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<mods:title id="6E7260103594084C4F4E63F10C921471">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<treatment id="D446F043FFE4FFCF8C92FECDFCC502A9" ID-DOI="http://doi.org/10.5281/zenodo.5043394" ID-GBIF-Taxon="183225131" ID-Zenodo-Dep="5043394" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9" httpUri="http://treatment.plazi.org/id/D446F043FFE4FFCF8C92FECDFCC502A9" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<taxonomicName id="9BEF3AD6FFE4FFCC8C92FECDFE05017E" ID-CoL="8KTGW" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[151,396,355,382]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="kovaci" status="sp. nov.">
<emphasis id="6E9B9D47FFE4FFCC8C92FECDFE05017E" bold="true" box="[151,396,355,382]" italics="true" pageId="17" pageNumber="18">Bellardithala kovaci</emphasis>
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<figureCitation id="C4D45DD0FFE4FFCC8C92FE29FF7B01A2" box="[151,242,391,418]" captionStart="FIGURE 4" captionStartId="10.[151,250,1550,1574]" captionTargetBox="[162,1423,181,1527]" captionTargetId="figure-19@10.[162,1425,181,1527]" captionTargetPageId="10" captionText="FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes &amp; Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes &amp; Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes &amp; Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037." figureDoi="http://doi.org/10.5281/zenodo.4921850" httpUri="https://zenodo.org/record/4921850/files/figure.png" pageId="17" pageNumber="18">Figs 4C</figureCitation>
,
<figureCitation id="C4D45DD0FFE4FFCC8CF8FE29FEA901A1" box="[253,288,391,417]" captionStart="FIGURE 6" captionStartId="16.[151,250,1717,1741]" captionTargetBox="[151,1436,181,1693]" captionTargetId="figure-19@16.[151,1436,181,1693]" captionTargetPageId="16" captionText="FIGURE 6. A1A2. Bellardithala boehmi (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360342. B1B2. Bellardithala boehmi (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0059/0001. C1C2. Bellardithala boehmi (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 1854/0035/0107. D1D2. Bellardithala kovaci nov. sp., holotype, Steinebrunn (Austria), NHMW 1846/0037/0114. E1E2. Bellardithala kovaci nov. sp., paratype, Sedlec (= Porzteich) (Czech Republic), NHMW 1858/0047/0007. F1F2. Bellardithala kovaci nov. sp., paratype, Forchtenau (Austria), NHMW 1866/0001/1019. G1G2. Bellardithala kovaci nov. sp., paratype, Steinebrunn (Austria), NHMW 1871/0010/0331. H1H2. Bellardithala kovaci nov. sp., paratype, Steinebrunn (Austria), NHMW 1871/0011/0004. I1I2. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001." figureDoi="http://doi.org/10.5281/zenodo.4921854" httpUri="https://zenodo.org/record/4921854/files/figure.png" pageId="17" pageNumber="18">6D</figureCitation>
<figureCitation id="C4D45DD0FFE4FFCC8D1AFE34FEC301A2" box="[287,330,392,424]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="17" pageNumber="18">
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D
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<figureCitation id="C4D45DD0FFE4FFCC8D4FFE34FEDB01A8" box="[330,338,410,424]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="17" pageNumber="18">
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, E
<figureCitation id="C4D45DD0FFE4FFCC8D74FE34FE1101A2" box="[369,408,392,424]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="17" pageNumber="18">
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E
</figureCitation>
<figureCitation id="C4D45DD0FFE4FFCC8D9DFE34FE2901A8" box="[408,416,410,424]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="17" pageNumber="18">
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, F
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F
</figureCitation>
<figureCitation id="C4D45DD0FFE4FFCC8DE1FE34FE6501A8" box="[484,492,410,424]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="17" pageNumber="18">
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, G
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G
</figureCitation>
<figureCitation id="C4D45DD0FFE4FFCC8E3DFE34FDC901A8" box="[568,576,410,424]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="17" pageNumber="18">
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, H
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H
</figureCitation>
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, I
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I
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<figureCitation id="C4D45DD0FFE4FFCC8EC9FE34FD5D01A8" box="[716,724,410,424]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="17" pageNumber="18">
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[
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<emphasis id="6E9B9D47FFE4FFCC8CA5FE69FF5301DE" box="[160,218,455,478]" italics="true" pageId="17" pageNumber="18">Mitra</emphasis>
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]
<emphasis id="6E9B9D47FFE4FFCC8CEDFE69FEC401DF" box="[232,333,455,479]" italics="true" pageId="17" pageNumber="18">candidula</emphasis>
[Partsch]—
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<bibRefCitation id="387E3CA4FFE4FFCC8DC2FE69FDF801DF" author="Hornes, M." box="[455,625,455,479]" pageId="17" pageNumber="18" refId="ref58087" refString="Hornes, M. (1848) Verzeichnis der Fossil-Reste aus 135 Fundorten des Tertiar-Beckens von Wien. Braumuller, Wien, 44 pp. [Appendix to Czjzek, J. (1849) Erlauterungen zur geognostischen Karte der Umgebungen Wiens. Braumuller, Wien, 105 pp., https: // books. google. at / books? id = 1 PL 0 QAAACAAJ &amp; printsec = frontcover &amp; hl = de &amp; source = gbs _ ge _ summary _ r &amp; cad = 0 # v = onepage &amp; q &amp; f = false]" type="book" year="1848">Hörnes 1848: 17</bibRefCitation>
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[
<emphasis id="6E9B9D47FFE4FFCC8E85FE69FC8601DF" box="[640,783,455,479]" italics="true" pageId="17" pageNumber="18">nomen nudum</emphasis>
].
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<taxonomicName id="9BEF3AD6FFE4FFCC8C92FE48FEFB01FD" authority="Bronn" authorityName="Bronn" box="[151,370,485,509]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8C92FE48FEA401FD" box="[151,301,485,509]" italics="true" pageId="17" pageNumber="18">Mitra obsoleta</emphasis>
Bronn
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[sic]—
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, pl. 10, figs 32ab [
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<taxonomicName id="9BEF3AD6FFE4FFCC8F7CFE4BFB3801FD" authority="Brocchi, 1814" authorityName="Brocchi" authorityYear="1814" box="[889,1201,485,509]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8F7CFE4BFB9F01FD" box="[889,1046,485,509]" italics="true" pageId="17" pageNumber="18">Voluta obsoleta</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8819FE4BFB3801FD" author="Brocchi, G." box="[1052,1201,485,509]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
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].
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[
<taxonomicName id="9BEF3AD6FFE4FFCC8CA5FDABFF53021C" ID-CoL="7P94F" box="[160,218,517,540]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFE4FFCC8CA5FDABFF53021C" box="[160,218,517,540]" italics="true" pageId="17" pageNumber="18">Mitra</emphasis>
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<taxonomicName id="9BEF3AD6FFE4FFCC8CEDFDAAFE0C021C" authority="Brocc." authorityName="Brocchi" authorityYear="1814" box="[232,389,516,540]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8CEDFDAAFEB7021C" box="[232,318,516,540]" italics="true" pageId="17" pageNumber="18">obsoleta</emphasis>
Brocc.
