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<mods:title id="90A9C12F49980432FD9C8A374845A54B">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<treatment id="D446F043FFE2FFC58C92FAB3FECF02F1" ID-DOI="http://doi.org/10.5281/zenodo.5044016" ID-GBIF-Taxon="183225136" ID-Zenodo-Dep="5044016" LSID="urn:lsid:plazi:treatment:D446F043FFE2FFC58C92FAB3FECF02F1" httpUri="http://treatment.plazi.org/id/D446F043FFE2FFC58C92FAB3FECF02F1" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<emphasis id="6E9B9D47FFE2FFCA8C92FAB3FC850538" bold="true" box="[151,780,1309,1336]" pageId="23" pageNumber="24">
<taxonomicName id="9BEF3AD6FFE2FFCA8C92FAB3FDF60537" ID-CoL="8KT36" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[151,639,1309,1336]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="kostejana" status="comb. nov.">
<emphasis id="6E9B9D47FFE2FFCA8C92FAB3FE3B0538" bold="true" box="[151,434,1309,1336]" italics="true" pageId="23" pageNumber="24">Bellardithala kostejana</emphasis>
(
<bibRefCitation id="387E3CA4FFE2FFCA8DC4FAB3FDFE0537" author="Boettger, O." box="[449,631,1309,1335]" pageId="23" pageNumber="24" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
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<taxonomicNameLabel id="75A8203CFFE2FFCA8E83FAB0FC850538" box="[646,780,1310,1336]" pageId="23" pageNumber="24" rank="species">nov. comb.</taxonomicNameLabel>
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<subSubSection id="14F512DEFFE2FFCA8C92FAECFE04055C" box="[151,397,1346,1378]" pageId="23" pageNumber="24" type="description">
<paragraph id="5C504155FFE2FFCA8C92FAECFE04055C" blockId="23.[151,780,1309,1378]" box="[151,397,1346,1378]" pageId="23" pageNumber="24">
<figureCitation id="C4D45DD0FFE2FFCA8C92FAECFF78055C" box="[151,241,1346,1372]" captionStart="FIGURE 8" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[151,1436,669,1777]" captionTargetId="figure-225@22.[151,1436,669,1777]" captionTargetPageId="22" captionText="FIGURE 8 A1–A2. Bellardithala dacica nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0105. B1–B2. Bellardithala dacica nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0061/0001. C1–C2. Bellardithala dacica nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0061/0002. D1–D2. Bellardithala kostejana (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360340. E1–E2. Bellardithala kostejana (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0064/0001. F. Bellardithala kostejana (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0064/0002. G. Bellardithala sp., Devínska Nová Ves (Slovakia), Z23408-P2003/27." figureDoi="http://doi.org/10.5281/zenodo.4921858" httpUri="https://zenodo.org/record/4921858/files/figure.png" pageId="23" pageNumber="24">Figs 8D</figureCitation>
<figureCitation id="C4D45DD0FFE2FFCA8CF4FAFAFE92055C" box="[241,283,1346,1378]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="23" pageNumber="24">
<subScript id="C06B4310FFE2FFCA8CF4FAFAFF700562" attach="left" box="[241,249,1364,1378]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
–D
</figureCitation>
<figureCitation id="C4D45DD0FFE2FFCA8D1EFAFAFEAA0562" box="[283,291,1364,1378]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="23" pageNumber="24">
<subScript id="C06B4310FFE2FFCA8D1EFAFAFEAA0562" attach="left" box="[283,291,1364,1378]" fontSize="6" pageId="23" pageNumber="24">2</subScript>
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, E
<figureCitation id="C4D45DD0FFE2FFCA8D47FAFAFEE3055C" box="[322,362,1346,1378]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="23" pageNumber="24">
<subScript id="C06B4310FFE2FFCA8D47FAFAFEC30562" attach="left" box="[322,330,1364,1378]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
–E
</figureCitation>
<figureCitation id="C4D45DD0FFE2FFCA8D6FFAFAFEFB0562" box="[362,370,1364,1378]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="23" pageNumber="24">
<subScript id="C06B4310FFE2FFCA8D6FFAFAFEFB0562" attach="left" box="[362,370,1364,1378]" fontSize="6" pageId="23" pageNumber="24">2</subScript>
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, F
</paragraph>
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<paragraph id="5C504155FFE2FFCA8C92FA29FCE8059E" blockId="23.[151,937,1414,1500]" box="[151,865,1414,1439]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="7CFF667BFFE2FFCA8C92FA29FCE8059E" box="[151,865,1414,1439]" pageId="23" pageNumber="24">
<emphasis id="6E9B9D47FFE2FFCA8C92FA29FF2D059E" box="[151,164,1415,1438]" italics="true" pageId="23" pageNumber="24">*</emphasis>
<taxonomicName id="9BEF3AD6FFE2FFCA8CAFFA29FDF8059E" box="[170,625,1414,1438]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="variety" species="partschi" subGenus="Thala" variety="kostejana">
<emphasis id="6E9B9D47FFE2FFCA8CAFFA29FF6D059E" box="[170,228,1415,1438]" italics="true" pageId="23" pageNumber="24">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFE2FFCA8CF6FA28FEA7059E" box="[243,302,1414,1438]" italics="true" pageId="23" pageNumber="24">Thala</emphasis>
)
<emphasis id="6E9B9D47FFE2FFCA8D38FA28FE19059E" box="[317,400,1414,1438]" italics="true" pageId="23" pageNumber="24">partschi</emphasis>
M. Hö. var.
<emphasis id="6E9B9D47FFE2FFCA8E15FA28FDF8059E" box="[528,625,1414,1438]" italics="true" pageId="23" pageNumber="24">kostejana</emphasis>
</taxonomicName>
n.—
<treatmentCitation id="DD4E6744FFE2FFCA8EA7FA29FCD4059E" author="Boettger, O." box="[674,861,1414,1439]" page="11" pageId="23" pageNumber="24" year="1906">
<bibRefCitation id="387E3CA4FFE2FFCA8EA7FA29FCD4059E" author="Boettger, O." box="[674,861,1414,1439]" pageId="23" pageNumber="24" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger 1906: 11</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="5C504155FFE2FFCA8C92FA08FC2005BD" blockId="23.[151,937,1414,1500]" box="[151,937,1445,1469]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="7CFF667BFFE2FFCA8C92FA08FC2005BD" box="[151,937,1445,1469]" pageId="23" pageNumber="24">
<taxonomicName id="9BEF3AD6FFE2FFCA8C92FA08FDD805BD" authority="(Boettger)" baseAuthorityName="Boettger" box="[151,593,1445,1469]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="subSpecies" species="partschi" subGenus="Thala" subSpecies="kostejana">
<emphasis id="6E9B9D47FFE2FFCA8C92FA08FF5805BD" box="[151,209,1446,1469]" italics="true" pageId="23" pageNumber="24">Pusia</emphasis>
(
<emphasis id="6E9B9D47FFE2FFCA8CE5FA0BFE9205BD" box="[224,283,1445,1469]" italics="true" pageId="23" pageNumber="24">Thala</emphasis>
)
<emphasis id="6E9B9D47FFE2FFCA8D2FFA0BFE6D05BD" box="[298,484,1445,1469]" italics="true" pageId="23" pageNumber="24">partschi kostejana</emphasis>
(Boettger)
</taxonomicName>
—
<treatmentCitation id="DD4E6744FFE2FFCA8E6EFA0BFC9905BD" author="Zilch, A." box="[619,784,1445,1469]" page="285" pageId="23" pageNumber="24" year="1934">
<bibRefCitation id="387E3CA4FFE2FFCA8E6EFA0BFC9905BD" author="Zilch, A." box="[619,784,1445,1469]" pageId="23" pageNumber="24" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch 1934: 285</bibRefCitation>
</treatmentCitation>
, pl. 17, fig. 13.
</treatmentCitationGroup>
</paragraph>
<paragraph id="5C504155FFE2FFCA8C92FA6AFC1705DC" blockId="23.[151,937,1414,1500]" box="[151,926,1475,1500]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="7CFF667BFFE2FFCA8C92FA6AFC1705DC" box="[151,926,1475,1500]" pageId="23" pageNumber="24">
<taxonomicName id="9BEF3AD6FFE2FFCA8C92FA6AFDB405DB" authority="(Boettger, 1906)" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[151,573,1475,1500]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="subSpecies" species="partschi" subSpecies="kostejana">
<emphasis id="6E9B9D47FFE2FFCA8C92FA6AFE1A05DC" box="[151,403,1476,1500]" italics="true" pageId="23" pageNumber="24">Thala partschi kostejana</emphasis>
(
<bibRefCitation id="387E3CA4FFE2FFCA8DA5FA6AFDBF05DB" author="Boettger, O." box="[416,566,1475,1500]" pageId="23" pageNumber="24" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
—
<treatmentCitation id="DD4E6744FFE2FFCA8E52FA6DFC1205DC" author="Rosenberg, G. &amp; Salisbury, R." box="[599,923,1475,1500]" page="3" pageId="23" pageNumber="24" year="2003">
<bibRefCitation id="387E3CA4FFE2FFCA8E52FA6DFC1205DC" author="Rosenberg, G. &amp; Salisbury, R." box="[599,923,1475,1500]" pageId="23" pageNumber="24" pagination="1 - 30" refId="ref61404" refString="Rosenberg, G. &amp; Salisbury, R. (2003) On Mitromica and Thala (Gastropoda: Costellariidae) with descriptions of new species from the Western Atlantic and the Indo-Pacific. Notulae Naturae, 478, 1 - 30." type="journal article" year="2003">Rosenberg &amp; Salisbury 2003: 3</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
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<paragraph id="5C504155FFE2FFCA8C92F9A4FEEE0691" blockId="23.[151,1437,1545,2040]" pageId="23" pageNumber="24">
<emphasis id="6E9B9D47FFE2FFCA8C92F9A4FECC0623" bold="true" box="[151,325,1545,1572]" pageId="23" pageNumber="24">Type material.</emphasis>
<materialsCitation id="EC874B08FFE2FFCA8D49F9A4FF650672" ID-GBIF-Occurrence="3311814368" collectingDate="2214-12" collectionCode="SMF" collectorName="de Sus" country="Germany" location="Costeiu de Sus" pageId="23" pageNumber="24" specimenCode="SMF 360340" specimenCount="1" typeStatus="lectotype">
<typeStatus id="8354FFF7FFE2FFCA8D49F9A4FE490624" box="[332,448,1546,1572]" pageId="23" pageNumber="24" type="lectotype">Lectotype</typeStatus>
:
<specimenCode id="0C49E92EFFE2FFCA8DCFF9A4FDD40624" box="[458,605,1546,1572]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="24">SMF 360340</specimenCode>
(=
<collectionCode id="3AFED990FFE2FFCA8E81F9A4FD350624" box="[644,700,1546,1572]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="24">SMF</collectionCode>
<date id="28516795FFE2FFCA8EC6F9A4FCA50624" box="[707,812,1546,1572]" pageId="23" pageNumber="24" value="2214-12">
<collectingDate id="38159E7DFFE2FFCA8EC6F9A4FCA50624" box="[707,812,1546,1572]" pageId="23" pageNumber="24" value="2214-12">XII.2214</collectingDate>
</date>
a), SL:
<quantity id="9B17ECB0FFE2FFCA8F79F9A4FC5D0624" box="[892,980,1546,1573]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="23" pageNumber="24" unit="mm" value="5.7">5.7 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE2FFCA881EF9A4FBFA0624" box="[1051,1139,1546,1572]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="23" pageNumber="24" unit="mm" value="2.1">2.1 mm</quantity>
, designated and illustrated by
<bibRefCitation id="387E3CA4FFE2FFCA8CBEF980FECB0648" author="Zilch, A." box="[187,322,1582,1608]" pageId="23" pageNumber="24" pagination="193 - 302" refId="ref63342" refString="Zilch, A. (1934) Zur Fauna des Mittel-Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. Boettger's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch (1934</bibRefCitation>
, pl. 17, fig. 13), Senckenberg Museum, Frankfurt/Main,
<collectingCountry id="24F801C5FFE2FFCA8FC3F980FBA60648" box="[966,1071,1582,1608]" name="Germany" pageId="23" pageNumber="24">Germany</collectingCountry>
,
<location id="5930178EFFE2FFCA883EF980FB6A0648" LSID="urn:lsid:plazi:treatment:D446F043FFE2FFC58C92FAB3FECF02F1:5930178EFFE2FFCA883EF980FB6A0648" box="[1083,1251,1582,1608]" country="Germany" name="Costeiu de Sus" pageId="23" pageNumber="24">
Coşteiu
<collectorName id="F11A2483FFE2FFCA889CF980FB6A0648" box="[1177,1251,1582,1608]" pageId="23" pageNumber="24">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFE2FFCA88F6F980FAD70649" box="[1267,1374,1582,1609]" name="Romania" pageId="23" pageNumber="24">Romania</collectingCountry>
), figs 8D
<subScript id="C06B4310FFE2FFCA8CBCF9CAFF480672" attach="left" box="[185,193,1636,1650]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
–D
<subScript id="C06B4310FFE2FFCA8CE1F9CAFF650672" attach="left" box="[228,236,1636,1650]" fontSize="6" pageId="23" pageNumber="24">2</subScript>
</materialsCitation>
.
<materialsCitation id="EC874B08FFE2FFCA8CFFF9FCFAB2066C" ID-GBIF-Occurrence="3311814358" box="[250,1339,1618,1644]" collectingDate="2214-12" collectionCode="SMF" country="Germany" county="Main" location="Senckenberg Museum" municipality="Frankfurt" pageId="23" pageNumber="24" specimenCount="28" typeStatus="paralectotype">
<typeStatus id="8354FFF7FFE2FFCA8CFFF9FCFE2B066C" box="[250,418,1618,1644]" pageId="23" pageNumber="24" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="4AE98ADCFFE2FFCA8DA9F9FCFDCF066C" box="[428,582,1618,1644]" pageId="23" pageNumber="24" type="generic">28 specimens</specimenCount>
,
<collectionCode id="3AFED990FFE2FFCA8E54F9FCFD00066C" box="[593,649,1618,1644]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="23" pageNumber="24">SMF</collectionCode>
<date id="28516795FFE2FFCA8E95F9FCFD71066C" box="[656,760,1618,1644]" pageId="23" pageNumber="24" value="2214-12">
<collectingDate id="38159E7DFFE2FFCA8E95F9FCFD71066C" box="[656,760,1618,1644]" pageId="23" pageNumber="24" value="2214-12">XII 2214</collectingDate>
</date>
b,
<location id="5930178EFFE2FFCA8F14F9FCFB84066C" LSID="urn:lsid:plazi:treatment:D446F043FFE2FFC58C92FAB3FECF02F1:5930178EFFE2FFCA8F14F9FCFB84066C" box="[785,1037,1618,1644]" country="Germany" county="Main" municipality="Frankfurt" name="Senckenberg Museum" pageId="23" pageNumber="24">Senckenberg Museum</location>
,
<collectingMunicipality id="BC34DB2FFFE2FFCA881DF9FCFB0C066C" box="[1048,1157,1618,1644]" pageId="23" pageNumber="24">Frankfurt</collectingMunicipality>
/
<collectingCounty id="B53139D9FFE2FFCA8888F9FCFB41066C" box="[1165,1224,1618,1644]" pageId="23" pageNumber="24">Main</collectingCounty>
,
<collectingCountry id="24F801C5FFE2FFCA88D7F9FCFAB2066C" box="[1234,1339,1618,1644]" name="Germany" pageId="23" pageNumber="24">Germany</collectingCountry>
</materialsCitation>
,
<materialsCitation id="EC874B08FFE2FFCA8942F9FCFEED0691" ID-GBIF-Occurrence="3311814379" collectorName="de Sus" country="Romania" location="Costeiu de Sus" pageId="23" pageNumber="24" specimenCount="1" typeStatus="paralectotype">
<location id="5930178EFFE2FFCA8942F9FCFF680690" LSID="urn:lsid:plazi:treatment:D446F043FFE2FFC58C92FAB3FECF02F1:5930178EFFE2FFCA8942F9FCFF680690" country="Romania" name="Costeiu de Sus" pageId="23" pageNumber="24">
Coşteiu
<collectorName id="F11A2483FFE2FFCA8C92F9D8FF680690" box="[151,225,1654,1680]" pageId="23" pageNumber="24">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFE2FFCA8CF5F9D8FED50691" box="[240,348,1654,1681]" name="Romania" pageId="23" pageNumber="24">Romania</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5C504155FFE2FFCA8CC2F937FB5006D8" blockId="23.[151,1437,1545,2040]" pageId="23" pageNumber="24">
<emphasis id="6E9B9D47FFE2FFCA8CC2F937FE3506B3" bold="true" box="[199,444,1689,1715]" pageId="23" pageNumber="24">Additional material.</emphasis>
<materialsCitation id="EC874B08FFE2FFCA8DC0F934FF6F06DE" ID-GBIF-Occurrence="3311814449" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="23" pageNumber="24" specimenCode="NHMW 2020" specimenCount="1">
<specimenCode id="0C49E92EFFE2FFCA8DC0F934FDEB06B4" box="[453,610,1690,1716]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24" type="Museum">NHMW 2020</specimenCode>
/0064/0001, SL:
<quantity id="9B17ECB0FFE2FFCA8F27F934FCF506B4" box="[802,892,1690,1717]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="23" pageNumber="24" unit="mm" value="5.8">5.8 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE2FFCA8FCDF934FBAB06B4" box="[968,1058,1690,1716]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="24" unit="mm" value="2.3">2.3 mm</quantity>
,
<location id="5930178EFFE2FFCA882AF934FB6806B4" LSID="urn:lsid:plazi:treatment:D446F043FFE2FFC58C92FAB3FECF02F1:5930178EFFE2FFCA882AF934FB6806B4" box="[1071,1249,1690,1716]" country="Romania" name="Lapugiu de Sus" pageId="23" pageNumber="24">
Lăpugiu
<collectorName id="F11A2483FFE2FFCA8893F934FB6806B4" box="[1174,1249,1690,1716]" pageId="23" pageNumber="24">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFE2FFCA88F7F934FAD406B5" box="[1266,1373,1690,1717]" name="Romania" pageId="23" pageNumber="24">Romania</collectingCountry>
), figs 8E
<subScript id="C06B4310FFE2FFCA8CB3F97EFF3706DE" attach="left" box="[182,190,1744,1758]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
–E
<subScript id="C06B4310FFE2FFCA8CDBF97EFF6F06DE" attach="left" box="[222,230,1744,1758]" fontSize="6" pageId="23" pageNumber="24">2</subScript>
</materialsCitation>
;
<materialsCitation id="EC874B08FFE2FFCA8CF1F910FB5006D8" ID-GBIF-Occurrence="3311814440" box="[244,1241,1726,1753]" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="23" pageNumber="24" specimenCode="NHMW 2020" specimenCount="8">
<specimenCode id="0C49E92EFFE2FFCA8CF1F910FE0606D8" box="[244,399,1726,1752]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24" type="Museum">NHMW 2020</specimenCode>
/0064/0002, SL:
<quantity id="9B17ECB0FFE2FFCA8E49F910FD2D06D8" box="[588,676,1726,1753]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="23" pageNumber="24" unit="mm" value="5.5">5.5 mm</quantity>
, MD:
<quantity id="9B17ECB0FFE2FFCA8EEEF910FCCA06D8" box="[747,835,1726,1752]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="24" unit="mm" value="2.3">2.3 mm</quantity>
,
<location id="5930178EFFE2FFCA8F4AF910FC7406D8" LSID="urn:lsid:plazi:treatment:D446F043FFE2FFC58C92FAB3FECF02F1:5930178EFFE2FFCA8F4AF910FC7406D8" box="[847,1021,1726,1752]" country="Romania" name="Lapugiu de Sus" pageId="23" pageNumber="24">
Lăpugiu
<collectorName id="F11A2483FFE2FFCA8FB6F910FC7406D8" box="[947,1021,1726,1752]" pageId="23" pageNumber="24">de Sus</collectorName>
</location>
(
<collectingCountry id="24F801C5FFE2FFCA8809F910FBFE06D9" box="[1036,1143,1726,1753]" name="Romania" pageId="23" pageNumber="24">Romania</collectingCountry>
), fig.
<specimenCount id="4AE98ADCFFE2FFCA88B2F910FB5006D8" box="[1207,1241,1726,1752]" pageId="23" pageNumber="24" type="generic">8F.</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="5C504155FFE2FFCA8CC2F94FFAD807F8" blockId="23.[151,1437,1545,2040]" pageId="23" pageNumber="24">
<emphasis id="6E9B9D47FFE2FFCA8CC2F94FFE3306FB" bold="true" box="[199,442,1761,1787]" pageId="23" pageNumber="24">Revised description.</emphasis>
Shell small, moderately broad fusiform with conical to slightly cyrtoconoid spire. Protoconch poorly preserved, high conical with three regularly convex whorls. Teleoconch of 4.5 weakly convex to nearly straight-sided whorls with periphery at abapical suture, separated by incised suture.Axial sculpture of densely spaced, orthocline axial ribs, crossed by densely spaced spiral cords, forming delicate, spirally elongated pits in the interspaces. Spiral sculpture most prominent in axial interspaces, but strongly reduced on axial ribs. Last whorl moderately constricted at base. Aperture moderately narrow, with indistinct anal sinus. Columellar callus narrow. Columella with four columellar folds, weakening rapidly abapically. Outer lip weakly thickened, bearing about nine prominent denticles within. Siphonal canal moderately short, bent slightly left, with shallow siphonal notch.
</paragraph>
<paragraph id="5C504155FFEDFFC58CC2FF38FE2B00D4" blockId="24.[151,1437,150,753]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58CC2FF38FDBC00B0" bold="true" box="[199,565,150,176]" pageId="24" pageNumber="25">Shell measurements and ratios</emphasis>
. SL:
<quantity id="9B17ECB0FFEDFFC58E71FF38FD7400B1" box="[628,765,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="5.8" metricValueMin="5.5" pageId="24" pageNumber="25" unit="mm" value="5.65" valueMax="5.8" valueMin="5.5">5.5–5.8 mm</quantity>
, MD: 2.1–2.3, mm, AA: 26–43°, SL/MD: 2.6–2.8, AL/AW: 3.4–5.0, AH/S: 2.4–2.6.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFEDFFC58CC2FF70FE1602A9" pageId="24" pageNumber="25" type="discussion">
<paragraph id="5C504155FFEDFFC58CC2FF70FE1601D1" blockId="24.[151,1437,150,753]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58CC2FF70FEC500F8" bold="true" box="[199,332,222,248]" pageId="24" pageNumber="25">Discussion.</emphasis>
<taxonomicName id="9BEF3AD6FFEDFFC58D53FF70FCAA00F9" authority="(Boettger, 1906)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[342,803,222,249]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="kostejana">
<emphasis id="6E9B9D47FFEDFFC58D53FF70FDD600F8" box="[342,607,222,248]" italics="true" pageId="24" pageNumber="25">Bellardithala kostejana</emphasis>
(
<bibRefCitation id="387E3CA4FFEDFFC58E75FF70FC9200F9" author="Boettger, O." box="[624,795,222,249]" pageId="24" pageNumber="25" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger, 1906</bibRefCitation>
)
</taxonomicName>
can be separated from all its Paratethyan congeners in having a strongly oblique suture. It is superficially similar to
<taxonomicName id="9BEF3AD6FFEDFFC58F3AFEADFBE5011D" authority="(Hornes, 1852)" baseAuthorityName="Hoernes" baseAuthorityYear="1852" box="[831,1132,258,285]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="partschi">
<emphasis id="6E9B9D47FFEDFFC58F3AFEADFC33011C" box="[831,954,258,284]" italics="true" pageId="24" pageNumber="25">B. partschi</emphasis>
(Hörnes, 1852)
</taxonomicName>
, which led
<bibRefCitation id="387E3CA4FFEDFFC588EEFEADFA15011D" author="Boettger, O." box="[1259,1436,258,285]" pageId="24" pageNumber="25" pagination="1 - 99" refId="ref54689" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso-Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99. [https: // www. biodiversitylibrary. org / item / 108176 # page / 59 / mode / 1 up]" type="journal article" year="1906">Boettger (1906)</bibRefCitation>
to described it as variety of the latter. However, it differs in its broader, regularly conical spire, its less incised suture, its finer sculpture, comprising a higher number of axial ribs and spiral cords, and the absence of a prominent subsutural spiral cord, as well as more oblique suture. In addition,
<taxonomicName id="9BEF3AD6FFEDFFC58F76FEC1FC750188" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[883,1020,366,392]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="kostejana">
<emphasis id="6E9B9D47FFEDFFC58F76FEC1FC750188" box="[883,1020,366,392]" italics="true" pageId="24" pageNumber="25">B. kostejana</emphasis>
</taxonomicName>
is constantly smaller. The occurrence of
<taxonomicName id="9BEF3AD6FFEDFFC58CB0FE3DFEB601AC" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[181,319,402,428]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="kostejana">
<emphasis id="6E9B9D47FFEDFFC58CB0FE3DFEB601AC" box="[181,319,402,428]" italics="true" pageId="24" pageNumber="25">B. kostejana</emphasis>
</taxonomicName>
in offshore marls indicates also an environmental separation from
<taxonomicName id="9BEF3AD6FFEDFFC5882FFE3DFB2C01AC" baseAuthorityName="Hoernes" baseAuthorityYear="1852" box="[1066,1189,402,428]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="partschi">
<emphasis id="6E9B9D47FFEDFFC5882FFE3DFB2C01AC" box="[1066,1189,402,428]" italics="true" pageId="24" pageNumber="25">B. partschi</emphasis>
</taxonomicName>
, which is restricted to near shore occurrences.
</paragraph>
<paragraph id="5C504155FFEDFFC58CC2FE74FDC40260" blockId="24.[151,1437,150,753]" pageId="24" pageNumber="25">
<bibRefCitation id="387E3CA4FFEDFFC58CC2FE74FEF601F4" author="Holzl, O." box="[199,383,474,500]" pageId="24" pageNumber="25" pagination="2" refId="ref58000" refString="Holzl, O. (1973) Faziostratotypus Kaltenbachgraben, Landkreis Miesbach-Aibling, Oberbayern, Bundesrepublik Deutschland (Profil der Gerner, Alberund Kirchberger Schichtenfolge). In: Papp, A., Rogl, F. &amp; Senes, J. (Eds.), M 2 Ottnangien. Die Innviertler, Salgotarjaner, Bantapusztaer Schichtgruppe und die Rzehakia Formation. Chronostratigraphie und Neostratotypen, Miozan der zentralen Paratethys. Vol. 3. Veda, Bratislava, pp. 155 - 196." type="book chapter" year="1973">Hölzl (1973: 93)</bibRefCitation>
mentioned
<taxonomicName id="9BEF3AD6FFEDFFC58E04FE74FCEB01F5" authority="(M. Hoernes)" baseAuthorityName="M. Hoernes" baseAuthorityYear="1852" box="[513,866,474,501]" class="Gastropoda" family="Costellariidae" genus="Thala" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="partschi">
<emphasis id="6E9B9D47FFEDFFC58E04FE74FDCB01F4" box="[513,578,474,500]" italics="true" pageId="24" pageNumber="25">Thala</emphasis>
cf.
<emphasis id="6E9B9D47FFEDFFC58E6EFE74FD4E01F4" box="[619,711,474,500]" italics="true" pageId="24" pageNumber="25">partschi</emphasis>
(M. Hoernes)
</taxonomicName>
from the Ottnangian (middle Burdigalian) of Gernergraben in
<collectingRegion id="9E2B8FB7FFEDFFC58D1BFE50FEFC0218" box="[286,373,510,536]" country="Germany" name="Bayern" pageId="24" pageNumber="25">Bavaria</collectingRegion>
(without illustration). This would be the oldest record for this group from the Paratethys Sea. We have not seen this specimen but the generally poor preservation of the material from Gernergraben usally does not allow unambiguous identifications.
</paragraph>
<paragraph id="5C504155FFEDFFC58CC2FDC4FE1602A9" blockId="24.[151,1437,150,753]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58CC2FDC4FE3F0284" bold="true" box="[199,438,618,644]" pageId="24" pageNumber="25">Palaeoenvironment.</emphasis>
The assemblages from Coşteiu de Sus suggests middle neritic palaeoenvironments with soft bottom (own data).
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFEDFFC58CC2FD1CFECF02F1" pageId="24" pageNumber="25" type="distribution">
<paragraph id="5C504155FFEDFFC58CC2FD1CFECF02F1" blockId="24.[151,1437,150,753]" pageId="24" pageNumber="25">
<emphasis id="6E9B9D47FFEDFFC58CC2FD1CFDE602CC" bold="true" box="[199,623,690,716]" pageId="24" pageNumber="25">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFEDFFC58FC2FD1DFBD802CC" box="[967,1105,691,717]" italics="true" pageId="24" pageNumber="25">Făget Basin</emphasis>
: Coşteiu de Sus, Lăpugiu de Sus (
<collectingCountry id="24F801C5FFEDFFC58CCBFD78FEB302F1" box="[206,314,726,753]" name="Romania" pageId="24" pageNumber="25">Romania</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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