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<mods:title id="CD6BCCF453A0A964E8AF4FA2C0CE27DC">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<taxonomicName id="9BEF3AD6FFFAFFD28C92FC31FDEC03BA" ID-CoL="8KSZT" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[151,613,927,954]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="boehmi" status="comb. nov.">
<emphasis id="6E9B9D47FFFAFFD28C92FC31FE1103BA" bold="true" box="[151,408,927,954]" italics="true" pageId="15" pageNumber="16">Bellardithala boehmi</emphasis>
(
<bibRefCitation id="387E3CA4FFFAFFD28DA2FC0EFDD503BA" author="Boettger, O." box="[423,604,928,954]" pageId="15" pageNumber="16" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
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<taxonomicNameLabel id="75A8203CFFFAFFD28E69FC0EFD7B03BA" box="[620,754,928,954]" pageId="15" pageNumber="16" rank="species">nov. comb.</taxonomicNameLabel>
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<subSubSection id="14F512DEFFFAFFD28C92FC6AFE4A03E4" box="[151,451,964,996]" pageId="15" pageNumber="16" type="description">
<paragraph id="5C504155FFFAFFD28C92FC6AFE4A03E4" blockId="15.[151,754,927,996]" box="[151,451,964,996]" pageId="15" pageNumber="16">
<figureCitation id="C4D45DD0FFFAFFD28C92FC6AFF7803DE" box="[151,241,964,990]" captionStart="FIGURE 6" captionStartId="16.[151,250,1717,1741]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-19@16.[151,1436,181,1693]" captionTargetPageId="16" captionText="FIGURE 6. A1A2. Bellardithala boehmi (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360342. B1B2. Bellardithala boehmi (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0059/0001. C1C2. Bellardithala boehmi (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 1854/0035/0107. D1D2. Bellardithala kovaci nov. sp., holotype, Steinebrunn (Austria), NHMW 1846/0037/0114. E1E2. Bellardithala kovaci nov. sp., paratype, Sedlec (= Porzteich) (Czech Republic), NHMW 1858/0047/0007. F1F2. Bellardithala kovaci nov. sp., paratype, Forchtenau (Austria), NHMW 1866/0001/1019. G1G2. Bellardithala kovaci nov. sp., paratype, Steinebrunn (Austria), NHMW 1871/0010/0331. H1H2. Bellardithala kovaci nov. sp., paratype, Steinebrunn (Austria), NHMW 1871/0011/0004. I1I2. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001." figureDoi="http://doi.org/10.5281/zenodo.4921854" httpUri="https://zenodo.org/record/4921854/files/figure.png" pageId="15" pageNumber="16">Figs 6A</figureCitation>
<figureCitation id="C4D45DD0FFFAFFD28CF4FC78FE9203DE" box="[241,283,964,996]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="15" pageNumber="16">
<subScript id="C06B4310FFFAFFD28CF4FC78FF7003E4" attach="left" box="[241,249,982,996]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
A
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<figureCitation id="C4D45DD0FFFAFFD28D1EFC78FEAA03E4" box="[283,291,982,996]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="15" pageNumber="16">
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, B
<figureCitation id="C4D45DD0FFFAFFD28D41FC78FEE503DF" box="[324,364,965,996]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="15" pageNumber="16">
<subScript id="C06B4310FFFAFFD28D41FC78FEC503E4" attach="left" box="[324,332,982,996]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
B
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, C
<figureCitation id="C4D45DD0FFFAFFD28D96FC78FE3503DE" box="[403,444,964,996]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="15" pageNumber="16">
<subScript id="C06B4310FFFAFFD28D96FC78FE1203E4" attach="left" box="[403,411,982,996]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
C
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<figureCitation id="C4D45DD0FFFAFFD28DBEFC78FE4A03E4" box="[443,451,982,996]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="15" pageNumber="16">
<subScript id="C06B4310FFFAFFD28DBEFC78FE4A03E4" attach="left" box="[443,451,982,996]" fontSize="6" pageId="15" pageNumber="16">2</subScript>
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</paragraph>
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<subSubSection id="14F512DEFFFAFFD28C92FBA7FCB3043F" pageId="15" pageNumber="16" type="reference_group">
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<treatmentCitationGroup id="7CFF667BFFFAFFD28C92FBA7FD5B0421" box="[151,722,1032,1057]" pageId="15" pageNumber="16">
<emphasis id="6E9B9D47FFFAFFD28C92FBA7FF2D0420" box="[151,164,1033,1056]" italics="true" pageId="15" pageNumber="16">*</emphasis>
<taxonomicName id="9BEF3AD6FFFAFFD28CAFFBA7FE490420" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[170,448,1032,1057]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="boehmi" status="sp. nov." subGenus="Micromitra">
<emphasis id="6E9B9D47FFFAFFD28CAFFBA7FF6D0420" box="[170,228,1033,1056]" italics="true" pageId="15" pageNumber="16">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFFAFFD28CF6FBA7FEEE0420" box="[243,359,1033,1056]" italics="true" pageId="15" pageNumber="16">Micromitra</emphasis>
)
<emphasis id="6E9B9D47FFFAFFD28D70FBA6FE490420" box="[373,448,1032,1056]" italics="true" pageId="15" pageNumber="16">boehmi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFFAFFD28DC3FBA7FE700421" box="[454,505,1033,1057]" pageId="15" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
<treatmentCitation id="DD4E6744FFFAFFD28E16FBA7FD470421" author="Boettger, O." box="[531,718,1032,1057]" page="11" pageId="15" pageNumber="16" year="1906">
<bibRefCitation id="387E3CA4FFFAFFD28E16FBA7FD470421" author="Boettger, O." box="[531,718,1032,1057]" pageId="15" pageNumber="16" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger 1906: 11</bibRefCitation>
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.
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</paragraph>
<paragraph id="5C504155FFFAFFD28C92FB86FCB3043F" blockId="15.[151,826,1032,1087]" box="[151,826,1063,1087]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="7CFF667BFFFAFFD28C92FB86FCB3043F" box="[151,826,1063,1087]" pageId="15" pageNumber="16">
<taxonomicName id="9BEF3AD6FFFAFFD28C92FB86FE68043F" authority="(Boettger)" baseAuthorityName="Boettger" box="[151,481,1063,1087]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="boehmi" subGenus="Thala">
<emphasis id="6E9B9D47FFFAFFD28C92FB86FF58043F" box="[151,209,1064,1087]" italics="true" pageId="15" pageNumber="16">Pusia</emphasis>
(
<emphasis id="6E9B9D47FFFAFFD28CE5FB89FE92043F" box="[224,283,1063,1087]" italics="true" pageId="15" pageNumber="16">Thala</emphasis>
)
<emphasis id="6E9B9D47FFFAFFD28D2FFB89FEFC043F" box="[298,373,1063,1087]" italics="true" pageId="15" pageNumber="16">boehmi</emphasis>
(Boettger)
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<treatmentCitation id="DD4E6744FFFAFFD28DFEFB89FD29043F" author="Zilch, A." box="[507,672,1063,1087]" page="258" pageId="15" pageNumber="16" year="1934">
<bibRefCitation id="387E3CA4FFFAFFD28DFEFB89FD29043F" author="Zilch, A." box="[507,672,1063,1087]" pageId="15" pageNumber="16" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch 1934: 258</bibRefCitation>
</treatmentCitation>
, pl. 17, fig. 14.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFAFFD28C92FBC3FE2B06C8" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="5C504155FFFAFFD28C92FBC3FAB204AC" blockId="15.[151,1437,1133,1988]" pageId="15" pageNumber="16">
<emphasis id="6E9B9D47FFFAFFD28C92FBC3FECC0487" bold="true" box="[151,325,1133,1159]" pageId="15" pageNumber="16">Type material.</emphasis>
<materialsCitation id="EC874B08FFFAFFD28D4EFBC0FAB204AC" ID-GBIF-Occurrence="3311814331" collectingDate="2217-12" collectionCode="SMF" collectorName="de Sus" country="Germany" location="Costeiu de Sus" pageId="15" pageNumber="16" specimenCode="SMF 360342" specimenCount="1" typeStatus="lectotype">
<typeStatus id="8354FFF7FFFAFFD28D4EFBC0FE360488" box="[331,447,1134,1160]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
:
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(=
<collectionCode id="3AFED990FFFAFFD28E87FBC3FD330487" box="[642,698,1133,1159]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="15" pageNumber="16">SMF</collectionCode>
<date id="28516795FFFAFFD28EC5FBC3FCA30487" box="[704,810,1133,1159]" pageId="15" pageNumber="16" value="2217-12">
<collectingDate id="38159E7DFFFAFFD28EC5FBC3FCA30487" box="[704,810,1133,1159]" pageId="15" pageNumber="16" value="2217-12">XII.2217</collectingDate>
</date>
a), SL:
<quantity id="9B17ECB0FFFAFFD28F7CFBC3FC590488" box="[889,976,1133,1160]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="15" pageNumber="16" unit="mm" value="6.4">6.4 mm</quantity>
, MD:
<quantity id="9B17ECB0FFFAFFD28812FBC3FBE70488" box="[1047,1134,1133,1160]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="16" unit="mm" value="2.3">2.3 mm</quantity>
, illustrated by
<bibRefCitation id="387E3CA4FFFAFFD28917FBC3FA110487" author="Zilch, A." box="[1298,1432,1133,1159]" pageId="15" pageNumber="16" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch (1934</bibRefCitation>
, pl. 22, fig. 14), Senckenberg Museum, Frankfurt/Main,
<collectingCountry id="24F801C5FFFAFFD28F02FB3FFCF904AB" box="[775,880,1169,1195]" name="Germany" pageId="15" pageNumber="16">Germany</collectingCountry>
,
<location id="5930178EFFFAFFD28F7EFB3FFBA804AB" LSID="urn:lsid:plazi:treatment:D446F043FFFAFFCC8C92FC31FE77011D:5930178EFFFAFFD28F7EFB3FFBA804AB" box="[891,1057,1169,1195]" country="Germany" name="Costeiu de Sus" pageId="15" pageNumber="16">
Coşteiu
<collectorName id="F11A2483FFFAFFD28FDDFB3FFBA804AB" box="[984,1057,1169,1195]" pageId="15" pageNumber="16">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFAFFD28834FB3FFB1504AC" box="[1073,1180,1169,1196]" name="Romania" pageId="15" pageNumber="16">Romania</collectingCountry>
), figs 6A
<subScript id="C06B4310FFFAFFD28907FB0AFA8304B2" attach="left" box="[1282,1290,1188,1202]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
A
<subScript id="C06B4310FFFAFFD28929FB0AFABD04B2" attach="left" box="[1324,1332,1188,1202]" fontSize="6" pageId="15" pageNumber="16">2</subScript>
.
</materialsCitation>
</paragraph>
<paragraph id="5C504155FFFAFFD28CC2FB1BFAE40518" blockId="15.[151,1437,1133,1988]" pageId="15" pageNumber="16">
<emphasis id="6E9B9D47FFFAFFD28CC2FB1BFE3504CF" bold="true" box="[199,444,1205,1231]" pageId="15" pageNumber="16">Additional material.</emphasis>
<materialsCitation id="EC874B08FFFAFFD28DC0FB18FF6004FA" ID-GBIF-Occurrence="3311814346" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="15" pageNumber="16" specimenCode="NHMW 2020" specimenCount="1">
<specimenCode id="0C49E92EFFFAFFD28DC0FB18FDEB04D0" box="[453,610,1205,1232]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="16" type="Museum">NHMW 2020</specimenCode>
/0059/0001, SL:
<quantity id="9B17ECB0FFFAFFD28F27FB1BFCF504D0" box="[802,892,1205,1232]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="15" pageNumber="16" unit="mm" value="7.9">7.9 mm</quantity>
, MD:
<quantity id="9B17ECB0FFFAFFD28FCDFB1BFBAB04D0" box="[968,1058,1205,1232]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="16" unit="mm" value="2.8">2.8 mm</quantity>
,
<location id="5930178EFFFAFFD2882AFB1BFB6804CF" LSID="urn:lsid:plazi:treatment:D446F043FFFAFFCC8C92FC31FE77011D:5930178EFFFAFFD2882AFB1BFB6804CF" box="[1071,1249,1205,1231]" country="Romania" name="Lapugiu de Sus" pageId="15" pageNumber="16">
Lăpugiu
