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<mods:title id="64886AE0786DF482284E63BD8F25006B">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="AE5D0FD30DA4EC9EF6D023CA0D86A875">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="A767CB33F3748F4F16A9B0B1CEEA5DFC">Landau, Bernard</mods:namePart>
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<treatment id="D446F043FFEDFFC48C92FCECFDF301D1" ID-DOI="http://doi.org/10.5281/zenodo.5044018" ID-GBIF-Taxon="183225128" ID-Zenodo-Dep="5044018" LSID="urn:lsid:plazi:treatment:D446F043FFEDFFC48C92FCECFDF301D1" httpUri="http://treatment.plazi.org/id/D446F043FFEDFFC48C92FCECFDF301D1" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<paragraph id="5C504155FFEDFFC58C92FCECFC26035C" blockId="24.[151,943,834,903]" box="[151,943,834,861]" pageId="24" pageNumber="25">
<heading id="0718F639FFEDFFC58C92FCECFC26035C" bold="true" box="[151,943,834,861]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="4">
<emphasis id="6E9B9D47FFEDFFC58C92FCECFC26035C" bold="true" box="[151,943,834,861]" pageId="24" pageNumber="25">
<taxonomicName id="9BEF3AD6FFEDFFC58C92FCECFCAB035C" ID-CoL="8KS63" authority="(Hoernes &amp; Auinger, 1880)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[151,802,834,861]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="lapugyensis" status="comb. nov.">
<emphasis id="6E9B9D47FFEDFFC58C92FCECFE45035D" bold="true" box="[151,460,834,861]" italics="true" pageId="24" pageNumber="25">Bellardithala lapugyensis</emphasis>
(
<bibRefCitation id="387E3CA4FFEDFFC58DDEFCECFC93035C" author="Hoernes, R. &amp; Auinger, M." box="[475,794,834,860]" pageId="24" pageNumber="25" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFEDFFC58F2CFCECFC26035C" box="[809,943,834,860]" pageId="24" pageNumber="25" rank="species">nov. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="14F512DEFFEDFFC58C92FCC8FE4403FF" pageId="24" pageNumber="25" type="description">
<paragraph id="5C504155FFEDFFC58C92FCC8FE370387" blockId="24.[151,943,834,903]" box="[151,446,870,903]" pageId="24" pageNumber="25">
<figureCitation id="C4D45DD0FFEDFFC58C92FCC8FF780380" box="[151,241,870,896]" captionStart="FIGURE 9" captionStartId="26.[151,250,1683,1707]" captionTargetBox="[151,1436,561,1660]" captionTargetId="figure-169@26.[151,1436,561,1660]" captionTargetPageId="26" captionText="FIGURE 9 A1–A2. Bellardithala lapugyensis (Hoernes &amp; Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0101. B1–B. Bellardithala lapugyensis (Hoernes &amp; Auinger, 1880), paralectotype, Lăpugiu de Sus (Romania), NHMW 2020/0053/0001. C1–C2. Bellardithala lapugyensis (Hoernes &amp; Auinger, 1880), paralectotype, Lăpugiu de Sus (Roma- nia), 2020/0053/0002. D1–D2. Bellardithala partschi (Hörnes, 1852), lectotype, Steinebrunn (Austria), NHMW 1846/0037/0113. E1–E2. Bellardithala partschi (Hörnes, 1852), paralectotype, Steinebrunn (Austria), NHMW 2020/0060/0004. F1–F2. Bellardithala partschi (Hörnes, 1852), paralectotype, Steinebrunn (Austria), NHMW 1860/0001/0110." figureDoi="http://doi.org/10.5281/zenodo.4921860" httpUri="https://zenodo.org/record/4921860/files/figure.png" pageId="24" pageNumber="25">Figs 9A</figureCitation>
<figureCitation id="C4D45DD0FFEDFFC58CF4FCD7FE920380" box="[241,283,870,903]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="24" pageNumber="25">
<subScript id="C06B4310FFEDFFC58CF4FCD7FF700387" attach="left" box="[241,249,889,903]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
–A
</figureCitation>
<figureCitation id="C4D45DD0FFEDFFC58D1EFCD7FEAA0387" box="[283,291,889,903]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="24" pageNumber="25">
<subScript id="C06B4310FFEDFFC58D1EFCD7FEAA0387" attach="left" box="[283,291,889,903]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
</figureCitation>
, B
<figureCitation id="C4D45DD0FFEDFFC58D41FCD7FE3F0380" box="[324,438,870,903]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="24" pageNumber="25">
<subScript id="C06B4310FFEDFFC58D41FCD7FEC50387" attach="left" box="[324,332,889,903]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
–B, C
<subScript id="C06B4310FFEDFFC58D88FCD7FE1C0387" attach="left" box="[397,405,889,903]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
–C
</figureCitation>
<figureCitation id="C4D45DD0FFEDFFC58DB3FCD7FE370387" box="[438,446,889,903]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="24" pageNumber="25">
<subScript id="C06B4310FFEDFFC58DB3FCD7FE370387" attach="left" box="[438,446,889,903]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
</figureCitation>
</paragraph>
<paragraph id="5C504155FFEDFFC58C92FC05FC0803C2" blockId="24.[151,1436,938,1116]" box="[151,897,938,963]" pageId="24" pageNumber="25">
[
<taxonomicName id="9BEF3AD6FFEDFFC58CA5FC05FF7703C3" authorityName="Swainson" authorityYear="1840" box="[160,254,939,963]" class="Insecta" family="Nymphalidae" genus="Callithea" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="6E9B9D47FFEDFFC58CA5FC05FF7703C3" box="[160,254,939,963]" italics="true" pageId="24" pageNumber="25">Callithea</emphasis>
</taxonomicName>
]
<emphasis id="6E9B9D47FFEDFFC58D08FC05FE0503C2" box="[269,396,939,962]" italics="true" pageId="24" pageNumber="25">Lapugyensis</emphasis>
nov. form.—Hoernes 1880: 125 [
<emphasis id="6E9B9D47FFEDFFC58EE6FC05FCFB03C3" box="[739,882,939,963]" italics="true" pageId="24" pageNumber="25">nomen nudum</emphasis>
].
</paragraph>
<paragraph id="5C504155FFEDFFC58C92FC64FE4403FF" blockId="24.[151,1436,938,1116]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58C92FC64FEE503E1" box="[151,364,970,993]" italics="true" pageId="24" pageNumber="25">
*
<taxonomicName id="9BEF3AD6FFEDFFC58CA9FC64FEE503E1" authority="Lapugyensis" authorityName="Lapugyensis" box="[172,364,970,993]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">Mitra Lapugyensis</taxonomicName>
</emphasis>
nov. form.—
<bibRefCitation id="387E3CA4FFEDFFC58DF3FC67FD7303E1" author="Hoernes, R. &amp; Auinger, M." box="[502,762,969,993]" pageId="24" pageNumber="25" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger 1880</bibRefCitation>
(pars): 89, pl. 10, figs 21a–c [
<emphasis id="6E9B9D47FFEDFFC58832FC64FBD403E1" box="[1079,1117,970,993]" italics="true" pageId="24" pageNumber="25">non</emphasis>
pl. 10., figs 22a–22c =
<taxonomicName id="9BEF3AD6FFEDFFC58950FC67FEE00400" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="fedosovi" status="sp. nov.">
<emphasis id="6E9B9D47FFEDFFC58950FC67FEE00400" italics="true" pageId="24" pageNumber="25">Bellardithala fedosovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFEDFFC58D75FC46FE4803FF" box="[368,449,1000,1023]" pageId="24" pageNumber="25" rank="species">nov. sp.</taxonomicNameLabel>
].
</paragraph>
</subSubSection>
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<paragraph id="5C504155FFEDFFC58C92FBA9FD44041F" blockId="24.[151,1436,938,1116]" box="[151,717,1031,1055]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="7CFF667BFFEDFFC58C92FBA9FD44041F" box="[151,717,1031,1055]" pageId="24" pageNumber="25">
<taxonomicName id="9BEF3AD6FFEDFFC58C92FBA9FE7C041F" authority="Ho. Au." authorityName="Ho. Au." box="[151,501,1031,1055]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="lapugyensis" subGenus="Thala">
<emphasis id="6E9B9D47FFEDFFC58C92FBA9FF58041E" box="[151,209,1031,1054]" italics="true" pageId="24" pageNumber="25">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFEDFFC58CE5FBA9FE92041F" box="[224,283,1031,1055]" italics="true" pageId="24" pageNumber="25">Thala</emphasis>
)
<emphasis id="6E9B9D47FFEDFFC58D2FFBA9FE2B041F" box="[298,418,1031,1055]" italics="true" pageId="24" pageNumber="25">lapugyensis</emphasis>
Hö. Au.
</taxonomicName>
—
<treatmentCitation id="DD4E6744FFEDFFC58E0BFBA9FD40041F" author="Boettger, O." box="[526,713,1031,1055]" page="11" pageId="24" pageNumber="25" year="1906">
<bibRefCitation id="387E3CA4FFEDFFC58E0BFBA9FD40041F" author="Boettger, O." box="[526,713,1031,1055]" pageId="24" pageNumber="25" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger 1906: 11</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="5C504155FFEDFFC58C92FB8BFC2F043D" blockId="24.[151,1436,938,1116]" box="[151,934,1061,1085]" pageId="24" pageNumber="25">
?
<taxonomicName id="9BEF3AD6FFEDFFC58CACFB8BFE42043D" ID-CoL="7PKC6" authority="Lapugyensis R. H." authorityName="Lapugyensis R. H." box="[169,459,1061,1085]" class="Gastropoda" family="Clavatulidae" genus="Turricula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFEDFFC58CACFB8BFE07043D" box="[169,398,1061,1085]" italics="true" pageId="24" pageNumber="25">Turricula Lapugyensis</emphasis>
R. H.
</taxonomicName>
i Auing.—Friedberg 1928: 579, pl. 37, fig. 28.
</paragraph>
<paragraph id="5C504155FFEDFFC58C92FBEAFC26045C" blockId="24.[151,1436,938,1116]" box="[151,943,1092,1116]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="7CFF667BFFEDFFC58C92FBEAFC26045C" box="[151,943,1092,1116]" pageId="24" pageNumber="25">
<taxonomicName id="9BEF3AD6FFEDFFC58C92FBEAFDE3045C" authority="(Hoernes &amp; Auinger, 1880)" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[151,618,1092,1116]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="lapugyensis">
<emphasis id="6E9B9D47FFEDFFC58C92FBEAFED8045C" box="[151,337,1092,1116]" italics="true" pageId="24" pageNumber="25">Thala lapugyensis</emphasis>
(
<bibRefCitation id="387E3CA4FFEDFFC58D58FBEAFDEA045C" author="Hoernes, R. &amp; Auinger, M." box="[349,611,1092,1116]" pageId="24" pageNumber="25" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
—
<treatmentCitation id="DD4E6744FFEDFFC58E81FBEAFCAA045C" author="Baluk, W." box="[644,803,1092,1116]" page="38" pageId="24" pageNumber="25" year="1997">
<bibRefCitation id="387E3CA4FFEDFFC58E81FBEAFCAA045C" author="Baluk, W." box="[644,803,1092,1116]" pageId="24" pageNumber="25" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997: 38</bibRefCitation>
</treatmentCitation>
, pl. 11, fig. 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFEDFFC58C92FB24FE2B0750" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="5C504155FFEDFFC58C92FB24FCE90535" blockId="24.[151,1437,1162,2017]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58C92FB24FECE04A4" bold="true" box="[151,327,1162,1188]" pageId="24" pageNumber="25">Type material.</emphasis>
<materialsCitation id="EC874B08FFEDFFC58D55FB25FBF604CF" ID-GBIF-Occurrence="3311814414" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="24" pageNumber="25" specimenCode="NHMW 1854" specimenCount="1" typeStatus="lectotype">
<typeStatus id="8354FFF7FFEDFFC58D55FB25FE4904A5" box="[336,448,1163,1189]" pageId="24" pageNumber="25" type="lectotype">Lectotype</typeStatus>
(designated herein):
<specimenCode id="0C49E92EFFEDFFC58EB7FB25FCC704A5" box="[690,846,1162,1189]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="25" type="Museum">NHMW 1854</specimenCode>
/0035/0101, SL:
<quantity id="9B17ECB0FFEDFFC5880AFB24FBE004A5" box="[1039,1129,1162,1189]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="24" pageNumber="25" unit="mm" value="5.4">5.4 mm</quantity>
, MD: 2.0 mm,
<location id="5930178EFFEDFFC58919FB24FF3604C8" LSID="urn:lsid:plazi:treatment:D446F043FFEDFFC48C92FCECFDF301D1:5930178EFFEDFFC58919FB24FF3604C8" country="Romania" name="Lapugiu de Sus" pageId="24" pageNumber="25">
Lăpugiu
<collectorName id="F11A2483FFEDFFC58987FB24FF3604C8" pageId="24" pageNumber="25">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFEDFFC58CCAFB00FEB304C9" box="[207,314,1198,1225]" name="Romania" pageId="24" pageNumber="25">Romania</collectingCountry>
), illustrated in
<bibRefCitation id="387E3CA4FFEDFFC58DE0FB01FC8704C8" author="Hoernes, R. &amp; Auinger, M." box="[485,782,1198,1225]" pageId="24" pageNumber="25" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger (1880</bibRefCitation>
, pl. 10, figs 21a–c), figs 9A
<subScript id="C06B4310FFEDFFC58849FB6FFBDD04CF" attach="left" box="[1100,1108,1217,1231]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
–A
<subScript id="C06B4310FFEDFFC58872FB6FFBF604CF" attach="left" box="[1143,1151,1217,1231]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
</materialsCitation>
.
<materialsCitation id="EC874B08FFEDFFC5888BFB00FBC904F3" ID-GBIF-Occurrence="3311814318" collectionCode="NHMW" country="Romania" location="Lapugiu de Sus" pageId="24" pageNumber="25" specimenCode="NHMW 2020" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="8354FFF7FFEDFFC5888BFB00FABF04C8" box="[1166,1334,1198,1224]" pageId="24" pageNumber="25" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCode id="0C49E92EFFEDFFC58944FB01FF5904ED" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="25" type="Museum">NHMW 2020</specimenCode>
/0053/0001, SL:
<quantity id="9B17ECB0FFEDFFC58D8FFB7CFE6804ED" box="[394,481,1234,1261]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="24" pageNumber="25" unit="mm" value="5.1">5.1 mm</quantity>
, MD: 2.0 mm,
<location id="5930178EFFEDFFC58E8DFB7CFCBD04EC" LSID="urn:lsid:plazi:treatment:D446F043FFEDFFC48C92FCECFDF301D1:5930178EFFEDFFC58E8DFB7CFCBD04EC" box="[648,820,1234,1260]" country="Romania" name="Lapugiu de Sus" pageId="24" pageNumber="25">Lăpugiu de Sus</location>
(
<collectingCountry id="24F801C5FFEDFFC58F47FB7CFC2404ED" box="[834,941,1234,1261]" name="Romania" pageId="24" pageNumber="25">Romania</collectingCountry>
), figs 9B
<subScript id="C06B4310FFEDFFC5880AFB4BFB9E04F3" attach="left" box="[1039,1047,1253,1267]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
–B
<subScript id="C06B4310FFEDFFC5883DFB4BFBC904F3" attach="left" box="[1080,1088,1253,1267]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
</materialsCitation>
;
<materialsCitation id="EC874B08FFEDFFC5884BFB7DFB830510" ID-GBIF-Occurrence="3311814389" collectionCode="NHMW" country="Romania" location="Lapugiu de Sus" pageId="24" pageNumber="25" specimenCode="NHMW 2020" specimenCount="2" typeStatus="paralectotype">
<specimenCode id="0C49E92EFFEDFFC5884BFB7DFB6104ED" box="[1102,1256,1234,1261]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="25" type="Museum">NHMW 2020</specimenCode>
/0053/0002, SL:
<quantity id="9B17ECB0FFEDFFC58C92FB58FF780511" box="[151,241,1270,1297]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="24" pageNumber="25" unit="mm" value="5.4">5.4 mm</quantity>
, MD:
<quantity id="9B17ECB0FFEDFFC58D39FB58FE1F0511" box="[316,406,1270,1297]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="24" pageNumber="25" unit="mm" value="2.1">2.1 mm</quantity>
,
<location id="5930178EFFEDFFC58DA1FB58FDDC0510" LSID="urn:lsid:plazi:treatment:D446F043FFEDFFC48C92FCECFDF301D1:5930178EFFEDFFC58DA1FB58FDDC0510" box="[420,597,1270,1296]" country="Romania" name="Lapugiu de Sus" pageId="24" pageNumber="25">Lăpugiu de Sus</location>
(
<collectingCountry id="24F801C5FFEDFFC58E63FB58FD580511" box="[614,721,1270,1297]" name="Romania" pageId="24" pageNumber="25">Romania</collectingCountry>
), figs 9C
<subScript id="C06B4310FFEDFFC58F3FFAA7FCCB0517" attach="left" box="[826,834,1289,1303]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
–C
<subScript id="C06B4310FFEDFFC58F66FAA7FCE20517" attach="left" box="[867,875,1289,1303]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
;
<specimenCount id="4AE98ADCFFEDFFC58F7EFB58FB830510" box="[891,1034,1270,1296]" pageId="24" pageNumber="25" type="generic">2 specimens</specimenCount>
</materialsCitation>
,
<materialsCitation id="EC874B08FFEDFFC58812FB59FF5C0535" ID-GBIF-Occurrence="3311814467" collectionCode="NHMW" country="Romania" pageId="24" pageNumber="25" specimenCode="NHMW 2020" specimenCount="2" typeStatus="paralectotype">
<specimenCode id="0C49E92EFFEDFFC58812FB59FB3D0511" box="[1047,1204,1270,1297]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="25" type="Museum">NHMW 2020</specimenCode>
/0053/0003,
<specimenCount id="4AE98ADCFFEDFFC58947FB58FF5C0535" pageId="24" pageNumber="25" type="generic">2 specimens</specimenCount>
</materialsCitation>
,
<materialsCitation id="EC874B08FFEDFFC58CE5FAB5FCD50535" ID-GBIF-Occurrence="3311814439" box="[224,860,1306,1333]" collectionCode="NHMW" country="Romania" location="Lapugiu de Sus" pageId="24" pageNumber="25" specimenCode="NHMW 2020" specimenCount="1" typeStatus="paralectotype">
<specimenCode id="0C49E92EFFEDFFC58CE5FAB5FEF50535" box="[224,380,1306,1333]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="24" pageNumber="25" type="Museum">NHMW 2020</specimenCode>
/0053/0002, all
<location id="5930178EFFEDFFC58E2EFAB4FD500534" LSID="urn:lsid:plazi:treatment:D446F043FFEDFFC48C92FCECFDF301D1:5930178EFFEDFFC58E2EFAB4FD500534" box="[555,729,1306,1332]" country="Romania" name="Lapugiu de Sus" pageId="24" pageNumber="25">Lăpugiu de Sus</location>
(
<collectingCountry id="24F801C5FFEDFFC58EEDFAB4FCDD0535" box="[744,852,1306,1333]" name="Romania" pageId="24" pageNumber="25">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFEDFFC58CC2FA90FD240708" blockId="24.[151,1437,1162,2017]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58CC2FA90FE300558" bold="true" box="[199,441,1342,1368]" pageId="24" pageNumber="25">Revised description.</emphasis>
Shell small, moderately broad fusiform with weakly cyrtoconoid spire. Protoconch high conical with 2.7 regularly convex whorls (height: 500 μm, diameter: 510 μm). Transition to teleoconch marked by onset of axial sculpture. Teleoconch of 4.5 weakly convex whorls, with periphery close above abapical suture, separated by deeply incised suture. Axial sculpture of about 15 prominent axial ribs on first teleoconch whorl, separated by slightly narrower interspaces. Spiral sculpture consisting of indistinct spiral grooves separating four spiral rows of blurred nodules. Cancellate sculpture on later whorls, with axial ribs and interspaces of similar width, crossed by four broad, regularly spaced spiral cords on second teleoconch whorl and five on subsequent whorl. Spiral cords form low subquadrate nodules at intersection with axial ribs, and well defined grooves in interspaces. Abapically, spiral interspaces reduced to rectangular pits. Last whorl weakly convex, with weakly constricted base, bearing about 13 spiral cords; 7 spirals on weak fasciole. Aperture narrow, elongate, with indistinct anal sinus. Columellar callus very narrow, indistinct. Columella with four columellar folds, weakening abapically. Outer lip only slightly thickened, bearing eight small denticles within. Siphonal canal moderately long, moderately wide, straight or bent slightly left, with wide, shallow siphonal notch.
</paragraph>
<paragraph id="5C504155FFEDFFC58CC2F8BCFE2B0750" blockId="24.[151,1437,1162,2017]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58CC2F8BCFDBE072C" bold="true" box="[199,567,1810,1836]" pageId="24" pageNumber="25">Shell measurements and ratios</emphasis>
. SL:
<quantity id="9B17ECB0FFEDFFC58E73F8BCFC89072D" box="[630,768,1810,1837]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.4" metricValueMin="5.1" pageId="24" pageNumber="25" unit="mm" value="5.25" valueMax="5.4" valueMin="5.1">5.1–5.4 mm</quantity>
, MD: 2.0–
<quantity id="9B17ECB0FFEDFFC58F79F8BCFC5A072D" box="[892,979,1810,1837]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="24" pageNumber="25" unit="mm" value="2.1">2.1 mm</quantity>
, AA: 40–43°, SL/MD: 2.5–2.7, AL/AW: 4.1–4.8, AH/S: 2.3–2.4.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFEDFFC48CC2F8F4FE5C0165" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="discussion">
<paragraph id="5C504155FFEDFFC48CC2F8F4FEF3011D" blockId="24.[151,1437,1162,2017]" lastBlockId="25.[151,1437,151,466]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58CC2F8F4FEC50774" bold="true" box="[199,332,1882,1908]" pageId="24" pageNumber="25">Discussion.</emphasis>
<bibRefCitation id="387E3CA4FFEDFFC58D56F8F5FDF70774" author="Hoernes, R. &amp; Auinger, M." box="[339,638,1882,1909]" pageId="24" pageNumber="25" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger (1880)</bibRefCitation>
based the description of
<taxonomicName id="9BEF3AD6FFEDFFC58F91F8F5FBD70774" box="[916,1118,1882,1908]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="lapugyensis">
<emphasis id="6E9B9D47FFEDFFC58F91F8F5FBD70774" box="[916,1118,1882,1908]" italics="true" pageId="24" pageNumber="25">Mitra lapugyensis</emphasis>
</taxonomicName>
on specimens from Lăpugiu de Sus (
<collectingCountry id="24F801C5FFEDFFC58CF5F8D0FED00799" box="[240,345,1918,1945]" name="Romania" pageId="24" pageNumber="25">Romania</collectingCountry>
) and from Steinebrunn (
<collectingCountry id="24F801C5FFEDFFC58E69F8D0FD4C0799" box="[620,709,1918,1945]" name="Austria" pageId="24" pageNumber="25">Austria</collectingCountry>
). In our opinion they are not conspecific, and as first revisers we designate the specimen illustrated by
<bibRefCitation id="387E3CA4FFEDFFC58E3BF80DFCEF07BC" author="Hoernes, R. &amp; Auinger, M." box="[574,870,1954,1981]" pageId="24" pageNumber="25" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger (1880</bibRefCitation>
, pl. 10, fig. 21) from Lăpugiu de Sus as
<typeStatus id="8354FFF7FFEDFFC5892AF80CFA1007BC" box="[1327,1433,1954,1980]" pageId="24" pageNumber="25" type="lectotype">lectotype</typeStatus>
. The specimen from Steinebrunn illustrated by
<bibRefCitation id="387E3CA4FFEDFFC58E9AF869FC4C07E0" author="Hoernes, R. &amp; Auinger, M." box="[671,965,1990,2017]" pageId="24" pageNumber="25" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger (1880</bibRefCitation>
, pl. 10, fig. 22) is described herein as
<taxonomicName id="9BEF3AD6FFEDFFC4896BF868FEE200B1" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" lastPageId="25" lastPageNumber="26" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="fedosovi" status="sp. nov.">
<emphasis id="6E9B9D47FFEDFFC4896BF868FEE200B1" italics="true" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">Bellardithala fedosovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFECFFC48D76FF39FE4000B1" box="[371,457,151,177]" pageId="25" pageNumber="26" rank="species">nov. sp.</taxonomicNameLabel>
<taxonomicName id="9BEF3AD6FFECFFC48DD4FF39FD7800B1" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[465,753,151,177]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="lapugyensis">
<emphasis id="6E9B9D47FFECFFC48DD4FF39FD7800B1" box="[465,753,151,177]" italics="true" pageId="25" pageNumber="26">Bellardithala lapugyensis</emphasis>
</taxonomicName>
differs from congeneric Paratethyan species in its cancellate sculpture and small size. The specimen described by Friedberg (1928) from the Badenian of
<collectingCountry id="24F801C5FFECFFC488B6FF15FA8400D5" box="[1203,1293,187,213]" name="Ukraine" pageId="25" pageNumber="26">Ukraine</collectingCountry>
differs from typical
<taxonomicName id="9BEF3AD6FFECFFC48CEEFF71FE1C00F9" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[235,405,223,249]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="lapugyensis">
<emphasis id="6E9B9D47FFECFFC48CEEFF71FE1C00F9" box="[235,405,223,249]" italics="true" pageId="25" pageNumber="26">M. lapugyensis</emphasis>
</taxonomicName>
in its more slender outline and the narrower spiral cords. It is doubtful that it is conspecific with
<taxonomicName id="9BEF3AD6FFECFFC48CD5FEADFEFA011D" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[208,371,259,285]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="lapugyensis">
<emphasis id="6E9B9D47FFECFFC48CD5FEADFEFA011D" box="[208,371,259,285]" italics="true" pageId="25" pageNumber="26">B. lapugyensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5C504155FFECFFC48CC2FE88FE5C0165" blockId="25.[151,1437,151,466]" pageId="25" pageNumber="26">
<emphasis id="6E9B9D47FFECFFC48CC2FE88FE3F0140" bold="true" box="[199,438,294,320]" pageId="25" pageNumber="26">Palaeoenvironment.</emphasis>
Unknown; the collections from Lăpugiu de Sus indicate a mixing of coastal-inner neritic and middle neritic elements.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFECFFC48CC2FEC0FDF301D1" pageId="25" pageNumber="26" type="distribution">
<paragraph id="5C504155FFECFFC48CC2FEC0FDF301D1" blockId="25.[151,1437,151,466]" pageId="25" pageNumber="26">
<emphasis id="6E9B9D47FFECFFC48CC2FEC0FDEF0189" bold="true" box="[199,614,366,393]" pageId="25" pageNumber="26">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFECFFC48FB4FEC1FBEE0188" box="[945,1127,367,392]" italics="true" pageId="25" pageNumber="26">Korytnica Basin</emphasis>
: Korytnica (
<collectingCountry id="24F801C5FFECFFC488F4FEC1FACB018A" box="[1265,1346,367,394]" name="Poland" pageId="25" pageNumber="26">Poland</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFECFFC4895DFEC1FF5A01AD" author="Baluk, W." pageId="25" pageNumber="26" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997</bibRefCitation>
),?
<emphasis id="6E9B9D47FFECFFC48CFCFE3DFDF001AC" box="[249,633,403,429]" italics="true" pageId="25" pageNumber="26">Ukrainian Fore-Carpathian Basin</emphasis>
: Zborów (
<emphasis id="6E9B9D47FFECFFC48EEAFE3DFCB301AD" box="[751,826,403,429]" italics="true" pageId="25" pageNumber="26">Zboriv</emphasis>
) (
<collectingCountry id="24F801C5FFECFFC48F54FE3DFC2601AE" box="[849,943,403,430]" name="Ukraine" pageId="25" pageNumber="26">Ukraine</collectingCountry>
) (Friedberg 1928);
<emphasis id="6E9B9D47FFECFFC4888DFE3DFA9901AC" box="[1160,1296,403,429]" italics="true" pageId="25" pageNumber="26">Făget Basin</emphasis>
: Lăpugiu de Sus (
<collectingCountry id="24F801C5FFECFFC48CCBFE19FEBE01D2" box="[206,311,439,466]" name="Romania" pageId="25" pageNumber="26">Romania</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFECFFC48D48FE19FDE401D1" author="Hoernes, R. &amp; Auinger, M." box="[333,621,439,465]" pageId="25" pageNumber="26" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger 1880</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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