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<document id="F59ECD1E5DC3F411CC6496A77A05C69C" ID-CLB-Dataset="5925" ID-DOI="10.11646/zootaxa.4982.1.1" ID-GBIF-Dataset="f68b381d-d0dd-44e2-b9fe-6cf3c1aab1fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921842" ID-ZooBank="58388FB8-128A-4381-83D1-3C508D0D3873" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623285106829" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2021" docId="D446F043FFF2FFD58C92F8FDFB7704C5" docLanguage="en" docName="zootaxa.4982.1.1.pdf" docOrigin="Zootaxa 4982 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Antithala claviformis Harzhauser &amp; Landau 2021, nov. sp." docType="treatment" docVersion="10" lastPageNumber="9" masterDocId="287F883BFFF5FFDD8C05FFAEFF890000" masterDocTitle="An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea" masterLastPageNumber="70" masterPageNumber="1" pageNumber="8" updateTime="1699009793899" updateUser="ExternalLinkService">
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<mods:title id="3F809BDE2917728F14E9E56250984714">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="FE6D44E3AD1A0F746D43D0A4582A411B">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="1838DC5124A5777634B025C0AD217D1E">Landau, Bernard</mods:namePart>
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<mods:title id="8BC85FB870C1797131A0077576A5C389">Zootaxa</mods:title>
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<mods:date id="5640554259711350C0CC750498BEA858">2021</mods:date>
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<mods:number id="A0C08750116C5DB3B0DAB49263DFB1F3">4982</mods:number>
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<treatment id="D446F043FFF2FFD58C92F8FDFB7704C5" ID-DOI="http://doi.org/10.5281/zenodo.5044004" ID-GBIF-Taxon="183225124" ID-Zenodo-Dep="5044004" LSID="urn:lsid:plazi:treatment:D446F043FFF2FFD58C92F8FDFB7704C5" httpUri="http://treatment.plazi.org/id/D446F043FFF2FFD58C92F8FDFB7704C5" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="14F512DEFFF2FFDA8C92F8FDFE74076D" box="[151,509,1875,1902]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="5C504155FFF2FFDA8C92F8FDFE74076D" blockId="7.[151,509,1875,1944]" box="[151,509,1875,1902]" pageId="7" pageNumber="8">
<heading id="0718F639FFF2FFDA8C92F8FDFE74076D" bold="true" box="[151,509,1875,1902]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="4">
<emphasis id="6E9B9D47FFF2FFDA8C92F8FDFE74076D" bold="true" box="[151,509,1875,1902]" pageId="7" pageNumber="8">
<taxonomicName id="9BEF3AD6FFF2FFDA8C92F8FDFE11076E" ID-CoL="8KSSP" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[151,408,1875,1902]" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="claviformis" status="sp. nov.">
<emphasis id="6E9B9D47FFF2FFDA8C92F8FDFE11076E" bold="true" box="[151,408,1875,1902]" italics="true" pageId="7" pageNumber="8">Antithala claviformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFF2FFDA8D9AF8FDFE74076D" box="[415,509,1875,1901]" pageId="7" pageNumber="8" rank="species">nov. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="14F512DEFFF2FFDA8C92F8D9FE6B0798" box="[151,482,1911,1944]" pageId="7" pageNumber="8" type="description">
<paragraph id="5C504155FFF2FFDA8C92F8D9FE6B0798" blockId="7.[151,509,1875,1944]" box="[151,482,1911,1944]" pageId="7" pageNumber="8">
<figureCitation id="C4D45DD0FFF2FFDA8C92F8D9FF780791" box="[151,241,1911,1937]" captionStart="FIGURE 3" captionStartId="9.[151,250,1824,1848]" captionTargetBox="[161,1426,848,1800]" captionTargetId="figure-291@9.[161,1426,848,1800]" captionTargetPageId="9" captionText="FIGURE 3. A1–A2. Antithala claviformis nov. sp., holotype, Coşteiu de Sus (Romania), NHMW 1869/0001/0670. B1–B2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0001. C1–C2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0002. D1–D2. Antithala filipescui nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0002. E1–E2. Antithala filipescui nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1876/0011/0012. F1–F2. Antithala filipescui nov. sp. paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0001." figureDoi="http://doi.org/10.5281/zenodo.4921848" httpUri="https://zenodo.org/record/4921848/files/figure.png" pageId="7" pageNumber="8">Figs 3A</figureCitation>