<collectorName id="F11A2483FFFAFFD28893FB1BFB6804CF" box="[1174,1249,1205,1231]" pageId="15" pageNumber="16">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFAFFD288F7FB1BFAD404D0" box="[1266,1373,1205,1232]" name="Romania" pageId="15" pageNumber="16">Romania</collectingCountry>
), figs 6B
<subScript id="C06B4310FFFAFFD28CBDFB42FF4904FA" attach="left" box="[184,192,1260,1274]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
B
<subScript id="C06B4310FFFAFFD28CE4FB42FF6004FA" attach="right" box="[225,233,1260,1274]" fontSize="6" pageId="15" pageNumber="16">2</subScript>
</materialsCitation>
;
<materialsCitation id="EC874B08FFFAFFD28CF3FB74FA1004F3" ID-GBIF-Occurrence="3311814327" box="[246,1433,1241,1274]" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="15" pageNumber="16" specimenCode="NHMW 1854" specimenCount="7">
<specimenCode id="0C49E92EFFFAFFD28CF3FB74FE0604F4" box="[246,399,1241,1268]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="16" type="Museum">NHMW 1854</specimenCode>
/0035/0107, SL:
<quantity id="9B17ECB0FFFAFFD28E4FFB77FD2804F4" box="[586,673,1241,1268]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="15" pageNumber="16" unit="mm" value="7.9">7.9 mm</quantity>
, MD:
<quantity id="9B17ECB0FFFAFFD28EE3FB77FCB404F4" box="[742,829,1241,1268]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="16" unit="mm" value="2.7">2.7 mm</quantity>
,
<location id="5930178EFFFAFFD28F42FB77FC7A04F3" LSID="urn:lsid:plazi:treatment:D446F043FFFAFFCC8C92FC31FE77011D:5930178EFFFAFFD28F42FB77FC7A04F3" box="[839,1011,1241,1267]" country="Romania" name="Lapugiu de Sus" pageId="15" pageNumber="16">
Lăpugiu
<collectorName id="F11A2483FFFAFFD28FAEFB77FC7A04F3" box="[939,1011,1241,1267]" pageId="15" pageNumber="16">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFAFFD28804FB77FBE504F4" box="[1025,1132,1241,1268]" name="Romania" pageId="15" pageNumber="16">Romania</collectingCountry>
), figs 6C
<subScript id="C06B4310FFFAFFD288CBFB42FB5F04FA" attach="left" box="[1230,1238,1260,1274]" fontSize="6" pageId="15" pageNumber="16">1</subScript>
C
<subScript id="C06B4310FFFAFFD288F2FB42FB7604FA" attach="left" box="[1271,1279,1260,1274]" fontSize="6" pageId="15" pageNumber="16">2</subScript>
;
<specimenCount id="4AE98ADCFFFAFFD28908FB74FA1004F3" box="[1293,1433,1241,1268]" pageId="15" pageNumber="16" type="generic">7 specimens</specimenCount>
</materialsCitation>
,
<materialsCitation id="EC874B08FFFAFFD28C92FB50FAE00518" ID-GBIF-Occurrence="3311814326" box="[151,1385,1277,1304]" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" municipality="Lapugiu de Sus" pageId="15" pageNumber="16" specimenCode="NHMW 2020" specimenCount="4">
<specimenCode id="0C49E92EFFFAFFD28C92FB50FEBB0518" box="[151,306,1277,1304]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="16" type="Museum">NHMW 2020</specimenCode>
/0059/0002,
<location id="5930178EFFFAFFD28DBAFB53FDE40517" LSID="urn:lsid:plazi:treatment:D446F043FFFAFFCC8C92FC31FE77011D:5930178EFFFAFFD28DBAFB53FDE40517" box="[447,621,1277,1303]" country="Romania" municipality="Lapugiu de Sus" name="Lapugiu de Sus" pageId="15" pageNumber="16">
Lăpugiu
<collectorName id="F11A2483FFFAFFD28E26FB53FDE40517" box="[547,621,1277,1303]" pageId="15" pageNumber="16">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFAFFD28E79FB53FD610518" box="[636,744,1277,1304]" name="Romania" pageId="15" pageNumber="16">Romania</collectingCountry>
); BL collection,
<specimenCount id="4AE98ADCFFFAFFD28FA4FB53FBA70517" box="[929,1070,1277,1303]" pageId="15" pageNumber="16" type="generic">4 specimens</specimenCount>
,
<collectingMunicipality id="BC34DB2FFFFAFFD2883CFB53FB6F0517" box="[1081,1254,1277,1303]" pageId="15" pageNumber="16">
Lăpugiu
<collectorName id="F11A2483FFFAFFD28898FB53FB6F0517" box="[1181,1254,1277,1303]" pageId="15" pageNumber="16">de Sus</collectorName>
</collectingMunicipality>
(
<collectingCountry id="24F801C5FFFAFFD288F3FB53FAEB0518" box="[1270,1378,1277,1304]" name="Romania" pageId="15" pageNumber="16">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFFAFFD28CC2FA8FFF54067F" blockId="15.[151,1437,1133,1988]" pageId="15" pageNumber="16">
<emphasis id="6E9B9D47FFFAFFD28CC2FA8FFE3D053B" bold="true" box="[199,436,1313,1339]" pageId="15" pageNumber="16">Revised description</emphasis>
. Shell medium-sized, moderately slender fusiform with conical spire. Protoconch high mammillate of three smooth, weakly convex whorls; height: 600 μm, diameter: 550 μm. Teleoconch of 5.5 weakly convex whorls with periphery slightly above abapical suture. Suture narrowly incised. First three teleoconch whorls with, convex axial ribs, separated by narrower interspaces. Axial sculpture intersected by about 8 spiral grooves. Sculpture weakening abruptly after third whorl. Penultimate whorl nearly smooth; last whorl with faint spiral cords. Prominent spiral cords on base and fasciole with delicate axial growth lines in interspaces. Aperture moderately narrow, elongate with narrow anal sinus; inner lip narrow. Columella with four prominent folds (adapical two being stronger). Outer lip thickened with 1012 small denticles, becoming nearly obsolete in abapical third of aperture. Last whorl convex, moderately constricted at base. Siphonal canal moderately short, straight with shallow anterior notch.
</paragraph>
<paragraph id="5C504155FFFAFFD28CC2F927FE2B06C8" blockId="15.[151,1437,1133,1988]" pageId="15" pageNumber="16">
<emphasis id="6E9B9D47FFFAFFD28CC2F927FDB406A3" bold="true" box="[199,573,1673,1699]" pageId="15" pageNumber="16">Shell measurements and ratios.</emphasis>
SL:
<quantity id="9B17ECB0FFFAFFD28E76F927FD7406A4" box="[627,765,1673,1700]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.9" metricValueMin="6.4" pageId="15" pageNumber="16" unit="mm" value="7.15" valueMax="7.9" valueMin="6.4">6.47.9 mm</quantity>
, MD:
<quantity id="9B17ECB0FFFAFFD28F40F927FC4706A4" box="[837,974,1673,1700]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="15" pageNumber="16" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.32.8 mm</quantity>
, AA= 3742°, SL/MD: 2.73.0, AL/AW: 4.45.0, AH/S: 2.52.7.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFAFFCC8CC2F97FFE5C00D4" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="discussion">
<paragraph id="5C504155FFFAFFD28CC2F97FFE0E0757" blockId="15.[151,1437,1133,1988]" pageId="15" pageNumber="16">
<emphasis id="6E9B9D47FFFAFFD28CC2F97FFEC506EB" bold="true" box="[199,332,1745,1771]" pageId="15" pageNumber="16">Discussion.</emphasis>
This rare species is reminiscent of
<taxonomicName id="9BEF3AD6FFFAFFD28EE0F97FFCF106EB" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[741,888,1745,1771]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFFAFFD28EE0F97FFCF106EB" box="[741,888,1745,1771]" italics="true" pageId="15" pageNumber="16">Thala kovaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFFAFFD28F84F97CFC5006EC" box="[897,985,1746,1772]" pageId="15" pageNumber="16" rank="species">nov. sp.</taxonomicNameLabel>
(=
<taxonomicName id="9BEF3AD6FFFAFFD28806F97FFA1106EC" authority="sensu Hornes, 1852" authorityName="Hornes" authorityYear="1852" box="[1027,1432,1745,1772]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFFAFFD28806F97FFB2406EB" box="[1027,1197,1745,1771]" italics="true" pageId="15" pageNumber="16">Thala obsoleta</emphasis>
sensu Hörnes, 1852
</taxonomicName>
, non
<bibRefCitation id="387E3CA4FFFAFFD28CCCF95BFEE3070F" author="Brocchi, G." box="[201,362,1781,1807]" pageId="15" pageNumber="16" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
) due to its reduced sculpture on the last whorl, but differs in its more slender fusiform outline, the much more prominent axial and spiral sculpture on the early teleoconch whorls, and the lower aperture/spire ratio (0.9 versus 1.1).
</paragraph>
<paragraph id="5C504155FFFAFFD28CC2F8CFFED407C4" blockId="15.[151,1437,1133,1988]" pageId="15" pageNumber="16">
<taxonomicName id="9BEF3AD6FFFAFFD28CC2F8CFFD03077B" authority="(Bellardi, 1888)" baseAuthorityName="Bellardi" baseAuthorityYear="1888" box="[199,650,1889,1915]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="seminuda">
<emphasis id="6E9B9D47FFFAFFD28CC2F8CFFE44077B" box="[199,461,1889,1915]" italics="true" pageId="15" pageNumber="16">Bellardithala seminuda</emphasis>
(
<bibRefCitation id="387E3CA4FFFAFFD28DD9F8CFFD08077B" author="Bellardi, L." box="[476,641,1889,1915]" pageId="15" pageNumber="16" refId="ref54491" refString="Bellardi, L. (1888) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte 5; Mitridae (fine). Ermanno Loescher, Torino, 54 pp. [https: // www. biodiversitylibrary. org / item / 45379 # page / 195 / mode / 1 up]" type="book" year="1888">Bellardi, 1888</bibRefCitation>
)
</taxonomicName>
, from the Burdigalian of the Colli Torinesi (
<collectingCountry id="24F801C5FFFAFFD28872F8CFFB26077C" box="[1143,1199,1889,1916]" name="Italy" pageId="15" pageNumber="16">Italy</collectingCountry>
), is reminiscent of
<taxonomicName id="9BEF3AD6FFFAFFD28981F8CCFF60079F" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="boehmi">
<emphasis id="6E9B9D47FFFAFFD28981F8CCFF60079F" italics="true" pageId="15" pageNumber="16">B. boehmi</emphasis>
</taxonomicName>
, but differs in its larger size and in the absence of spiral sculpture on the last whorl (see
<bibRefCitation id="387E3CA4FFFAFFD288D8F82BFA1E07A0" author="Bellardi, L." box="[1245,1431,1925,1952]" pageId="15" pageNumber="16" refId="ref54491" refString="Bellardi, L. (1888) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte 5; Mitridae (fine). Ermanno Loescher, Torino, 54 pp. [https: // www. biodiversitylibrary. org / item / 45379 # page / 195 / mode / 1 up]" type="book" year="1888">Bellardi 1888: 7</bibRefCitation>
, pl. 6, figs 46ab).
</paragraph>
<caption id="089011DDFFE5FFCD8C92F91BFECC07C9" ID-DOI="http://doi.org/10.5281/zenodo.4921854" ID-Zenodo-Dep="4921854" httpUri="https://zenodo.org/record/4921854/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1717,1741]" targetBox="[151,1436,181,1693]" targetPageId="16">
<paragraph id="5C504155FFE5FFCD8C92F91BFECC07C9" blockId="16.[151,1437,1717,1993]" pageId="16" pageNumber="17">
<emphasis id="6E9B9D47FFE5FFCD8C92F91BFED606CD" bold="true" box="[151,351,1717,1747]" pageId="16" pageNumber="17">
FIGURE 6. A
<subScript id="C06B4310FFE5FFCD8D2EF96BFEBB06D3" attach="left" box="[299,306,1733,1747]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
A
<subScript id="C06B4310FFE5FFCD8D54F96BFED106D3" attach="left" box="[337,344,1733,1747]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD8D63F91BFD6506CD" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[358,748,1717,1741]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="boehmi">
<emphasis id="6E9B9D47FFE5FFCD8D63F91BFDB406CD" box="[358,573,1717,1741]" italics="true" pageId="16" pageNumber="17">Bellardithala boehmi</emphasis>
(
<bibRefCitation id="387E3CA4FFE5FFCD8E4EF91BFD6C06CD" author="Boettger, O." box="[587,741,1717,1741]" pageId="16" pageNumber="17" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
, lectotype, Coşteiu de Sus (Romania), SMF 360342.
<emphasis id="6E9B9D47FFE5FFCD890EF91BFAC706CD" bold="true" box="[1291,1358,1717,1747]" pageId="16" pageNumber="17">
B
<subScript id="C06B4310FFE5FFCD8919F96BFAAA06D3" attach="left" box="[1308,1315,1733,1747]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
B
<subScript id="C06B4310FFE5FFCD8944F96BFAC106D3" attach="right" box="[1345,1352,1733,1747]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD8950F91BFE5406F1" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="boehmi">
<emphasis id="6E9B9D47FFE5FFCD8950F91BFEB906F1" italics="true" pageId="16" pageNumber="17">Bellardithala boehmi</emphasis>
(
<bibRefCitation id="387E3CA4FFE5FFCD8D39F977FE5F06F1" author="Boettger, O." box="[316,470,1753,1777]" pageId="16" pageNumber="17" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
, Lăpugiu de Sus (Romania), NHMW 2020/0059/0001.
<emphasis id="6E9B9D47FFE5FFCD880BF977FBDD06F1" bold="true" box="[1038,1108,1753,1783]" pageId="16" pageNumber="17">
C
<subScript id="C06B4310FFE5FFCD8824F947FBA106F7" attach="right" box="[1057,1064,1769,1783]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
C
<subScript id="C06B4310FFE5FFCD8842F947FBC706F7" attach="left" box="[1095,1102,1769,1783]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD885EF977FF5D0715" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="boehmi">
<emphasis id="6E9B9D47FFE5FFCD885EF977FABB06F1" box="[1115,1330,1753,1777]" italics="true" pageId="16" pageNumber="17">Bellardithala boehmi</emphasis>
(
<bibRefCitation id="387E3CA4FFE5FFCD893BF977FF440715" author="Boettger, O." pageId="16" pageNumber="17" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
, Lăpugiu de Sus (Romania), NHMW 1854/0035/0107.
<emphasis id="6E9B9D47FFE5FFCD8F01F953FCC30715" bold="true" box="[772,842,1789,1819]" pageId="16" pageNumber="17">
D
<subScript id="C06B4310FFE5FFCD8F13F8A3FC94071B" attach="left" box="[790,797,1805,1819]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
D
<subScript id="C06B4310FFE5FFCD8F38F8A3FCCD071B" attach="left" box="[829,836,1805,1819]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD8F55F953FB970715" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[848,1054,1789,1813]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFE5FFCD8F55F953FB970715" box="[848,1054,1789,1813]" italics="true" pageId="16" pageNumber="17">Bellardithala kovaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFE5FFCD8821F950FBFA0715" box="[1060,1139,1790,1813]" pageId="16" pageNumber="17" rank="species">nov. sp.</taxonomicNameLabel>
, holotype, Steinebrunn (Austria), NHMW 1846/0037/0114.
<emphasis id="6E9B9D47FFE5FFCD8DDCF88FFD950739" bold="true" box="[473,540,1825,1855]" pageId="16" pageNumber="17">
E
<subScript id="C06B4310FFE5FFCD8DEFF89FFE78073F" attach="left" box="[490,497,1841,1855]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
E
<subScript id="C06B4310FFE5FFCD8E0AF89FFD9F073F" attach="right" box="[527,534,1841,1855]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD8E26F88FFD780739" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[547,753,1825,1849]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFE5FFCD8E26F88FFD780739" box="[547,753,1825,1849]" italics="true" pageId="16" pageNumber="17">Bellardithala kovaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFE5FFCD8EF2F88CFCCE0739" box="[759,839,1826,1849]" pageId="16" pageNumber="17" rank="species">nov. sp.</taxonomicNameLabel>
, paratype, Sedlec (= Porzteich) (Czech Republic), NHMW 1858/0047/0007.
<emphasis id="6E9B9D47FFE5FFCD8D4EF8EBFE05075D" bold="true" box="[331,396,1861,1891]" pageId="16" pageNumber="17">
F
<subScript id="C06B4310FFE5FFCD8D5EF8FBFEEB0763" attach="right" box="[347,354,1877,1891]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
F
<subScript id="C06B4310FFE5FFCD8D7BF8FBFE0C0763" attach="left" box="[382,389,1877,1891]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD8D91F8EBFDED075D" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[404,612,1861,1885]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFE5FFCD8D91F8EBFDED075D" box="[404,612,1861,1885]" italics="true" pageId="16" pageNumber="17">Bellardithala kovaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFE5FFCD8E69F8E8FD34075D" box="[620,701,1862,1885]" pageId="16" pageNumber="17" rank="species">nov. sp.</taxonomicNameLabel>
, paratype, Forchtenau (Austria), NHMW 1866/0001/1019.
<emphasis id="6E9B9D47FFE5FFCD8924F8EBFAE3075D" bold="true" box="[1313,1386,1861,1891]" pageId="16" pageNumber="17">
G
<subScript id="C06B4310FFE5FFCD8930F8FBFAB50763" attach="left" box="[1333,1340,1877,1891]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
G
<subScript id="C06B4310FFE5FFCD8959F8FBFAEA0763" attach="left" box="[1372,1379,1877,1891]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD8977F8EBFECE0781" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFE5FFCD8977F8EBFECE0781" italics="true" pageId="16" pageNumber="17">Bellardithala kovaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFE5FFCD8D55F8C4FE2A0781" box="[336,419,1898,1921]" pageId="16" pageNumber="17" rank="species">nov. sp.</taxonomicNameLabel>
, paratype, Steinebrunn (Austria), NHMW 1871/0010/0331.
<emphasis id="6E9B9D47FFE5FFCD881DF8C7FBE80781" bold="true" box="[1048,1121,1897,1927]" pageId="16" pageNumber="17">
H
<subScript id="C06B4310FFE5FFCD882EF8D7FBBB0787" attach="left" box="[1067,1074,1913,1927]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
H
<subScript id="C06B4310FFE5FFCD8856F8D7FBD30787" attach="left" box="[1107,1114,1913,1927]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD886FF8C7FAB50781" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[1130,1340,1897,1921]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFE5FFCD886FF8C7FAB50781" box="[1130,1340,1897,1921]" italics="true" pageId="16" pageNumber="17">Bellardithala kovaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFE5FFCD8940F8C4FA110781" box="[1349,1432,1898,1921]" pageId="16" pageNumber="17" rank="species">nov. sp.</taxonomicNameLabel>
, paratype, Steinebrunn (Austria), NHMW 1871/0011/0004.
<emphasis id="6E9B9D47FFE5FFCD8EF4F823FCAE07A5" bold="true" box="[753,807,1933,1963]" pageId="16" pageNumber="17">
I
<subScript id="C06B4310FFE5FFCD8EFEF833FC8B07AB" attach="left" box="[763,770,1949,1963]" fontSize="6" pageId="16" pageNumber="17">1</subScript>
I
<subScript id="C06B4310FFE5FFCD8F1CF833FCA907AB" attach="left" box="[793,800,1949,1963]" fontSize="6" pageId="16" pageNumber="17">2</subScript>
.
</emphasis>
<taxonomicName id="9BEF3AD6FFE5FFCD8F2BF823FC7407A5" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[814,1021,1933,1957]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFE5FFCD8F2BF823FC7407A5" box="[814,1021,1933,1957]" italics="true" pageId="16" pageNumber="17">Bellardithala kovaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFE5FFCD8801F820FBDD07A5" box="[1028,1108,1934,1957]" pageId="16" pageNumber="17" rank="species">nov. sp.</taxonomicNameLabel>
, Steinebrunn (Austria), NHMW 2020/0062/0001.
</paragraph>
</caption>
<paragraph id="5C504155FFE4FFCC8CC2FF38FE5C00D4" blockId="17.[151,1437,150,285]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FF38FE3F00B0" bold="true" box="[199,438,150,176]" pageId="17" pageNumber="18">Palaeoenvironment.</emphasis>
Unknown; the collections from Lăpugiu de Sus indicate a mixing of coastal-inner neritic and middle neritic elements.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFE4FFCC8CC2FF70FE77011D" pageId="17" pageNumber="18" type="distribution">
<paragraph id="5C504155FFE4FFCC8CC2FF70FE77011D" blockId="17.[151,1437,150,285]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FF70FDE600F8" bold="true" box="[199,623,222,248]" pageId="17" pageNumber="18">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFE4FFCC8FC3FF71FBD900F8" box="[966,1104,223,249]" italics="true" pageId="17" pageNumber="18">Făget Basin</emphasis>
: Lăpugiu de Sus (own data), Coşteiu de Sus (
<bibRefCitation id="387E3CA4FFE4FFCC8D49FEACFE78011D" author="Boettger, O." box="[332,497,258,285]" pageId="17" pageNumber="18" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger 1906</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>