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<treatmentCitation id="DD4E6744FFE4FFCC8D9AFDAAFDCE021C" author="Auinger, M." box="[415,583,516,540]" page="8" pageId="17" pageNumber="18" year="1871">
<bibRefCitation id="387E3CA4FFE4FFCC8D9AFDAAFDCE021C" author="Auinger, M." box="[415,583,516,540]" pageId="17" pageNumber="18" pagination="1 - 27" refId="ref53961" refString="Auinger, M. (1871) Tabellarisches Verzeichnis der bisher aus den Tertiarbildungen der Markgrafschaft Mahren bekannt gewordenen fossilen Conchylien auf Grundlage der Sammlung des k. k. Hof-Mineralien-Cabinetes. Naturforschender Verein Brunn, Verhandlungen, 9, 1 - 27. [https: // opac. geologie. ac. at / ais 312 / dokumente / Auinger _ 1871 _ Tertiaerbildungen. pdf]" type="journal article" year="1871">Auinger 1871: 8</bibRefCitation>
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[
<emphasis id="6E9B9D47FFE4FFCC8E50FDABFDF2021C" box="[597,635,517,540]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8E87FDAAFC9F021C" author="Brocchi, G." box="[642,790,516,540]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
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<paragraph id="5C504155FFE4FFCC8C92FD8DFCE9023B" blockId="17.[151,1211,455,785]" box="[151,864,546,571]" pageId="17" pageNumber="18">
[
<taxonomicName id="9BEF3AD6FFE4FFCC8CA5FD8DFF77023B" ID-CoL="924DW" authorityName="Swainson" authorityYear="1840" box="[160,254,547,571]" class="Insecta" family="Nymphalidae" genus="Callithea" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="6E9B9D47FFE4FFCC8CA5FD8DFF77023B" box="[160,254,547,571]" italics="true" pageId="17" pageNumber="18">Callithea</emphasis>
</taxonomicName>
]
<taxonomicName id="9BEF3AD6FFE4FFCC8D08FD8DFE23023A" authority="Brocc." authorityName="Brocchi" authorityYear="1814" box="[269,426,547,571]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8D08FD8DFEEA023B" box="[269,355,547,571]" italics="true" pageId="17" pageNumber="18">obsoleta</emphasis>
Brocc.
</taxonomicName>
—Hoernes 1880: 125 [
<emphasis id="6E9B9D47FFE4FFCC8E90FD8DFD32023A" box="[661,699,547,570]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8EC4FD8DFCDF023B" author="Brocchi, G." box="[705,854,547,571]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
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<emphasis id="6E9B9D47FFE4FFCC8C92FDECFEA40259" box="[151,301,577,601]" italics="true" pageId="17" pageNumber="18">Mitra obsoleta</emphasis>
Brocc.
</taxonomicName>
<treatmentCitation id="DD4E6744FFE4FFCC8D8BFDECFD3C0259" author="Hoernes, R. &amp; Auinger, M." box="[398,693,577,601]" page="88" pageId="17" pageNumber="18" year="1880">
<bibRefCitation id="387E3CA4FFE4FFCC8D8BFDECFD3C0259" author="Hoernes, R. &amp; Auinger, M." box="[398,693,577,601]" pageId="17" pageNumber="18" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger 1880: 88</bibRefCitation>
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[
<emphasis id="6E9B9D47FFE4FFCC8EC1FDECFD630259" box="[708,746,578,601]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8EF4FDEFFC0F0259" author="Brocchi, G." box="[753,902,577,601]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
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</paragraph>
<paragraph id="5C504155FFE4FFCC8C92FDCFFCAA0278" blockId="17.[151,1211,455,785]" box="[151,803,608,632]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="7CFF667BFFE4FFCC8C92FDCFFCAA0278" box="[151,803,608,632]" pageId="17" pageNumber="18">
<taxonomicName id="9BEF3AD6FFE4FFCC8C92FDCFFEFF0278" authority="Braun." authorityName="Braun." box="[151,374,608,632]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8C92FDCFFEA40278" box="[151,301,608,632]" italics="true" pageId="17" pageNumber="18">Mitra obsoleta</emphasis>
Braun.
</taxonomicName>
<treatmentCitation id="DD4E6744FFE4FFCC8D8AFDCEFDC00278" author="Rzehak, A." box="[399,585,608,632]" page="256" pageId="17" pageNumber="18" year="1894">
<bibRefCitation id="387E3CA4FFE4FFCC8D8AFDCEFDC00278" author="Rzehak, A." box="[399,585,608,632]" pageId="17" pageNumber="18" pagination="252 - 262" refId="ref61539" refString="Rzehak, A. (1894) Uber einige neue Fossilienfundorte im mahrischen Miocan. Verhandlungen des naturforschenden Vereines in Brunn, 33, 252 - 262. [http: // www. zobodat. at / pdf / Verh-naturf-Ver-Bruenn _ 33 _ 0252 - 0262. pdf]" type="journal article" year="1894">Rzehak 1894: 256</bibRefCitation>
</treatmentCitation>
[
<emphasis id="6E9B9D47FFE4FFCC8E5DFDCFFDF70278" box="[600,638,609,632]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8E81FDCEFC900278" author="Brocchi, G." box="[644,793,608,632]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
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</paragraph>
<paragraph id="5C504155FFE4FFCC8C92FDD1FCC70297" blockId="17.[151,1211,455,785]" box="[151,846,639,663]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="7CFF667BFFE4FFCC8C92FDD1FCC70297" box="[151,846,639,663]" pageId="17" pageNumber="18">
<taxonomicName id="9BEF3AD6FFE4FFCC8C92FDD1FE0A0297" ID-CoL="8V284" authority="Brocch." authorityName="Brocch." box="[151,387,639,663]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8C92FDD1FEA60297" box="[151,303,639,663]" italics="true" pageId="17" pageNumber="18">Thala obsoleta</emphasis>
Brocch.
</taxonomicName>
<treatmentCitation id="DD4E6744FFE4FFCC8D99FDD1FDFD0297" author="Meznerics, I." box="[412,628,639,663]" page="344" pageId="17" pageNumber="18" year="1933">
<bibRefCitation id="387E3CA4FFE4FFCC8D99FDD1FDFD0297" author="Meznerics, I." box="[412,628,639,663]" pageId="17" pageNumber="18" pagination="319 - 359" refId="ref59495" refString="Meznerics, I. (1933) Die Minutien der tortonischen Ablagerungen von Steinabrunn in Niederosterreich. Annalen des Naturhistorischen Museums Wien, 46, 319 - 359. [https: // www. zobodat. at / pdf / ANNA _ 46 _ 0319 - 0359. pdf]" type="journal article" year="1933">Meznerics 1933: 344</bibRefCitation>
</treatmentCitation>
[
<emphasis id="6E9B9D47FFE4FFCC8E87FDD1FD210296" box="[642,680,639,662]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8EAAFDD1FCCA0297" author="Brocchi, G." box="[687,835,639,663]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
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</paragraph>
<paragraph id="5C504155FFE4FFCC8C92FD33FC6802B5" blockId="17.[151,1211,455,785]" box="[151,993,669,693]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="7CFF667BFFE4FFCC8C92FD33FC6802B5" box="[151,993,669,693]" pageId="17" pageNumber="18">
<taxonomicName id="9BEF3AD6FFE4FFCC8C92FD33FDB502B5" authority="(Brocc.)" baseAuthorityName="Brocc." box="[151,572,669,693]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoletum" subGenus="Costellaria">
<emphasis id="6E9B9D47FFE4FFCC8C92FD33FF6702B5" box="[151,238,669,693]" italics="true" pageId="17" pageNumber="18">Vexillum</emphasis>
(
<emphasis id="6E9B9D47FFE4FFCC8CF8FD33FEE602B5" box="[253,367,669,693]" italics="true" pageId="17" pageNumber="18">Costellaria</emphasis>
)
<emphasis id="6E9B9D47FFE4FFCC8D7BFD33FE6F02B5" box="[382,486,669,693]" italics="true" pageId="17" pageNumber="18">obsoletum</emphasis>
(Brocc.)
</taxonomicName>
<treatmentCitation id="DD4E6744FFE4FFCC8E52FD33FC8E02B5" author="Sieber, R." box="[599,775,669,693]" page="244" pageId="17" pageNumber="18" year="1956">
<bibRefCitation id="387E3CA4FFE4FFCC8E52FD33FC8E02B5" author="Sieber, R." box="[599,775,669,693]" pageId="17" pageNumber="18" pagination="236 - 249" refId="ref61969" refString="Sieber, R. (1956) Die Tortonfauna von Mattersburg und Forchtenau (Burgenland). Verhandlungen der Geologischen Bunsdesanstalt, 1956, 236 - 249. [https: // opac. geologie. ac. at / ais 312 / dokumente / VH 1956 _ 236 _ A. pdf]" type="journal article" year="1956">Sieber 1956: 244</bibRefCitation>
</treatmentCitation>
[
<emphasis id="6E9B9D47FFE4FFCC8F10FD30FCB202B5" box="[789,827,670,693]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8F47FD33FC5E02B5" author="Brocchi, G." box="[834,983,669,693]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
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</paragraph>
<paragraph id="5C504155FFE4FFCC8C92FD12FCD002D4" blockId="17.[151,1211,455,785]" box="[151,857,700,724]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8C92FD12FF3B02D4" box="[151,178,700,724]" italics="true" pageId="17" pageNumber="18">Th</emphasis>
[
<emphasis id="6E9B9D47FFE4FFCC8CBFFD12FF5302D4" box="[186,218,700,724]" italics="true" pageId="17" pageNumber="18">ala</emphasis>
].
<taxonomicName id="9BEF3AD6FFE4FFCC8CF5FD12FE2202D4" ID-CoL="8V284" authority="(Brocchi)" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[240,427,700,724]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8CF5FD12FECF02D4" box="[240,326,700,724]" italics="true" pageId="17" pageNumber="18">obsoleta</emphasis>
(Brocchi)
</taxonomicName>
<bibRefCitation id="387E3CA4FFE4FFCC8DC0FD12FDF602D4" author="Sieber, R." box="[453,639,700,724]" pageId="17" pageNumber="18" pagination="123 - 192" refId="ref62023" refString="Sieber, R. (1958 a) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]" type="journal article" year="1958">Sieber 1958a: 153</bibRefCitation>
[
<emphasis id="6E9B9D47FFE4FFCC8E8BFD13FD3D02D4" box="[654,692,701,724]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8EBFFD12FCC602D4" author="Brocchi, G." box="[698,847,700,724]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
</paragraph>
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<emphasis id="6E9B9D47FFE4FFCC8C92FD75FEA602F3" box="[151,303,731,755]" italics="true" pageId="17" pageNumber="18">Thala obsoleta</emphasis>
(Brocc.)
</taxonomicName>
<treatmentCitation id="DD4E6744FFE4FFCC8D9AFD75FDD502F3" author="Sieber, R." box="[415,604,730,755]" page="149" pageId="17" pageNumber="18" year="1958">
<bibRefCitation id="387E3CA4FFE4FFCC8D9AFD75FDD502F3" author="Sieber, R." box="[415,604,730,755]" pageId="17" pageNumber="18" pagination="142 - 155" refId="ref62086" refString="Sieber, R. (1958 b) Die Tortonfauna von Steinabrunn bei Drasenhofen (Bez. Mistelbach, N. - O.). Verhandlungen der Geologischen Bundesanstalt, 1958, 142 - 155. [https: // opac. geologie. ac. at / ais 312 / dokumente / VH 1958 _ 142 _ A. pdf]" type="journal article" year="1958">Sieber 1958b: 149</bibRefCitation>
</treatmentCitation>
[
<emphasis id="6E9B9D47FFE4FFCC8E6FFD75FD1902F2" box="[618,656,731,754]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8E92FD75FCA502F3" author="Brocchi, G." box="[663,812,731,755]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
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</paragraph>
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<treatmentCitationGroup id="7CFF667BFFE4FFCC8C92FD57FC650311" box="[151,1004,761,785]" pageId="17" pageNumber="18">
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<emphasis id="6E9B9D47FFE4FFCC8C92FD57FEA60311" box="[151,303,761,785]" italics="true" pageId="17" pageNumber="18">Thala obsoleta</emphasis>
(
<bibRefCitation id="387E3CA4FFE4FFCC8D39FD57FE450311" author="Brocchi, G." box="[316,460,761,785]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
)
</taxonomicName>
<treatmentCitation id="DD4E6744FFE4FFCC8DE8FD57FD050311" author="Baluk, W." box="[493,652,761,785]" page="39" pageId="17" pageNumber="18" year="1997">
<bibRefCitation id="387E3CA4FFE4FFCC8DE8FD57FD050311" author="Baluk, W." box="[493,652,761,785]" pageId="17" pageNumber="18" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997: 39</bibRefCitation>
</treatmentCitation>
, pl. 11, fig. 2 [
<emphasis id="6E9B9D47FFE4FFCC8F24FD54FCCE0311" box="[801,839,762,785]" italics="true" pageId="17" pageNumber="18">non</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8F48FD57FC6B0311" author="Brocchi, G." box="[845,994,761,785]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFE4FFCC8C92FC94FD490499" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="5C504155FFE4FFCC8C92FC94FA9803E5" blockId="17.[151,1437,826,2041]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8C92FC94FECF0354" bold="true" box="[151,326,826,852]" pageId="17" pageNumber="18">Type material.</emphasis>
<materialsCitation id="EC874B08FFE4FFCC8D4BFC94FD68037F" ID-GBIF-Occurrence="3311814427" collectionCode="NHMW" country="Austria" location="Steinebrunn" pageId="17" pageNumber="18" specimenCode="NHMW 1846" specimenCount="1" typeStatus="holotype">
<typeStatus id="8354FFF7FFE4FFCC8D4BFC94FE320354" box="[334,443,826,852]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
:
<specimenCode id="0C49E92EFFE4FFCC8DC3FC95FDE90354" box="[454,608,826,853]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW 1846</specimenCode>
/0037/0114, SL:
<quantity id="9B17ECB0FFE4FFCC8F25FC94FCF00355" box="[800,889,826,853]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="17" pageNumber="18" unit="mm" value="8.3">8.3 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE4FFCC8FC6FC94FB950355" box="[963,1052,826,853]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="17" pageNumber="18" unit="mm" value="3.1">3.1 mm</quantity>
,
<location id="5930178EFFE4FFCC882CFC94FB390354" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC882CFC94FB390354" box="[1065,1200,826,852]" country="Austria" name="Steinebrunn" pageId="17" pageNumber="18">Steinebrunn</location>
(
<collectingCountry id="24F801C5FFE4FFCC88C5FC94FA910355" box="[1216,1304,826,853]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
), illustrated in
<bibRefCitation id="387E3CA4FFE4FFCC8CB2FCF1FEED0379" author="Hornes, M." box="[183,356,862,889]" pageId="17" pageNumber="18" pagination="43 - 208" refId="ref58243" refString="Hornes, M. (1852 b) Die fossilen Mollusken des Tertiar-Beckens von Wien. Abhandlungen der k. k Geologischen Reichsanstalt, 3 - 4, 43 - 208, pls. 6 - 20. [https: // opac. geologie. ac. at / ais 312 / dokumente / Abh- 3 - GRA-gesamt. pdf]" type="journal article" year="1852">Hörnes (1852b</bibRefCitation>
, pl. 10, figs 32ab), figs 6D
<subScript id="C06B4310FFE4FFCC8EAAFCDFFD3E037F" attach="left" box="[687,695,881,895]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
D
<subScript id="C06B4310FFE4FFCC8EDCFCDFFD68037F" attach="left" box="[729,737,881,895]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
</materialsCitation>
.
<materialsCitation id="EC874B08FFE4FFCC8EF7FCF1FCAA03A3" ID-GBIF-Occurrence="3311814364" collectionCode="NHMW" country="Czech Republic" location="Sedlec" municipality="Porzteich" pageId="17" pageNumber="18" specimenCode="NHMW 1858" specimenCount="1" typeStatus="paratype">
<typeStatus id="8354FFF7FFE4FFCC8EF7FCF1FCEA0379" box="[754,867,863,889]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="0C49E92EFFE4FFCC8F74FCF1FB850379" box="[881,1036,862,889]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW 1858</specimenCode>
/0047/0007, SL:
<quantity id="9B17ECB0FFE4FFCC88D4FCF0FAA50379" box="[1233,1324,862,889]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="17" pageNumber="18" unit="mm" value="8.9">8.9 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE4FFCC897CFCF0FF4C039D" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="17" pageNumber="18" unit="mm" value="3.3">3.3 mm</quantity>
,
<location id="5930178EFFE4FFCC8CD1FC2CFE96039C" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC8CD1FC2CFE96039C" box="[212,287,898,924]" country="Czech Republic" municipality="Porzteich" name="Sedlec" pageId="17" pageNumber="18">Sedlec</location>
(=
<collectingMunicipality id="BC34DB2FFFE4FFCC8D4EFC2CFE3E039D" box="[331,439,898,925]" pageId="17" pageNumber="18">Porzteich</collectingMunicipality>
) (
<collectingCountry id="24F801C5FFE4FFCC8DD5FC2CFD02039D" box="[464,651,898,925]" name="Czech Republic" pageId="17" pageNumber="18">Czech Republic</collectingCountry>
), figs 6E
<subScript id="C06B4310FFE4FFCC8EF1FC3BFD7503A3" attach="left" box="[756,764,917,931]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
E
<subScript id="C06B4310FFE4FFCC8F1EFC3BFCAA03A3" attach="left" box="[795,803,917,931]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
</materialsCitation>
;
<materialsCitation id="EC874B08FFE4FFCC8F30FC2DFD8203C7" ID-GBIF-Occurrence="3311814450" collectionCode="NHMW" country="Austria" location="Forchtenau" pageId="17" pageNumber="18" specimenCode="NHMW 1866" specimenCount="1" typeStatus="paratype">
<specimenCode id="0C49E92EFFE4FFCC8F30FC2DFC5B039C" box="[821,978,898,925]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW 1866</specimenCode>
/0001/1019, SL:
<quantity id="9B17ECB0FFE4FFCC8890FC2CFB79039D" box="[1173,1264,898,925]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="17" pageNumber="18" unit="mm" value="6.8">6.8 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE4FFCC8938FC2CFA11039D" box="[1341,1432,898,925]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="17" pageNumber="18" unit="mm" value="2.7">2.7 mm</quantity>
,
<location id="5930178EFFE4FFCC8C92FC08FE9C03C0" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC8C92FC08FE9C03C0" box="[151,277,934,960]" country="Austria" name="Forchtenau" pageId="17" pageNumber="18">Forchtenau</location>
(
<collectingCountry id="24F801C5FFE4FFCC8D21FC08FEF503C1" box="[292,380,934,961]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
), figs 6F
<subScript id="C06B4310FFE4FFCC8DD8FC17FE6C03C7" attach="left" box="[477,485,953,967]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
F
<subScript id="C06B4310FFE4FFCC8E06FC17FD8203C7" attach="right" box="[515,523,953,967]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
</materialsCitation>
;
<materialsCitation id="EC874B08FFE4FFCC8E1CFC09FE9303EB" ID-GBIF-Occurrence="3311814488" collectionCode="NHMW" country="Austria" location="Steinebrunn" pageId="17" pageNumber="18" specimenCode="NHMW 1871" specimenCount="1" typeStatus="paratype">
<specimenCode id="0C49E92EFFE4FFCC8E1CFC09FD3803C0" box="[537,689,934,961]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW 1871</specimenCode>
/0010/0331, SL: 6.0 mm, MD:
<quantity id="9B17ECB0FFE4FFCC880BFC08FBEF03C1" box="[1038,1126,934,961]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="18" unit="mm" value="2.2">2.2 mm</quantity>
,
<location id="5930178EFFE4FFCC8874FC08FB7103C0" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC8874FC08FB7103C0" box="[1137,1272,934,960]" country="Austria" name="Steinebrunn" pageId="17" pageNumber="18">Steinebrunn</location>
(
<collectingCountry id="24F801C5FFE4FFCC8902FC08FAD603C1" box="[1287,1375,934,961]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
), figs 4C, 6G
<subScript id="C06B4310FFE4FFCC8CEDFC73FF7903EB" attach="left" box="[232,240,989,1003]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
G
<subScript id="C06B4310FFE4FFCC8D17FC73FE9303EB" attach="left" box="[274,282,989,1003]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
</materialsCitation>
;
<materialsCitation id="EC874B08FFE4FFCC8D2CFC65FA8303EB" ID-GBIF-Occurrence="3311814468" box="[297,1290,970,1003]" collectionCode="NHMW" country="Austria" location="Steinebrunn" pageId="17" pageNumber="18" specimenCode="NHMW 1871" specimenCount="1" typeStatus="paratype">
<specimenCode id="0C49E92EFFE4FFCC8D2CFC65FE4803E4" box="[297,449,970,997]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW 1871</specimenCode>
/0011/0004, SL:
<quantity id="9B17ECB0FFE4FFCC8E85FC64FD5103E5" box="[640,728,970,997]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="17" pageNumber="18" unit="mm" value="5.9">5.9 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE4FFCC8F1AFC64FCFE03E5" box="[799,887,970,997]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="17" pageNumber="18" unit="mm" value="2.2">2.2 mm</quantity>
,
<location id="5930178EFFE4FFCC8F86FC64FB8303E4" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC8F86FC64FB8303E4" box="[899,1034,970,996]" country="Austria" name="Steinebrunn" pageId="17" pageNumber="18">Steinebrunn</location>
(
<collectingCountry id="24F801C5FFE4FFCC881CFC64FBF803E5" box="[1049,1137,970,997]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
), figs 6H
<subScript id="C06B4310FFE4FFCC88D2FC73FB5603EB" attach="left" box="[1239,1247,989,1003]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
H
<subScript id="C06B4310FFE4FFCC8907FC73FA8303EB" attach="left" box="[1282,1290,989,1003]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFE4FFCC8CC2FC40FCF5042D" blockId="17.[151,1437,826,2041]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FC40FE320408" bold="true" box="[199,443,1006,1032]" pageId="17" pageNumber="18">Additional material.</emphasis>
<materialsCitation id="EC874B08FFE4FFCC8DC7FC41FEA6042C" ID-GBIF-Occurrence="3311814361" collectionCode="NHMW" country="Austria" location="Steinebrunn" pageId="17" pageNumber="18" specimenCode="NHMW 2020" specimenCount="18">
<specimenCode id="0C49E92EFFE4FFCC8DC7FC41FDD70408" box="[450,606,1006,1033]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW 2020</specimenCode>
/0062/0001, SL:
<quantity id="9B17ECB0FFE4FFCC8F1EFC40FCFD0409" box="[795,884,1006,1033]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="17" pageNumber="18" unit="mm" value="5.7">5.7 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE4FFCC8FB8FC40FB9F0409" box="[957,1046,1006,1033]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="17" pageNumber="18" unit="mm" value="2.1">2.1 mm</quantity>
,
<location id="5930178EFFE4FFCC8827FC40FB200408" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC8827FC40FB200408" box="[1058,1193,1006,1032]" country="Austria" name="Steinebrunn" pageId="17" pageNumber="18">Steinebrunn</location>
(
<collectingCountry id="24F801C5FFE4FFCC88BCFC40FA980409" box="[1209,1297,1006,1033]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
), figs 6I
<subScript id="C06B4310FFE4FFCC896BFBAFFAFF040F" attach="right" box="[1390,1398,1025,1039]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
I
<subScript id="C06B4310FFE4FFCC8988FBAFFA1C040F" attach="left" box="[1421,1429,1025,1039]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
.
<specimenCount id="4AE98ADCFFE4FFCC8C92FBBCFEA6042C" box="[151,303,1042,1068]" pageId="17" pageNumber="18" type="generic">18 specimens</specimenCount>
</materialsCitation>
<materialsCitation id="EC874B08FFE4FFCC8D33FBBDFCF1042D" ID-GBIF-Occurrence="3311814351" box="[310,888,1042,1069]" collectionCode="NHMW" country="Austria" location="Steinebrunn" pageId="17" pageNumber="18" specimenCode="NHMW 2020" specimenCount="1">
<specimenCode id="0C49E92EFFE4FFCC8D33FBBDFE59042C" box="[310,464,1042,1069]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18" type="Museum">NHMW 2020</specimenCode>
/0062/0002, all
<location id="5930178EFFE4FFCC8E84FBBCFC81042C" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC8E84FBBCFC81042C" box="[641,776,1042,1068]" country="Austria" name="Steinebrunn" pageId="17" pageNumber="18">Steinebrunn</location>
(
<collectingCountry id="24F801C5FFE4FFCC8F12FBBCFCF9042D" box="[791,880,1042,1069]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFE4FFCC8CC2FB98FDEE0451" blockId="17.[151,1437,826,2041]" box="[199,615,1078,1105]" pageId="17" pageNumber="18">
<materialsCitation id="EC874B08FFE4FFCC8CC2FB98FDEA0451" ID-GBIF-Occurrence="3311814323" box="[199,611,1078,1105]" country="Austria" location="Steinebrunn" pageId="17" pageNumber="18" specimenCount="1" typeStatus="holotype">
<emphasis id="6E9B9D47FFE4FFCC8CC2FB98FEEF0450" bold="true" box="[199,358,1078,1104]" pageId="17" pageNumber="18">
<typeStatus id="8354FFF7FFE4FFCC8CC2FB98FE8B0450" box="[199,258,1078,1104]" pageId="17" pageNumber="18">Type</typeStatus>
locality.
</emphasis>
<location id="5930178EFFE4FFCC8D69FB98FE7A0450" LSID="urn:lsid:plazi:treatment:D446F043FFE4FFCF8C92FECDFCC502A9:5930178EFFE4FFCC8D69FB98FE7A0450" box="[364,499,1078,1104]" country="Austria" name="Steinebrunn" pageId="17" pageNumber="18">Steinebrunn</location>
(
<collectingCountry id="24F801C5FFE4FFCC8E06FB98FDD50451" box="[515,604,1078,1105]" name="Austria" pageId="17" pageNumber="18">Austria</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFE4FFCC8CC2FBF4FD770474" blockId="17.[151,1437,826,2041]" box="[199,766,1114,1140]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FBF4FEE60474" bold="true" box="[199,367,1114,1140]" pageId="17" pageNumber="18">
<typeStatus id="8354FFF7FFE4FFCC8CC2FBF4FE8B0474" box="[199,258,1114,1140]" pageId="17" pageNumber="18">Type</typeStatus>
stratum.
</emphasis>
Silty sand of the Baden Formation.
</paragraph>
<paragraph id="5C504155FFE4FFCC8CC2FBD0FD490499" blockId="17.[151,1437,826,2041]" box="[199,704,1150,1177]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FBD0FF740498" bold="true" box="[199,253,1150,1176]" pageId="17" pageNumber="18">Age.</emphasis>
Middle Miocene, Badenian (Langhian).
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFE4FFCC8CC2FB0CFC3604E0" pageId="17" pageNumber="18" type="etymology">
<paragraph id="5C504155FFE4FFCC8CC2FB0CFC3604E0" blockId="17.[151,1437,826,2041]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FB0CFEC704BC" bold="true" box="[199,334,1186,1212]" pageId="17" pageNumber="18">Etymology.</emphasis>
In honour of Michal Kováč (Comenius University, Bratislava,
<collectingCountry id="24F801C5FFE4FFCC880EFB0CFBF804BD" box="[1035,1137,1186,1213]" name="Slovakia" pageId="17" pageNumber="18">Slovakia</collectingCountry>
), in recognition of his contributions on the stratigraphy and paleogeography of the Paratethys Sea.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFE4FFCC8CC2FB44FBCD0528" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="5C504155FFE4FFCC8CC2FB44FBCD0528" blockId="17.[151,1437,826,2041]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FB44FECB0504" bold="true" box="[199,322,1258,1284]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Medium-sized, moderately slender fusiform shell differing from other
<taxonomicName id="9BEF3AD6FFE4FFCC886DFB44FADF0504" box="[1128,1366,1258,1284]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="6E9B9D47FFE4FFCC886DFB44FB750504" box="[1128,1276,1258,1284]" italics="true" pageId="17" pageNumber="18">Bellardithala</emphasis>
species
</taxonomicName>
by its weak spiral sculpture and broad flat axial ribs separated by very narrow interspaces.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFE4FFCC8CC2FA9CFDFD0720" pageId="17" pageNumber="18" type="description">
<paragraph id="5C504155FFE4FFCC8CC2FA9CFDBC06D8" blockId="17.[151,1437,826,2041]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2FA9CFED0054C" bold="true" box="[199,345,1330,1356]" pageId="17" pageNumber="18">Description.</emphasis>
Shell medium-sized, moderately slender fusiform with slightly cyrtoconoid spire. Protoconch (fig. 4C) high conical of 3.5 weakly convex whorls; height: 660 μm, diameter: 580 μm. Teleoconch of moderately convex 5.5 whorls with periphery slightly below mid-whorl; suture narrowly incised. Last whorl high, weakly convex with weak basal constriction. Sculpture of first teleoconch whorl of broad, slightly opisthocline axial ribs with flattened tops separated by narrow interspaces. Spiral sculpture reduced to five spiral threads in axial interspaces. Axial ribs becoming broad and low on later spire whorls with irregularly spaced narrow interspaces. Spiral threads restricted to axial interspaces causing tiny pits between axial and spiral elements. Interspaces very narrow, present throughout growth. Spiral grooves somewhat stronger in adapical third of last whorl causing tiny pits at intersections with axial interspaces. Axial sculpture weakening towards fasciole. Aperture elongate, moderately narrow with distinct, narrow anal sinus. Inner lip narrow, indistinct. Columella with four broad folds, adapical two most prominent. Outer lip thickened with 10 denticles on inner side. Siphonal canal moderately short, moderately wide, straight with shallow siphonal notch.
</paragraph>
<paragraph id="5C504155FFE4FFCC8CC2F94CFDFD0720" blockId="17.[151,1437,826,2041]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2F94CFDB506FC" bold="true" box="[199,572,1762,1788]" pageId="17" pageNumber="18">Shell measurements and ratios</emphasis>
. NHMW specimens: SL:
<quantity id="9B17ECB0FFE4FFCC8F6CF94CFC7D06FD" box="[873,1012,1762,1789]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.5" metricValueMin="6.8" pageId="17" pageNumber="18" unit="mm" value="7.65" valueMax="8.5" valueMin="6.8">6.88.5 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE4FFCC8845F94CFB4506FD" box="[1088,1228,1762,1789]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.3" metricValueMin="2.7" pageId="17" pageNumber="18" unit="mm" value="3.0" valueMax="3.3" valueMin="2.7">2.73.3 mm</quantity>
; AA: 4446°, SL/ MD: 2.8, AL/AW: 4.75.3, AH/S: 2.42.6.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFE4FFCF8CC2F884FE0C0219" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph id="5C504155FFE4FFCC8CC2F884FA0C0768" blockId="17.[151,1437,826,2041]" pageId="17" pageNumber="18">
<emphasis id="6E9B9D47FFE4FFCC8CC2F884FEB20744" bold="true" box="[199,315,1834,1860]" pageId="17" pageNumber="18">Remarks.</emphasis>
<taxonomicName id="9BEF3AD6FFE4FFCC8D44F884FDAC0744" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[321,549,1834,1860]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="kovaci">
<emphasis id="6E9B9D47FFE4FFCC8D44F884FDAC0744" box="[321,549,1834,1860]" italics="true" pageId="17" pageNumber="18">Bellardithala kovaci</emphasis>
</taxonomicName>
is documented also by subadult specimens (figs 6G
<subScript id="C06B4310FFE4FFCC8861F893FBE5074B" attach="left" box="[1124,1132,1853,1867]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
G
<subScript id="C06B4310FFE4FFCC888AF893FB1E074B" attach="left" box="[1167,1175,1853,1867]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
, H
<subScript id="C06B4310FFE4FFCC88BDF893FB49074B" attach="left" box="[1208,1216,1853,1867]" fontSize="6" pageId="17" pageNumber="18">1</subScript>
H
<subScript id="C06B4310FFE4FFCC88E6F893FB62074B" attach="left" box="[1251,1259,1853,1867]" fontSize="6" pageId="17" pageNumber="18">2</subScript>
) illustrating the development of denticles in juveniles, which was also observed in
<taxonomicName id="9BEF3AD6FFE4FFCC8F87F8E1FC770768" authorityName="Hornes" authorityYear="1852" box="[898,1022,1870,1896]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="partschi">
<emphasis id="6E9B9D47FFE4FFCC8F87F8E1FC770768" box="[898,1022,1870,1896]" italics="true" pageId="17" pageNumber="18">B. partschi</emphasis>
</taxonomicName>
by
<bibRefCitation id="387E3CA4FFE4FFCC8822F8E0FA080768" author="Rosenberg, G. &amp; Salisbury, R." box="[1063,1409,1870,1896]" pageId="17" pageNumber="18" pagination="1 - 30" refId="ref61404" refString="Rosenberg, G. &amp; Salisbury, R. (2003) On Mitromica and Thala (Gastropoda: Costellariidae) with descriptions of new species from the Western Atlantic and the Indo-Pacific. Notulae Naturae, 478, 1 - 30." type="journal article" year="2003">Rosenberg &amp; Salisbury (2003)</bibRefCitation>
.
</paragraph>
<paragraph id="5C504155FFE4FFCF8CC2F8DCFE0E0189" blockId="17.[151,1437,826,2041]" lastBlockId="18.[151,1437,151,682]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
This species was misidentified so far as
<taxonomicName id="9BEF3AD6FFE4FFCC8E81F8DCFBBF078C" authority="(Brocchi, 1814)" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[644,1078,1906,1932]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8E81F8DCFCF7078C" box="[644,894,1906,1932]" italics="true" pageId="17" pageNumber="18">Bellardithala obsoleta</emphasis>
(
<bibRefCitation id="387E3CA4FFE4FFCC8F8EF8DCFBA6078C" author="Brocchi, G." box="[907,1071,1906,1932]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
)
</taxonomicName>
, which was originally described from the Italian Pliocene. The
<typeStatus id="8354FFF7FFE4FFCC8DE8F838FDC607B0" box="[493,591,1942,1968]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
of
<taxonomicName id="9BEF3AD6FFE4FFCC8E71F838FC4307B0" authority="Brocchi, 1814" authorityName="Brocchi" authorityYear="1814" box="[628,970,1942,1968]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8E71F838FCAB07B0" box="[628,802,1942,1968]" italics="true" pageId="17" pageNumber="18">Voluta obsoleta</emphasis>
<bibRefCitation id="387E3CA4FFE4FFCC8F2CF838FC4307B0" author="Brocchi, G." box="[809,970,1942,1968]" pageId="17" pageNumber="18" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
</taxonomicName>
, illustrated in Rossi Ronchetti (1955) and
<bibRefCitation id="387E3CA4FFE4FFCC8C92F814FE1407D4" author="Pinna G. &amp; Spezia L." box="[151,413,1978,2004]" pageId="17" pageNumber="18" pagination="125 - 180" refId="ref60642" refString="Pinna G. &amp; Spezia L. (1978) Catalogo dei tipi del Museo Civico di Storia Naturale di Milano. V. I tipi dei gasteropodi fossili. Atti della Societa Italiana di Scienze Naturali, 119, 125 - 180." type="journal article" year="1978">Pinna &amp; Spezia (1978)</bibRefCitation>
, differs clearly from the Paratethyan species in having prominent axial sculpture, on spire whorls and last whorl. Pliocene specimens of
<taxonomicName id="9BEF3AD6FFE4FFCC8E98F871FC9507F8" authorityName="Brocchi" authorityYear="1814" box="[669,796,2014,2040]" class="Gastropoda" family="Volutidae" genus="Voluta" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE4FFCC8E98F871FC9507F8" box="[669,796,2014,2040]" italics="true" pageId="17" pageNumber="18">B. obsoleta</emphasis>
</taxonomicName>
from Altavilla (
<collectingCountry id="24F801C5FFE4FFCC8FD7F870FB8307F9" box="[978,1034,2014,2041]" name="Italy" pageId="17" pageNumber="18">Italy</collectingCountry>
), described by
<bibRefCitation id="387E3CA4FFE4FFCF88B0F870FEA700B1" author="Giannuzzi-Savelli, R. &amp; Reina, M." lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" pagination="227 - 236" refId="ref56518" refString="Giannuzzi-Savelli, R. &amp; Reina, M. (1983) Thala obsoleta (Brocchi, 1814) nel Pliocene di Altavilla ed alcune considerazioni evolutive e paleoecologiche sul genere. Bollettino Malacologico, 19, 227 - 236. [https: // www. biodiversitylibrary. org / item / 199216 # page / 163 / mode / 1 up]" type="journal article" year="1983">Giannuzzi-Savelli &amp; Reina (1983)</bibRefCitation>
, differ from the Paratethyan species also in the more prominent axial and spiral sculpture and have a more slender fusiform outline. Serravallian specimens from the
<collectingRegion id="9E2B8FB7FFE7FFCF8F85FF15FC6E00D5" box="[896,999,187,213]" country="Turkey" name="Karaman" pageId="18" pageNumber="19">Karaman</collectingRegion>
Basin in
<collectingCountry id="24F801C5FFE7FFCF8851FF15FB2C00D5" box="[1108,1189,187,213]" name="Turkey" pageId="18" pageNumber="19">Turkey</collectingCountry>
, described by
<bibRefCitation id="387E3CA4FFE7FFCF894DFF15FEA100F9" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." pageId="18" pageNumber="19" pagination="3 - 576" refId="ref58833" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576." type="journal article" year="2013">
Landau
<emphasis id="6E9B9D47FFE7FFCF8C92FF71FF4500F9" box="[151,204,223,249]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (2013)
</bibRefCitation>
as
<taxonomicName id="9BEF3AD6FFE7FFCF8D51FF71FE7600F9" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[340,511,223,249]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="obsoleta">
<emphasis id="6E9B9D47FFE7FFCF8D51FF71FE7600F9" box="[340,511,223,249]" italics="true" pageId="18" pageNumber="19">Thala obsoleta</emphasis>
</taxonomicName>
, differ from the Paratethyan species clearly in the prominent axial ribs and the subcylindrical spire whorls. The same features distinguish Tortonian specimens from Montegibbio in
<collectingCountry id="24F801C5FFE7FFCF892CFEADFAD5011D" box="[1321,1372,259,285]" name="Italy" pageId="18" pageNumber="19">Italy</collectingCountry>
from
<taxonomicName id="9BEF3AD6FFE7FFCF8C92FE89FEF50141" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[151,380,295,321]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="kovaci">
<emphasis id="6E9B9D47FFE7FFCF8C92FE89FEF50141" box="[151,380,295,321]" italics="true" pageId="18" pageNumber="19">Bellardithala kovaci</emphasis>
</taxonomicName>
(see
<bibRefCitation id="387E3CA4FFE7FFCF8DB2FE89FDC70141" author="Davoli, F." box="[439,590,295,321]" pageId="18" pageNumber="19" pagination="165 - 215" refId="ref55395" refString="Davoli, F. (2000) I gasteropodi mitriformi del Tortoniano di Montegibbio (Subappennino modenese). Bolletino della Societa Paleontologica Italiana, 39 / 2, 165 - 215." type="journal article" year="2000">Davoli, 2000</bibRefCitation>
).
<taxonomicName id="9BEF3AD6FFE7FFCF8E64FE89FB870142" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[609,1038,295,322]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="boehmi">
<emphasis id="6E9B9D47FFE7FFCF8E64FE89FCC70141" box="[609,846,295,321]" italics="true" pageId="18" pageNumber="19">Bellardithala boehmi</emphasis>
(
<bibRefCitation id="387E3CA4FFE7FFCF8F58FE89FB8F0142" author="Boettger, O." box="[861,1030,295,322]" pageId="18" pageNumber="19" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
differs in its more slender fusiform outline, much more prominent axial and spiral sculpture on the early teleoconch whorls, and the lower aperture/spire ratio (0.9 versus 1.1).
</paragraph>
<paragraph id="5C504155FFE7FFCF8CC2FE3CFE0C0219" blockId="18.[151,1437,151,682]" pageId="18" pageNumber="19">
<emphasis id="6E9B9D47FFE7FFCF8CC2FE3CFE3F01AC" bold="true" box="[199,438,402,428]" pageId="18" pageNumber="19">Palaeoenvironment.</emphasis>
Coastal, inner neritic. The assemblage from Steinebrunn suggests shallow sublittoral environments with sea grass (
<bibRefCitation id="387E3CA4FFE7FFCF8DC6FE19FD7401D1" author="Harzhauser, M. &amp; Landau, B. M." box="[451,765,439,465]" pageId="18" pageNumber="19" pagination="1 - 136" refId="ref57199" refString="Harzhauser, M. &amp; Landau, B. M. (2019) Turritellidae (Gastropoda) of the Miocene Paratethys Sea with considerations about turritellid genera. Zootaxa, 4681 (1), 1 - 136. https: // doi. org / 10.11646 / zootaxa. 4681.1.1" type="journal article" year="2019">Harzhauser &amp; Landau 2019</bibRefCitation>
).At the Niederleis section, sediment and fossils from coastal and lagoonal environments not exceeding
<quantity id="9B17ECB0FFE7FFCF8E71FE75FD2701F5" box="[628,686,475,501]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="m" value="30.0">30 m</quantity>
water depth were transported by tempestites into offshore settings (
<bibRefCitation id="387E3CA4FFE7FFCF8C9AFE51FEF00219" author="Mandic, O. &amp; Harzhauser, M. &amp; Spezzaferri, S. &amp; Zuschin, M." box="[159,377,511,537]" pageId="18" pageNumber="19" pagination="193 - 206" refId="ref59240" refString="Mandic, O., Harzhauser, M., Spezzaferri, S., Zuschin, M. (2002) The Palaeoenvironment of an early Middle Miocene Paratethys sequence in NE Austria with special emphasis on Palaeoecology of mollusks and foraminifera. Geobios, Memoirs Special, 24, 193 - 206. https: // doi. org / 10.1016 / S 0016 - 6995 (02) 00059 - 1" type="journal article" year="2002">
Mandic
<emphasis id="6E9B9D47FFE7FFCF8CF8FE51FEA70219" box="[253,302,511,537]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. 2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFE7FFCF8CC2FD8CFCC502A9" pageId="18" pageNumber="19" type="distribution">
<paragraph id="5C504155FFE7FFCF8CC2FD8CFCC502A9" blockId="18.[151,1437,151,682]" pageId="18" pageNumber="19">
<emphasis id="6E9B9D47FFE7FFCF8CC2FD8CFDE4023D" bold="true" box="[199,621,546,573]" pageId="18" pageNumber="19">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFE7FFCF8FC4FD8DFA1C023C" box="[961,1429,547,573]" italics="true" pageId="18" pageNumber="19">North Alpine-Carpathian Foreland Basin</emphasis>
: Unkovice (
<collectingCountry id="24F801C5FFE7FFCF8D13FDE9FE590262" box="[278,464,583,610]" name="Czech Republic" pageId="18" pageNumber="19">Czech Republic</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFE7FFCF8DEFFDE9FD0F0261" author="Rzehak, A." box="[490,646,583,609]" pageId="18" pageNumber="19" pagination="252 - 262" refId="ref61539" refString="Rzehak, A. (1894) Uber einige neue Fossilienfundorte im mahrischen Miocan. Verhandlungen des naturforschenden Vereines in Brunn, 33, 252 - 262. [http: // www. zobodat. at / pdf / Verh-naturf-Ver-Bruenn _ 33 _ 0252 - 0262. pdf]" type="journal article" year="1894">Rzehak 1894</bibRefCitation>
);
<emphasis id="6E9B9D47FFE7FFCF8E9BFDE9FCD00260" box="[670,857,583,608]" italics="true" pageId="18" pageNumber="19">Korytnica Basin</emphasis>
: Korytnica (
<collectingCountry id="24F801C5FFE7FFCF8FEBFDE9FBB60262" box="[1006,1087,583,610]" name="Poland" pageId="18" pageNumber="19">Poland</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFE7FFCF885CFDE9FB6C0261" author="Baluk, W." box="[1113,1253,583,609]" pageId="18" pageNumber="19" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk 1997</bibRefCitation>
);
<emphasis id="6E9B9D47FFE7FFCF88F8FDE9FA1C0260" box="[1277,1429,583,608]" italics="true" pageId="18" pageNumber="19">
<collectingRegion id="9E2B8FB7FFE7FFCF88F8FDE9FAC30260" box="[1277,1354,583,608]" country="Austria" name="Wien" pageId="18" pageNumber="19">Vienna</collectingRegion>
Basin
</emphasis>
: Niederleis, Steinebrunn (
<collectingCountry id="24F801C5FFE7FFCF8DB0FDC5FD870286" box="[437,526,619,646]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
); Mikulov, Sedlec (= Porzteich) (
<collectingCountry id="24F801C5FFE7FFCF8F97FDC5FBC20286" box="[914,1099,619,646]" name="Czech Republic" pageId="18" pageNumber="19">Czech Republic</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFE7FFCF8860FDC5FA060285" author="Hoernes, R. &amp; Auinger, M." box="[1125,1423,619,645]" pageId="18" pageNumber="19" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger 1880</bibRefCitation>
);
<emphasis id="6E9B9D47FFE7FFCF8C92FD21FE2502A8" box="[151,428,655,681]" italics="true" pageId="18" pageNumber="19">Eisenstadt-Sopron Basin</emphasis>
: Forchtenau (
<collectingCountry id="24F801C5FFE7FFCF8E42FD21FD1402AA" box="[583,669,655,682]" name="Austria" pageId="18" pageNumber="19">Austria</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFE7FFCF8EB6FD21FCC902A9" author="Sieber, R." box="[691,832,654,681]" pageId="18" pageNumber="19" pagination="236 - 249" refId="ref61969" refString="Sieber, R. (1956) Die Tortonfauna von Mattersburg und Forchtenau (Burgenland). Verhandlungen der Geologischen Bunsdesanstalt, 1956, 236 - 249. [https: // opac. geologie. ac. at / ais 312 / dokumente / VH 1956 _ 236 _ A. pdf]" type="journal article" year="1956">Sieber 1956</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>