" captionStartId="45.[151,250,757,779]" captionTargetBox="[158,1431,197,730]" captionTargetId="figure@45.[152,1435,193,736]" captionTargetPageId="45" captionText="FIGURE 27. Amaeana accraensis holotype ZMH V- 1710. A, notochaetae, segment 8; B, notochaetae from anterior row, segment 8; C, tips of notochaetae from posterior row, segment 8; D, neuropodial spines, segment 42. Scale bars: A = 30 µm; B – D = 10 µm." httpUri="https://zenodo.org/record/289164/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
|
||
D).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF91FFB6FF36FE74D851FE3F" blockId="46.[151,1437,151,710]" pageId="46" pageNumber="47">Nephridial and genital papillae ventral and slightly anterior to bases of all notopodia, all relatively small. Pygidium unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1DC56526FF91FFB6FF36FE31D9CBFD42" pageId="46" pageNumber="47" type="discussion">
|
||
<paragraph id="556036ADFF91FFB6FF36FE31DD13FD9B" blockId="46.[151,1437,151,710]" pageId="46" pageNumber="47">
|
||
<emphasis id="67ABEABFFF91FFB6FF36FE31D811FE59" bold="true" box="[199,315,452,477]" pageId="46" pageNumber="47">Remarks.</emphasis>
|
||
Members of
|
||
<taxonomicName id="92DF4D2EFF91FFB6FE2DFE31DB5DFE58" box="[476,631,452,476]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="accraensis">
|
||
<emphasis id="67ABEABFFF91FFB6FE2DFE31DB5DFE58" box="[476,631,452,476]" italics="true" pageId="46" pageNumber="47">A. accraensis</emphasis>
|
||
</taxonomicName>
|
||
are unique among all species of
|
||
<taxonomicName id="92DF4D2EFF91FFB6FBF1FE31DD40FE5F" box="[1024,1130,452,475]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="genus">
|
||
<emphasis id="67ABEABFFF91FFB6FBF1FE31DD40FE5F" box="[1024,1130,452,475]" italics="true" pageId="46" pageNumber="47">Amaeana</emphasis>
|
||
</taxonomicName>
|
||
in having distally pinnate chaetae in posterior row of notochaetae, instead of acicular narrowly-winged chaetae.
|
||
<taxonomicName id="92DF4D2EFF91FFB6FB81FE13DC1DFE79" box="[1136,1335,485,509]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="hsiehae" status="sp. nov.">
|
||
<emphasis id="67ABEABFFF91FFB6FB81FE13DC1DFE79" box="[1136,1335,485,509]" italics="true" pageId="46" pageNumber="47">Amaeana hsiehae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="67ABEABFFF91FFB6FAB0FE10DCB6FE7A" bold="true" box="[1345,1436,485,510]" pageId="46" pageNumber="47">
|
||
<taxonomicNameLabel id="7C9857C4FF91FFB6FAB0FE10DCB6FE7A" box="[1345,1436,485,510]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also has pinnate chaetae, but in this species they are present in both rows (
|
||
<tableCitation id="185D0316FF91FFB6FC25FDF3DD02FD9B" box="[980,1064,518,543]" captionStart="TABLE 1" captionStartId="4.[119,199,561,582]" captionText="TABLE 1. Comparison of selected morphological characters between species of Amaeana." pageId="46" pageNumber="47">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="556036ADFF91FFB6FF36FDD2D9CBFD42" blockId="46.[151,1437,151,710]" pageId="46" pageNumber="47">
|
||
Besides, although it is not clear if 12 or 13 pairs of notopodia are present, as the
|
||
<typeStatus id="8A64880FFF91FFB6FB7EFDD2DDDBFDC4" box="[1167,1265,551,576]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
have different numbers from right to left sides of the body, this is a large number of pairs for a species of
|
||
<taxonomicName id="92DF4D2EFF91FFB6FB4CFDBCDC0DFDE4" box="[1213,1319,585,608]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="genus">
|
||
<emphasis id="67ABEABFFF91FFB6FB4CFDBCDC0DFDE4" box="[1213,1319,585,608]" italics="true" pageId="46" pageNumber="47">Amaeana</emphasis>
|
||
</taxonomicName>
|
||
. No other species has 13 pairs, and only two others have 12 pairs,
|
||
<taxonomicName id="92DF4D2EFF91FFB6FCEDFD9EDAEEFD07" box="[796,964,619,643]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF91FFB6FCEDFD9EDAEEFD07" box="[796,964,619,643]" italics="true" pageId="46" pageNumber="47">A. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="92DF4D2EFF91FFB6FC0FFD9EDD5AFD06" box="[1022,1136,619,642]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="yirrarn">
|
||
<emphasis id="67ABEABFFF91FFB6FC0FFD9EDD5AFD06" box="[1022,1136,619,642]" italics="true" pageId="46" pageNumber="47">A. yirrarn</emphasis>
|
||
</taxonomicName>
|
||
, as already discussed (see above). The differences between members of
|
||
<taxonomicName id="92DF4D2EFF91FFB6FD69FD79DA05FD20" box="[664,815,652,676]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="accraensis" status="sp. nov.">
|
||
<emphasis id="67ABEABFFF91FFB6FD69FD79DA05FD20" box="[664,815,652,676]" italics="true" pageId="46" pageNumber="47">A. accraensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="67ABEABFFF91FFB6FCC7FD79DABAFD21" bold="true" box="[822,912,652,677]" pageId="46" pageNumber="47">
|
||
<taxonomicNameLabel id="7C9857C4FF91FFB6FCC7FD79DABAFD21" box="[822,912,652,677]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="92DF4D2EFF91FFB6FC6FFD79DD6EFD20" box="[926,1092,652,676]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="occidentalis">
|
||
<emphasis id="67ABEABFFF91FFB6FC6FFD79DD6EFD20" box="[926,1092,652,676]" italics="true" pageId="46" pageNumber="47">A. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="92DF4D2EFF91FFB6FB8DFD79DDC6FD20" box="[1148,1260,652,676]" class="Polychaeta" family="Terebellidae" genus="Amaeana" kingdom="Animalia" order="Terebellida" pageId="46" pageNumber="47" phylum="Annelida" rank="species" species="yirrarn">
|
||
<emphasis id="67ABEABFFF91FFB6FB8DFD79DDC6FD20" box="[1148,1260,652,676]" italics="true" pageId="46" pageNumber="47">A. yirrarn</emphasis>
|
||
</taxonomicName>
|
||
were discussed above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |