298 lines
37 KiB
XML
298 lines
37 KiB
XML
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<mods:title id="A398C38BD9B2AB3A6C8165573A4D1A39">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="EC72934923C8708013707043629DAF6C">Landau, Bernard</mods:namePart>
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<treatment id="D446F043FFE7FFCF8C92FD41FA9007F2" ID-DOI="http://doi.org/10.5281/zenodo.5043396" ID-GBIF-Taxon="183225132" ID-Zenodo-Dep="5043396" LSID="urn:lsid:plazi:treatment:D446F043FFE7FFCF8C92FD41FA9007F2" httpUri="http://treatment.plazi.org/id/D446F043FFE7FFCF8C92FD41FA9007F2" lastPageNumber="19" pageId="18" pageNumber="19">
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<taxonomicName id="9BEF3AD6FFE7FFCF8C92FD41FE02030A" ID-CoL="8KRZ4" authority="Harzhauser & Landau, 2021" authorityName="Harzhauser & Landau" authorityYear="2021" box="[151,395,751,778]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="baluki" status="sp. nov.">
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<emphasis id="6E9B9D47FFE7FFCF8C92FD41FE02030A" bold="true" box="[151,395,751,778]" italics="true" pageId="18" pageNumber="19">Bellardithala baluki</emphasis>
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<taxonomicNameLabel id="75A8203CFFE7FFCF8D97FD5EFE79030A" box="[402,496,752,778]" pageId="18" pageNumber="19" rank="species">nov. sp.</taxonomicNameLabel>
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<paragraph id="5C504155FFE7FFCF8C92FCBAFE4F0334" blockId="18.[151,496,751,820]" box="[151,454,788,820]" pageId="18" pageNumber="19">
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<figureCitation id="C4D45DD0FFE7FFCF8C92FCBAFF78032E" box="[151,241,788,814]" captionStart="FIGURE 7" captionStartId="20.[151,250,1753,1777]" captionTargetBox="[185,1403,181,1729]" captionTargetId="figure-19@20.[185,1403,181,1729]" captionTargetPageId="20" captionText="FIGURE 7 A1–A2. Bellardithala baluki nov. sp., holotype, Vienna/Pötzleinsdorf (Austria), NHMW 1859/0047/0015. B1–B2. Bellardithala baluki nov. sp., paratype, Vienna/Pötzleinsdorf (Austria), NHMW 1859/0047/0015b. C1–C2. Bellardithala baluki nov. sp., paratype, Vienna/Pötzleinsdorf (Austria), NHMW 2020/0063/0001. D1–D2. Bellardithala fedosovi nov. sp., holotype, Steinebrunn (Austria), NHMW 1864/0001/0563. E1–E2. Bellardithala fedosovi nov. sp., paratype, Steinebrunn (Austria), NHMW 2020/0052/0001. F1–F2. Bellardithala laubei (Hoernes & Auinger, 1880), lectotype, Bad Vöslau (Austria), NHMW 1999z0076/0002. G1–G2. Bellardithala cf. dacica nov. sp., Niederleis (Austria), NHMW 1866/0001/0806. H1–H2. Bellardithala cf. dacica nov. sp., Niederleis (Austria), NHMW 2020/0054/0001. I1–I2. Bellardithala cf. dacica nov. sp., Niederleis (Austria), NHMW 2020/0054/0002." figureDoi="http://doi.org/10.5281/zenodo.4921856" httpUri="https://zenodo.org/record/4921856/files/figure.png" pageId="18" pageNumber="19">Figs 7A</figureCitation>
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<figureCitation id="C4D45DD0FFE7FFCF8CF4FC88FE92032E" box="[241,283,788,820]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="18" pageNumber="19">
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–A
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, B
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–B
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, C
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–C
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<subScript id="C06B4310FFE7FFCF8DBBFC88FE4F0334" attach="left" box="[446,454,806,820]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
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<paragraph id="5C504155FFE7FFCF8C92FCF7FACC0371" blockId="18.[151,1349,856,881]" box="[151,1349,856,881]" pageId="18" pageNumber="19">
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<treatmentCitationGroup id="7CFF667BFFE7FFCF8C92FCF7FACC0371" box="[151,1349,856,881]" pageId="18" pageNumber="19">
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<taxonomicName id="9BEF3AD6FFE7FFCF8C92FCF7FE940370" ID-CoL="7P94F" authority="Laubei" authorityName="Laubei" box="[151,285,856,880]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFE7FFCF8C92FCF7FE940370" box="[151,285,856,880]" italics="true" pageId="18" pageNumber="19">Mitra Laubei</emphasis>