<figureCitation id="C4D45DD0FFF2FFDA8CF4F824FE920791" box="[241,283,1911,1944]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="7" pageNumber="8">
<subScript id="C06B4310FFF2FFDA8CF4F824FF700798" attach="left" box="[241,249,1930,1944]" fontSize="6" pageId="7" pageNumber="8">1</subScript>
–A
</figureCitation>
<figureCitation id="C4D45DD0FFF2FFDA8D1EF824FEAA0798" box="[283,291,1930,1944]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="7" pageNumber="8">
<subScript id="C06B4310FFF2FFDA8D1EF824FEAA0798" attach="left" box="[283,291,1930,1944]" fontSize="6" pageId="7" pageNumber="8">2</subScript>
</figureCitation>
,
<figureCitation id="C4D45DD0FFF2FFDA8D34F8D9FED80791" box="[305,337,1911,1937]" captionStart="FIGURE 3" captionStartId="9.[151,250,1824,1848]" captionTargetBox="[161,1426,848,1800]" captionTargetId="figure-291@9.[161,1426,848,1800]" captionTargetPageId="9" captionText="FIGURE 3. A1–A2. Antithala claviformis nov. sp., holotype, Coşteiu de Sus (Romania), NHMW 1869/0001/0670. B1–B2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0001. C1–C2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0002. D1–D2. Antithala filipescui nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0002. E1–E2. Antithala filipescui nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1876/0011/0012. F1–F2. Antithala filipescui nov. sp. paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0001." figureDoi="http://doi.org/10.5281/zenodo.4921848" httpUri="https://zenodo.org/record/4921848/files/figure.png" pageId="7" pageNumber="8">3B</figureCitation>
<figureCitation id="C4D45DD0FFF2FFDA8D57F824FEF30792" box="[338,378,1912,1944]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="7" pageNumber="8">
<subScript id="C06B4310FFF2FFDA8D57F824FED30798" attach="right" box="[338,346,1930,1944]" fontSize="6" pageId="7" pageNumber="8">1</subScript>
–B
</figureCitation>
<figureCitation id="C4D45DD0FFF2FFDA8D7EF824FE0A0798" box="[379,387,1930,1944]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="7" pageNumber="8">
<subScript id="C06B4310FFF2FFDA8D7EF824FE0A0798" attach="right" box="[379,387,1930,1944]" fontSize="6" pageId="7" pageNumber="8">2</subScript>
</figureCitation>
,
<figureCitation id="C4D45DD0FFF2FFDA8D94F8D9FE3B0791" box="[401,434,1911,1937]" captionStart="FIGURE 3" captionStartId="9.[151,250,1824,1848]" captionTargetBox="[161,1426,848,1800]" captionTargetId="figure-291@9.[161,1426,848,1800]" captionTargetPageId="9" captionText="FIGURE 3. A1–A2. Antithala claviformis nov. sp., holotype, Coşteiu de Sus (Romania), NHMW 1869/0001/0670. B1–B2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0001. C1–C2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0002. D1–D2. Antithala filipescui nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0002. E1–E2. Antithala filipescui nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1876/0011/0012. F1–F2. Antithala filipescui nov. sp. paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0001." figureDoi="http://doi.org/10.5281/zenodo.4921848" httpUri="https://zenodo.org/record/4921848/files/figure.png" pageId="7" pageNumber="8">3C</figureCitation>
<figureCitation id="C4D45DD0FFF2FFDA8DB7F824FE530791" box="[434,474,1911,1944]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="7" pageNumber="8">
<subScript id="C06B4310FFF2FFDA8DB7F824FE330798" attach="left" box="[434,442,1930,1944]" fontSize="6" pageId="7" pageNumber="8">1</subScript>
–C
</figureCitation>
<figureCitation id="C4D45DD0FFF2FFDA8DDFF824FE6B0798" box="[474,482,1930,1944]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="7" pageNumber="8">
<subScript id="C06B4310FFF2FFDA8DDFF824FE6B0798" attach="left" box="[474,482,1930,1944]" fontSize="6" pageId="7" pageNumber="8">2</subScript>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFF2FFDA8C92F812FE9B07F2" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="5C504155FFF2FFDA8C92F812FE9B07F2" blockId="7.[151,1437,1980,2034]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="7CFF667BFFF2FFDA8C92F812FE9B07F2" pageId="7" pageNumber="8">
<taxonomicName id="9BEF3AD6FFF2FFDA8C92F812FE8107D3" ID-CoL="7P94F" authority="Sturi" authorityName="Sturi" box="[151,264,1980,2003]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFF2FFDA8C92F812FE8107D3" box="[151,264,1980,2003]" italics="true" pageId="7" pageNumber="8">Mitra Sturi</emphasis>
</taxonomicName>
nov. form.—
<treatmentCitation id="DD4E6744FFF2FFDA8D95F812FD3E07D4" author="Hoernes, R. &amp; Auinger, M." box="[400,695,1980,2004]" page="90" pageId="7" pageNumber="8" year="1880">
<bibRefCitation id="387E3CA4FFF2FFDA8D95F812FD3E07D4" author="Hoernes, R. &amp; Auinger, M." box="[400,695,1980,2004]" pageId="7" pageNumber="8" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger 1880: 90</bibRefCitation>
</treatmentCitation>
(
<emphasis id="6E9B9D47FFF2FFDA8EC3F812FD7A07D3" box="[710,755,1980,2003]" italics="true" pageId="7" pageNumber="8">pars</emphasis>
;
<typeStatus id="8354FFF7FFF2FFDA8F05F812FCD007D4" box="[768,857,1980,2004]" pageId="7" pageNumber="8" type="syntype">syntypes</typeStatus>
not illustrated) [
<emphasis id="6E9B9D47FFF2FFDA8807F812FBA107D3" box="[1026,1064,1980,2003]" italics="true" pageId="7" pageNumber="8">non</emphasis>
<taxonomicName id="9BEF3AD6FFF2FFDA882AF812FE8107F2" ID-CoL="8KRWF" authority="(Hoernes &amp; Auinger, 1880)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="sturi">
<emphasis id="6E9B9D47FFF2FFDA882AF812FB4807D3" box="[1071,1217,1980,2004]" italics="true" pageId="7" pageNumber="8">Antithala sturi</emphasis>
(
<bibRefCitation id="387E3CA4FFF2FFDA88CAF812FF7607F2" author="Hoernes, R. &amp; Auinger, M." pageId="7" pageNumber="8" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFDFFD58C92FF39FDB40189" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="5C504155FFFDFFD58C92FF39FD03011E" blockId="8.[151,1437,150,1222]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58C92FF39FECE00B0" bold="true" box="[151,327,150,177]" pageId="8" pageNumber="9">Type material.</emphasis>
<materialsCitation id="EC874B08FFFDFFD58D55FF39FF6500DB" ID-GBIF-Occurrence="3311814428" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="8" pageNumber="9" specimenCode="NHMW 1869" specimenCount="1" typeStatus="holotype">
<typeStatus id="8354FFF7FFFDFFD58D55FF39FE3400B1" box="[336,445,151,177]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
:
<specimenCode id="0C49E92EFFFDFFD58DCFFF39FDEC00B1" box="[458,613,151,177]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW 1869</specimenCode>
/0001/0670, SL:
<quantity id="9B17ECB0FFFDFFD58F2DFF39FC0B00B1" box="[808,898,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="9" unit="mm" value="4.4">4.4 mm</quantity>
,
<location id="5930178EFFFDFFD58F95FF39FBB300B1" LSID="urn:lsid:plazi:treatment:D446F043FFF2FFD58C92F8FDFB7704C5:5930178EFFFDFFD58F95FF39FBB300B1" box="[912,1082,151,177]" country="Romania" name="Costeiu de Sus" pageId="8" pageNumber="9">
Coşteiu
<collectorName id="F11A2483FFFDFFD58FEAFF39FBB300B1" box="[1007,1082,151,177]" pageId="8" pageNumber="9">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFDFFD58849FF39FB3E00B2" box="[1100,1207,151,178]" name="Romania" pageId="8" pageNumber="9">Romania</collectingCountry>
), MD:
<quantity id="9B17ECB0FFFDFFD5890FFF39FAED00B1" box="[1290,1380,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="9" unit="mm" value="1.8">1.8 mm</quantity>
, figs 3A
<subScript id="C06B4310FFFDFFD58CBCFF63FF4800DB" attach="left" box="[185,193,205,219]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
–A
<subScript id="C06B4310FFFDFFD58CE1FF63FF6500DB" attach="left" box="[228,236,205,219]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
</materialsCitation>
.
<materialsCitation id="EC874B08FFFDFFD58CFEFF15FA1D00DB" ID-GBIF-Occurrence="3311814394" box="[251,1428,186,219]" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="8" pageNumber="9" specimenCode="NHMW 2020" specimenCount="1" typeStatus="paratype">
<typeStatus id="8354FFF7FFFDFFD58CFEFF15FEE500D5" box="[251,364,187,213]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="0C49E92EFFFDFFD58D7DFF15FD9D00D5" box="[376,532,186,213]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW 2020</specimenCode>
/0056/0001, SL:
<quantity id="9B17ECB0FFFDFFD58ED7FF14FCA200D5" box="[722,811,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="9" unit="mm" value="4.5">4.5 mm</quantity>
, MD:
<quantity id="9B17ECB0FFFDFFD58F71FF15FC4400D5" box="[884,973,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="9" unit="mm" value="1.9">1.9 mm</quantity>
,
<location id="5930178EFFFDFFD58FDFFF15FB0B00D5" LSID="urn:lsid:plazi:treatment:D446F043FFF2FFD58C92F8FDFB7704C5:5930178EFFFDFFD58FDFFF15FB0B00D5" box="[986,1154,187,213]" country="Romania" name="Costeiu de Sus" pageId="8" pageNumber="9">
Coşteiu
<collectorName id="F11A2483FFFDFFD58832FF15FB0B00D5" box="[1079,1154,187,213]" pageId="8" pageNumber="9">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFDFFD58897FF15FB7400D6" box="[1170,1277,187,214]" name="Romania" pageId="8" pageNumber="9">Romania</collectingCountry>
), figs 3B
<subScript id="C06B4310FFFDFFD58966FF63FAE200DB" attach="left" box="[1379,1387,205,219]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
–B
<subScript id="C06B4310FFFDFFD58989FF63FA1D00DB" attach="left" box="[1420,1428,205,219]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
</materialsCitation>
;
<materialsCitation id="EC874B08FFFDFFD58C92FF71FAB100F9" ID-GBIF-Occurrence="3311814448" box="[151,1336,222,255]" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="8" pageNumber="9" specimenCode="NHMW 2020" specimenCount="11" typeStatus="paratype">
<specimenCode id="0C49E92EFFFDFFD58C92FF71FEB900F9" box="[151,304,222,249]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW 2020</specimenCode>
/0056/0002, SL:
<quantity id="9B17ECB0FFFDFFD58DEFFF71FDC900F9" box="[490,576,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="8" pageNumber="9" unit="mm" value="4.1">4.1 mm</quantity>
, MD:
<quantity id="9B17ECB0FFFDFFD58E81FF71FD5500F9" box="[644,732,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="9" unit="mm" value="1.9">1.9 mm</quantity>
.
<location id="5930178EFFFDFFD58EE0FF71FC0E00F9" LSID="urn:lsid:plazi:treatment:D446F043FFF2FFD58C92F8FDFB7704C5:5930178EFFFDFFD58EE0FF71FC0E00F9" box="[741,903,223,249]" country="Romania" name="Costeiu de Sus" pageId="8" pageNumber="9">
Coşteiu
<collectorName id="F11A2483FFFDFFD58F3AFF71FC0E00F9" box="[831,903,223,249]" pageId="8" pageNumber="9">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFDFFD58F90FF71FB8900FA" box="[917,1024,223,250]" name="Romania" pageId="8" pageNumber="9">Romania</collectingCountry>
), figs 3C
<subScript id="C06B4310FFFDFFD58864FF5FFBE000FF" attach="left" box="[1121,1129,241,255]" fontSize="6" pageId="8" pageNumber="9">1</subScript>
–C
<subScript id="C06B4310FFFDFFD5888FFF5FFB1B00FF" attach="left" box="[1162,1170,241,255]" fontSize="6" pageId="8" pageNumber="9">2</subScript>
;
<specimenCount id="4AE98ADCFFFDFFD588A5FF71FAB100F9" box="[1184,1336,223,249]" pageId="8" pageNumber="9" type="generic">11 specimens</specimenCount>
</materialsCitation>
,
<materialsCitation id="EC874B08FFFDFFD58944FF71FD0F011E" ID-GBIF-Occurrence="3311814315" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="8" pageNumber="9" specimenCode="NHMW 2020" specimenCount="1" typeStatus="paratype">
<specimenCode id="0C49E92EFFFDFFD58944FF71FF46011D" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="9" type="Museum">NHMW 2020</specimenCode>
/0056/0004,
<location id="5930178EFFFDFFD58D58FEADFD8A011D" LSID="urn:lsid:plazi:treatment:D446F043FFF2FFD58C92F8FDFB7704C5:5930178EFFFDFFD58D58FEADFD8A011D" box="[349,515,259,285]" country="Romania" name="Costeiu de Sus" pageId="8" pageNumber="9">
Coşteiu
<collectorName id="F11A2483FFFDFFD58DBCFEADFD8A011D" box="[441,515,259,285]" pageId="8" pageNumber="9">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFDFFD58E17FEADFDF7011E" box="[530,638,259,286]" name="Romania" pageId="8" pageNumber="9">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFFDFFD58CC2FE89FCA50141" blockId="8.[151,1437,150,1222]" box="[199,812,294,322]" pageId="8" pageNumber="9">
<materialsCitation id="EC874B08FFFDFFD58CC2FE89FCA10141" ID-GBIF-Occurrence="3311814483" box="[199,808,294,322]" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="8" pageNumber="9" specimenCount="1" typeStatus="holotype">
<emphasis id="6E9B9D47FFFDFFD58CC2FE89FEEF0140" bold="true" box="[199,358,294,321]" pageId="8" pageNumber="9">
<typeStatus id="8354FFF7FFFDFFD58CC2FE89FE8B0141" box="[199,258,295,321]" pageId="8" pageNumber="9">Type</typeStatus>
locality.
</emphasis>
<location id="5930178EFFFDFFD58D69FE89FD9B0141" LSID="urn:lsid:plazi:treatment:D446F043FFF2FFD58C92F8FDFB7704C5:5930178EFFFDFFD58D69FE89FD9B0141" box="[364,530,295,321]" country="Romania" name="Costeiu de Sus" pageId="8" pageNumber="9">
Coşteiu
<collectorName id="F11A2483FFFDFFD58DCCFE89FD9B0141" box="[457,530,295,321]" pageId="8" pageNumber="9">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFFDFFD58E24FE89FD050142" box="[545,652,295,322]" name="Romania" pageId="8" pageNumber="9">Romania</collectingCountry>
), Făget Basin
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFFDFFD58CC2FEE5FD750165" blockId="8.[151,1437,150,1222]" box="[199,764,331,357]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FEE5FEE60165" bold="true" box="[199,367,331,357]" pageId="8" pageNumber="9">
<typeStatus id="8354FFF7FFFDFFD58CC2FEE5FE8B0165" box="[199,258,331,357]" pageId="8" pageNumber="9">Type</typeStatus>
stratum.
</emphasis>
Silt and clay of the Dej Formation.
</paragraph>
<paragraph id="5C504155FFFDFFD58CC2FEC0FDB40189" blockId="8.[151,1437,150,1222]" box="[199,573,366,393]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FEC0FF740188" bold="true" box="[199,253,366,392]" pageId="8" pageNumber="9">Age.</emphasis>
Middle Miocene, Badenian.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFDFFD58CC2FE3DFC3A01AD" box="[199,947,403,430]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="5C504155FFFDFFD58CC2FE3DFC3A01AD" blockId="8.[151,1437,150,1222]" box="[199,947,403,430]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FE3DFEC701AD" bold="true" box="[199,334,403,429]" pageId="8" pageNumber="9">Etymology.</emphasis>
“
<emphasis id="6E9B9D47FFFDFFD58D64FE3DFE2D01AD" box="[353,420,403,429]" italics="true" pageId="8" pageNumber="9">Clava</emphasis>
” (Latin: club), referring to the club-like shape.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFDFFD58CC2FE18FE6C01F5" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="5C504155FFFDFFD58CC2FE18FE6C01F5" blockId="8.[151,1437,150,1222]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FE18FECB01D0" bold="true" box="[199,322,438,464]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
<taxonomicName id="9BEF3AD6FFFDFFD58D4EFE19FD8601D1" box="[331,527,439,465]" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="6E9B9D47FFFDFFD58D4EFE19FE3A01D1" box="[331,435,439,465]" italics="true" pageId="8" pageNumber="9">Antithala</emphasis>
species
</taxonomicName>
of small size, with prominently beaded sculpture, three beaded cords on spire whorls, four columellar folds.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFDFFD58CC2FE50FE2B03A5" pageId="8" pageNumber="9" type="description">
<paragraph id="5C504155FFFDFFD58CC2FE50FB17035D" blockId="8.[151,1437,150,1222]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FE50FED00218" bold="true" box="[199,345,510,536]" pageId="8" pageNumber="9">Description.</emphasis>
Shell small, stout fusiform. Protoconch unknown. Teleoconch of 4.5 straight-sided whorls with periphery at abapical suture; suture distinctly incised. Spire weakly gradate, conical to slightly cyrtoconoid. First teleoconch whorl with prominent axial ribs intersected by two weaker spiral grooves forming three spiral rows of nodes; spiral rows strengthen abapically, forming prominent subquadrate nodes on later whorls. Fourth spiral cord at abapical suture largely covered by succeeding whorl. Last whorl regularly convex at periphery, passing via distinct concavity into strongly constricted base. Seven to eight spiral rows of nodes on last whorl forming orthocline to weakly sigmoidal axial ribs. Fasciole weak, bearing about five spiral cords with reduced nodes. Aperture elongate, narrow with distinct anal sinus narrowed by broad, low parietal denticle. Columella narrow, sharply delimited, bearing four folds, abapically rapidly decreasing in strength. Outer lip thickened, with six small denticles within. Siphonal canal moderately long, recurved, bent slightly to left, with shallow siphonal notch.
</paragraph>
<paragraph id="5C504155FFFDFFD58CC2FCC8FE2B03A5" blockId="8.[151,1437,150,1222]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FCC8FDB70380" bold="true" box="[199,574,870,897]" pageId="8" pageNumber="9">Shell measurements and ratios.</emphasis>
SL:
<quantity id="9B17ECB0FFFDFFD58E73FCC8FC890381" box="[630,768,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="8" pageNumber="9" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
, MD:
<quantity id="9B17ECB0FFFDFFD58F4CFCC9FC5A0381" box="[841,979,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="8" pageNumber="9" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
; AA: 45–51°, SL/MD: 2.2–2.6, AL/AW: 4.8–5.6, AH/S: 2.2–2.5.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFDFFD58CC2FC00FE1604A2" pageId="8" pageNumber="9" type="discussion">
<paragraph id="5C504155FFFDFFD58CC2FC00FAE60459" blockId="8.[151,1437,150,1222]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FC00FEC503C8" bold="true" box="[199,332,942,968]" pageId="8" pageNumber="9">Discussion.</emphasis>
The specimens here described as
<taxonomicName id="9BEF3AD6FFFDFFD58ED1FC01FC4A03C9" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[724,963,943,969]" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="claviformis" status="sp. nov.">
<emphasis id="6E9B9D47FFFDFFD58ED1FC01FC4A03C9" box="[724,963,943,969]" italics="true" pageId="8" pageNumber="9">Antithala claviformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFFDFFD58FCEFC01FBAA03C9" box="[971,1059,943,969]" pageId="8" pageNumber="9" rank="species">nov. sp.</taxonomicNameLabel>
were probably identified by
<bibRefCitation id="387E3CA4FFFDFFD58974FC01FE2F03ED" author="Hoernes, R. &amp; Auinger, M." pageId="8" pageNumber="9" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger (1880)</bibRefCitation>
as
<taxonomicName id="9BEF3AD6FFFDFFD58DC8FC7DFC0103ED" authority="(Hoernes &amp; Auinger, 1880)" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[461,904,979,1005]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="sturi">
<emphasis id="6E9B9D47FFFDFFD58DC8FC7DFDC103EC" box="[461,584,979,1004]" italics="true" pageId="8" pageNumber="9">Mitra sturi</emphasis>
(
<bibRefCitation id="387E3CA4FFFDFFD58E5DFC7DFCF603ED" author="Hoernes, R. &amp; Auinger, M." box="[600,895,979,1005]" pageId="8" pageNumber="9" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
due to them having almost identical sculpture.
<taxonomicName id="9BEF3AD6FFFDFFD58C92FC59FE0E0411" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[151,391,1015,1041]" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="claviformis">
<emphasis id="6E9B9D47FFFDFFD58C92FC59FE0E0411" box="[151,391,1015,1041]" italics="true" pageId="8" pageNumber="9">Antithala claviformis</emphasis>
</taxonomicName>
, however, is constantly smaller, broader and has a strongly constricted base, resulting in a club-like last whorl. Most importantly,
<taxonomicName id="9BEF3AD6FFFDFFD58E5EFBB5FD720435" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[603,763,1051,1077]" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="claviformis">
<emphasis id="6E9B9D47FFFDFFD58E5EFBB5FD720435" box="[603,763,1051,1077]" italics="true" pageId="8" pageNumber="9">A. claviformis</emphasis>
</taxonomicName>
has four columellar folds whereas
<taxonomicName id="9BEF3AD6FFFDFFD58891FBB5FB600434" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[1172,1257,1051,1077]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="sturi">
<emphasis id="6E9B9D47FFFDFFD58891FBB5FB600434" box="[1172,1257,1051,1077]" italics="true" pageId="8" pageNumber="9">A. sturi</emphasis>
</taxonomicName>
has only three.
<taxonomicName id="9BEF3AD6FFFDFFD58C92FB91FEE40459" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[151,365,1087,1113]" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="filipescui" status="sp. nov.">
<emphasis id="6E9B9D47FFFDFFD58C92FB91FEE40459" box="[151,365,1087,1113]" italics="true" pageId="8" pageNumber="9">Antithala filipescui</emphasis>
</taxonomicName>
<emphasis id="6E9B9D47FFFDFFD58D71FB91FE470459" bold="true" box="[372,462,1087,1113]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="75A8203CFFFDFFD58D71FB91FE470459" box="[372,462,1087,1113]" pageId="8" pageNumber="9" rank="species">nov. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="9BEF3AD6FFFDFFD58E01FB91FD270459" authorityName="Hoernes &amp; Auinger" authorityYear="1880" box="[516,686,1087,1113]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="neugeboreni">
<emphasis id="6E9B9D47FFFDFFD58E01FB91FD270459" box="[516,686,1087,1113]" italics="true" pageId="8" pageNumber="9">A. neugeboreni</emphasis>
</taxonomicName>
both differ in having four rows of nodules on the spire whorls.
</paragraph>
<paragraph id="5C504155FFFDFFD58CC2FBCCFE1604A2" blockId="8.[151,1437,150,1222]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FBCCFE3F047C" bold="true" box="[199,438,1122,1148]" pageId="8" pageNumber="9">Palaeoenvironment.</emphasis>
The assemblages from Coşteiu de Sus suggests middle neritic palaeoenvironments with soft bottom (own data).
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFFDFFD58CC2FB04FB7704C5" box="[199,1278,1194,1222]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="5C504155FFFDFFD58CC2FB04FB7704C5" blockId="8.[151,1437,150,1222]" box="[199,1278,1194,1222]" pageId="8" pageNumber="9">
<emphasis id="6E9B9D47FFFDFFD58CC2FB04FDE304C5" bold="true" box="[199,618,1194,1221]" pageId="8" pageNumber="9">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFFDFFD58FBCFB05FBCB04C4" box="[953,1090,1195,1221]" italics="true" pageId="8" pageNumber="9">Făget Basin</emphasis>
: Coşteiu de Sus.
</paragraph>
</subSubSection>
</treatment>
</document>
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