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nov. form.—
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<treatmentCitation id="DD4E6744FFE7FFCF8DA0FCF6FD580371" author="Hoernes, R. & Auinger, M." box="[421,721,856,881]" page="89" pageId="18" pageNumber="19" year="1880">
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<bibRefCitation id="387E3CA4FFE7FFCF8DA0FCF6FD580371" author="Hoernes, R. & Auinger, M." box="[421,721,856,881]" pageId="18" pageNumber="19" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes & Auinger, 1880: 89</bibRefCitation>
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(pars) [non
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<taxonomicName id="9BEF3AD6FFE7FFCF8F4BFCF6FAB20371" ID-CoL="8KRR7" authority="(Hoernes & Auinger, 1880)" authorityName="Harzhauser & Landau" authorityYear="2021" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" box="[846,1339,856,881]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="laubei">
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<emphasis id="6E9B9D47FFE7FFCF8F4BFCF6FB900370" box="[846,1049,856,880]" italics="true" pageId="18" pageNumber="19">Bellardithala laubei</emphasis>
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(
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<bibRefCitation id="387E3CA4FFE7FFCF8822FCF7FABB0371" author="Hoernes, R. & Auinger, M." box="[1063,1330,856,881]" pageId="18" pageNumber="19" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes & Auinger, 1880</bibRefCitation>
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)
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</taxonomicName>
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].
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</treatmentCitationGroup>
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<subSubSection id="14F512DEFFE7FFCF8C92FC31FD57046E" pageId="18" pageNumber="19" type="materials_examined">
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<emphasis id="6E9B9D47FFE7FFCF8C92FC31FEC103B8" bold="true" box="[151,328,926,953]" pageId="18" pageNumber="19">Type material.</emphasis>
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<typeStatus id="8354FFF7FFE7FFCF8D54FC31FE3703B9" box="[337,446,927,953]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="0C49E92EFFE7FFCF8DCEFC31FDEE03B9" box="[459,615,926,953]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="19" type="Museum">NHMW 1859</specimenCode>
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/0047/0015, SL:
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, MD:
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,
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<collectingRegion id="9E2B8FB7FFE7FFCF883FFC31FB0003B9" box="[1082,1161,927,953]" country="Austria" name="Wien" pageId="18" pageNumber="19">Vienna</collectingRegion>
|
||
/
|
||
<location id="5930178EFFE7FFCF8897FC31FAAE03B9" LSID="urn:lsid:plazi:treatment:D446F043FFE7FFCF8C92FD41FA9007F2:5930178EFFE7FFCF8897FC31FAAE03B9" box="[1170,1319,927,953]" country="Austria" name="Potzleinsdorf" pageId="18" pageNumber="19" stateProvince="Wien">Pötzleinsdorf</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE7FFCF893DFC31FA1903BA" box="[1336,1424,927,954]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
|
||
), figs 7A
|
||
<subScript id="C06B4310FFE7FFCF8CEFFC7BFF7B03E3" attach="left" box="[234,242,981,995]" fontSize="6" pageId="18" pageNumber="19">1</subScript>
|
||
–A
|
||
<subScript id="C06B4310FFE7FFCF8D10FC7BFE9403E3" attach="left" box="[277,285,981,995]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="EC874B08FFE7FFCF8D2EFC6DFF600407" ID-GBIF-Occurrence="3311814442" collectionCode="NHMW" country="Austria" location="Potzleinsdorf" pageId="18" pageNumber="19" specimenCode="NHMW 1859" specimenCount="1" stateProvince="Wien" typeStatus="paratype">
|
||
<typeStatus id="8354FFF7FFE7FFCF8D2EFC6DFE1503DD" box="[299,412,963,989]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCode id="0C49E92EFFE7FFCF8DA3FC6DFDC903DD" box="[422,576,962,989]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="19" type="Museum">NHMW 1859</specimenCode>
|
||
/0047/0015a, SL:
|
||
<quantity id="9B17ECB0FFE7FFCF8F0FFC6CFCEB03DD" box="[778,866,962,989]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="18" pageNumber="19" unit="mm" value="8.5">8.5 mm</quantity>
|
||
, MD:
|
||
<quantity id="9B17ECB0FFE7FFCF8FACFC6DFB8803DD" box="[937,1025,963,989]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="18" pageNumber="19" unit="mm" value="2.9">2.9 mm</quantity>
|
||
,
|
||
<collectingRegion id="9E2B8FB7FFE7FFCF8809FC6DFBD203DD" box="[1036,1115,963,989]" country="Austria" name="Wien" pageId="18" pageNumber="19">Vienna</collectingRegion>
|
||
/
|
||
<location id="5930178EFFE7FFCF8861FC6DFB7003DD" LSID="urn:lsid:plazi:treatment:D446F043FFE7FFCF8C92FD41FA9007F2:5930178EFFE7FFCF8861FC6DFB7003DD" box="[1124,1273,963,989]" country="Austria" name="Potzleinsdorf" pageId="18" pageNumber="19" stateProvince="Wien">Pötzleinsdorf</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE7FFCF8902FC6DFAD603DE" box="[1287,1375,963,990]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
|
||
), figs 7B
|
||
<subScript id="C06B4310FFE7FFCF8CBDFC57FF490407" attach="left" box="[184,192,1017,1031]" fontSize="6" pageId="18" pageNumber="19">1</subScript>
|
||
–B
|
||
<subScript id="C06B4310FFE7FFCF8CE4FC57FF600407" attach="right" box="[225,233,1017,1031]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="EC874B08FFE7FFCF8CFDFC49FAB30407" ID-GBIF-Occurrence="3311814311" box="[248,1338,998,1031]" collectionCode="NHMW" country="Austria" location="Potzleinsdorf" pageId="18" pageNumber="19" specimenCode="NHMW 2020" specimenCount="1" stateProvince="Wien" typeStatus="paratype">
|
||
<specimenCode id="0C49E92EFFE7FFCF8CFDFC49FE1A0401" box="[248,403,999,1025]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="18" pageNumber="19" type="Museum">NHMW 2020</specimenCode>
|
||
/0063/0001, SL:
|
||
<quantity id="9B17ECB0FFE7FFCF8E4AFC48FD2E0401" box="[591,679,998,1025]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="18" pageNumber="19" unit="mm" value="8.5">8.5 mm</quantity>
|
||
, MD:
|
||
<quantity id="9B17ECB0FFE7FFCF8EEBFC49FCCF0401" box="[750,838,999,1025]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="18" pageNumber="19" unit="mm" value="2.9">2.9 mm</quantity>
|
||
,
|
||
<collectingRegion id="9E2B8FB7FFE7FFCF8F54FC49FC290401" box="[849,928,999,1025]" country="Austria" name="Wien" pageId="18" pageNumber="19">Vienna</collectingRegion>
|
||
/
|
||
<location id="5930178EFFE7FFCF8FACFC49FBB70401" LSID="urn:lsid:plazi:treatment:D446F043FFE7FFCF8C92FD41FA9007F2:5930178EFFE7FFCF8FACFC49FBB70401" box="[937,1086,999,1025]" country="Austria" name="Potzleinsdorf" pageId="18" pageNumber="19" stateProvince="Wien">Pötzleinsdorf</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE7FFCF8848FC49FB2C0402" box="[1101,1189,999,1026]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
|
||
), figs 7C
|
||
<subScript id="C06B4310FFE7FFCF890CFC57FA980407" attach="left" box="[1289,1297,1017,1031]" fontSize="6" pageId="18" pageNumber="19">1</subScript>
|
||
–C
|
||
<subScript id="C06B4310FFE7FFCF8937FC57FAB30407" attach="left" box="[1330,1338,1017,1031]" fontSize="6" pageId="18" pageNumber="19">2</subScript>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE7FFCF8CC2FBA5FD420426" blockId="18.[151,1437,926,2034]" box="[199,715,1034,1062]" pageId="18" pageNumber="19">
|
||
<materialsCitation id="EC874B08FFE7FFCF8CC2FBA5FD4E0426" ID-GBIF-Occurrence="3311814456" box="[199,711,1034,1062]" country="Austria" location="Potzleinsdorf" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Wien" typeStatus="holotype">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2FBA5FEEF0424" bold="true" box="[199,358,1034,1061]" pageId="18" pageNumber="19">
|
||
<typeStatus id="8354FFF7FFE7FFCF8CC2FBA5FE8B0425" box="[199,258,1035,1061]" pageId="18" pageNumber="19">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingRegion id="9E2B8FB7FFE7FFCF8D69FBA5FE320425" box="[364,443,1035,1061]" country="Austria" name="Wien" pageId="18" pageNumber="19">Vienna</collectingRegion>
|
||
/
|
||
<location id="5930178EFFE7FFCF8DC6FBA5FDD10425" LSID="urn:lsid:plazi:treatment:D446F043FFE7FFCF8C92FD41FA9007F2:5930178EFFE7FFCF8DC6FBA5FDD10425" box="[451,600,1035,1061]" country="Austria" name="Potzleinsdorf" pageId="18" pageNumber="19" stateProvince="Wien">Pötzleinsdorf</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE7FFCF8E62FBA5FD360426" box="[615,703,1035,1062]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE7FFCF8CC2FB81FD580449" blockId="18.[151,1437,926,2034]" box="[199,721,1071,1097]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2FB81FEE60449" bold="true" box="[199,367,1071,1097]" pageId="18" pageNumber="19">
|
||
<typeStatus id="8354FFF7FFE7FFCF8CC2FB81FE8B0449" box="[199,258,1071,1097]" pageId="18" pageNumber="19">Type</typeStatus>
|
||
stratum.
|
||
</emphasis>
|
||
Silty sand of the Baden Group.
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE7FFCF8CC2FBFCFD57046E" blockId="18.[151,1437,926,2034]" box="[199,734,1106,1134]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2FBFCFF74046C" bold="true" box="[199,253,1106,1132]" pageId="18" pageNumber="19">Age.</emphasis>
|
||
Middle Miocene, Badenian (Serravallian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE7FFCF8CC2FBD9FD3F04B5" pageId="18" pageNumber="19" type="etymology">
|
||
<paragraph id="5C504155FFE7FFCF8CC2FBD9FD3F04B5" blockId="18.[151,1437,926,2034]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2FBD9FEC70491" bold="true" box="[199,334,1143,1169]" pageId="18" pageNumber="19">Etymology.</emphasis>
|
||
In honour of Wacław Bałuk (University of Warsaw, Warszawa,
|
||
<collectingCountry id="24F801C5FFE7FFCF8824FBD9FBF80492" box="[1057,1137,1143,1170]" name="Poland" pageId="18" pageNumber="19">Poland</collectingCountry>
|
||
) in recognition of his contributions on Paratethyan gastropod systematics.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE7FFCF8CC2FB10FAD304FD" pageId="18" pageNumber="19" type="diagnosis">
|
||
<paragraph id="5C504155FFE7FFCF8CC2FB10FAD304FD" blockId="18.[151,1437,926,2034]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2FB10FECB04D8" bold="true" box="[199,322,1214,1240]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8D4EFB11FDB004D9" box="[331,569,1215,1241]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="6E9B9D47FFE7FFCF8D4EFB11FE5604D9" box="[331,479,1215,1241]" italics="true" pageId="18" pageNumber="19">Bellardithala</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of medium size, moderately slender fusiform shell, with cyrtoconoid spire, and beaded sculpture differing from congeneric species by its slender bullet-like spire and delicate sculpture.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE7FFCF8CC2FAA8FE2B06AD" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="5C504155FFE7FFCF8CC2FAA8FD310665" blockId="18.[151,1437,926,2034]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2FAA8FED00520" bold="true" box="[199,345,1286,1312]" pageId="18" pageNumber="19">Description.</emphasis>
|
||
Shell medium-sized, moderately slender fusiform with cyrtoconoid spire and narrowly incised suture. Protoconch unknown. Teleoconch of 4.5 weakly convex whorls, with periphery at abapical third of whorl, rapidly increasing in height. First teleoconch whorl with blunt axial ribs, crossed by six weak spiral threads, most prominent in axial interspaces. Regularly spaced axial ribs crossed by ten spiral cords on penultimate whorl, forming low beads at intersections. Last whorl weakly convex, with faint subsutural concavity, weakly constricted at base, bearing about 22 beaded spiral cords, adapical subsutural spiral cord most prominent, bearing prosocline nodes. Cords over fasciole prominent, where they develop as extensions of columellar folds. Aperture moderately narrow with shallow anal sinus. Columellar callus indistinct, poorly delimited. Columella with four prominent columellar folds. Outer lip weakly thickened; ten delicate denticles within. Siphonal canal moderately long, straight or bent slightly to left, with distinct siphonal notch.
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE7FFCF8CC2F9C0FE2B06AD" blockId="18.[151,1437,926,2034]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2F9C0FDBE0688" bold="true" box="[199,567,1646,1673]" pageId="18" pageNumber="19">Shell measurements and ratios</emphasis>
|
||
. SL:
|
||
<quantity id="9B17ECB0FFE7FFCF8E73F9C0FC890689" box="[630,768,1646,1673]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.5" metricValueMin="8.1" pageId="18" pageNumber="19" unit="mm" value="8.3" valueMax="8.5" valueMin="8.1">8.1–8.5 mm</quantity>
|
||
, MD:
|
||
<quantity id="9B17ECB0FFE7FFCF8F4CF9C1FC5A0689" box="[841,979,1647,1673]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="18" pageNumber="19" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.7–2.9 mm</quantity>
|
||
, AA: 40–45°, SL/MD: 3.0–3.1, AL/AW: 4.6–5.2, AH/S: 2.1–2.2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE7FFCF8CC2F918FEA407CD" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="5C504155FFE7FFCF8CC2F918FEA407CD" blockId="18.[151,1437,926,2034]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2F918FEC506D0" bold="true" box="[199,332,1718,1744]" pageId="18" pageNumber="19">Discussion.</emphasis>
|
||
Specimens of
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8DF8F919FD6B06D1" authority="Harzhauser & Landau, 2021" authorityName="Harzhauser & Landau" authorityYear="2021" box="[509,738,1719,1745]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="baluki" status="sp. nov.">
|
||
<emphasis id="6E9B9D47FFE7FFCF8DF8F919FD6B06D1" box="[509,738,1719,1745]" italics="true" pageId="18" pageNumber="19">Bellardithala baluki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="75A8203CFFE7FFCF8EE8F919FCCE06D1" box="[749,839,1719,1745]" pageId="18" pageNumber="19" rank="species">nov. sp.</taxonomicNameLabel>
|
||
were mixed with
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF881AF919FB0106D1" authorityName="Harzhauser & Landau" authorityYear="2021" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" box="[1055,1160,1719,1745]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE7FFCF881AF919FB0106D1" box="[1055,1160,1719,1745]" italics="true" pageId="18" pageNumber="19">B. laubei</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="387E3CA4FFE7FFCF88BFF919FF6C06F5" author="Hoernes, R. & Auinger, M." pageId="18" pageNumber="19" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes & Auinger (1880)</bibRefCitation>
|
||
. However,
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8D63F975FE4406F5" authority="Harzhauser & Landau, 2021" authorityName="Harzhauser & Landau" authorityYear="2021" box="[358,461,1755,1781]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="baluki" status="sp. nov.">
|
||
<emphasis id="6E9B9D47FFE7FFCF8D63F975FE4406F5" box="[358,461,1755,1781]" italics="true" pageId="18" pageNumber="19">B. baluki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="75A8203CFFE7FFCF8DD3F975FDA406F5" box="[470,557,1755,1781]" pageId="18" pageNumber="19" rank="species">nov. sp.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8EC3F975FBE506F5" authority="(Hoernes & Auinger, 1880)" authorityName="Harzhauser & Landau" authorityYear="2021" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" box="[710,1132,1755,1781]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE7FFCF8EC3F975FCA506F5" box="[710,812,1755,1781]" italics="true" pageId="18" pageNumber="19">B. laubei</emphasis>
|
||
(
|
||
<bibRefCitation id="387E3CA4FFE7FFCF8F39F975FBED06F5" author="Hoernes, R. & Auinger, M." box="[828,1124,1755,1781]" pageId="18" pageNumber="19" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes & Auinger, 1880</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, as redefined herein, in its broader shell, beaded sculpture especially on the last whorl, and it develops only 4.5 teleoconch whorls instead of
|
||
<quantity id="9B17ECB0FFE7FFCF8C92F88CFF50073D" box="[151,217,1826,1853]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" pageId="18" pageNumber="19" unit="in" value="5.5">5.5 in</quantity>
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8CE7F88DFEC3073D" authorityName="Harzhauser & Landau" authorityYear="2021" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" box="[226,330,1827,1853]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CE7F88DFEC3073D" box="[226,330,1827,1853]" italics="true" pageId="18" pageNumber="19">B. laubei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8D5FF88DFDDD073D" authority="Harzhauser & Landau, 2021" authorityName="Harzhauser & Landau" authorityYear="2021" box="[346,596,1827,1853]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="fedosovi" status="sp. nov.">
|
||
<emphasis id="6E9B9D47FFE7FFCF8D5FF88DFDDD073D" box="[346,596,1827,1853]" italics="true" pageId="18" pageNumber="19">Bellardithala fedosovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="75A8203CFFE7FFCF8E5BF88DFD3F073D" box="[606,694,1827,1853]" pageId="18" pageNumber="19" rank="species">nov. sp.</taxonomicNameLabel>
|
||
is similar in shell shape, but differs in its smaller size, the more convex last whorl, the wider interspaces between the axial ribs, and the overall coarser sculpture. The
|
||
<typeStatus id="8354FFF7FFE7FFCF8908F8E9FAB40761" box="[1293,1341,1863,1889]" pageId="18" pageNumber="19">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8C92F8C5FE5A0785" authority="(Brocchi, 1814)" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[151,467,1899,1925]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="obsoleta">
|
||
<emphasis id="6E9B9D47FFE7FFCF8C92F8C5FE9E0785" box="[151,279,1899,1925]" italics="true" pageId="18" pageNumber="19">B. obsoleta</emphasis>
|
||
(
|
||
<bibRefCitation id="387E3CA4FFE7FFCF8D2DF8C5FE420785" author="Brocchi, G." box="[296,459,1899,1925]" pageId="18" pageNumber="19" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Mediterranean Pliocene, differs in being slightly broader and the axial ribs are well developed and elevated. In
|
||
<taxonomicName id="9BEF3AD6FFE7FFCF8E29F821FD1807A9" authorityName="Harzhauser & Landau" authorityYear="2021" box="[556,657,1935,1961]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="baluki">
|
||
<emphasis id="6E9B9D47FFE7FFCF8E29F821FD1807A9" box="[556,657,1935,1961]" italics="true" pageId="18" pageNumber="19">B. baluki</emphasis>
|
||
</taxonomicName>
|
||
axial sculpture consists of axial rows of small beads, but do not form elevated ribs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE7FFCF8CC2F878FA9007F2" box="[199,1305,2006,2034]" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="5C504155FFE7FFCF8CC2F878FA9007F2" blockId="18.[151,1437,926,2034]" box="[199,1305,2006,2034]" pageId="18" pageNumber="19">
|
||
<emphasis id="6E9B9D47FFE7FFCF8CC2F878FDE307F1" bold="true" box="[199,618,2006,2033]" pageId="18" pageNumber="19">Distribution in Central Paratethys.</emphasis>
|
||
Badenian (middle Miocene):
|
||
<collectingRegion id="9E2B8FB7FFE7FFCF8FBCF879FB8107F1" box="[953,1032,2007,2033]" country="Austria" name="Wien" pageId="18" pageNumber="19">Vienna</collectingRegion>
|
||
/Pötzleinsdorf (
|
||
<collectingCountry id="24F801C5FFE7FFCF88B1F879FA8407F2" box="[1204,1293,2007,2034]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |