328 lines
39 KiB
XML
328 lines
39 KiB
XML
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<mods:title id="DA45526B39BD4D29CC9C363C598F5E11">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="EA09238151518CD949FC7691A1205C44">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="42FD0BA0CCD435A495FE4E12BBD59F71">Landau, Bernard</mods:namePart>
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<mods:title id="DF8F45CD385F6960D67C46B17D91FE4A">Zootaxa</mods:title>
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<treatment id="D446F043FFE0FFCB8C92FC58FBAD0286" ID-DOI="http://doi.org/10.5281/zenodo.5044012" ID-GBIF-Taxon="183225134" ID-Zenodo-Dep="5044012" LSID="urn:lsid:plazi:treatment:D446F043FFE0FFCB8C92FC58FBAD0286" httpUri="http://treatment.plazi.org/id/D446F043FFE0FFCB8C92FC58FBAD0286" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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<subSubSection id="14F512DEFFE0FFC88C92FC58FE7B0411" box="[151,498,1014,1041]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="5C504155FFE0FFC88C92FC58FE7B0411" blockId="21.[151,498,1014,1083]" box="[151,498,1014,1041]" pageId="21" pageNumber="22">
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<heading id="0718F639FFE0FFC88C92FC58FE7B0411" bold="true" box="[151,498,1014,1041]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
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<emphasis id="6E9B9D47FFE0FFC88C92FC58FE7B0411" bold="true" box="[151,498,1014,1041]" pageId="21" pageNumber="22">
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<taxonomicName id="9BEF3AD6FFE0FFC88C92FC58FE050411" ID-CoL="8KSC9" authority="Harzhauser & Landau, 2021" authorityName="Harzhauser & Landau" authorityYear="2021" box="[151,396,1014,1041]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="dacica" status="sp. nov.">
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<emphasis id="6E9B9D47FFE0FFC88C92FC58FE050411" bold="true" box="[151,396,1014,1041]" italics="true" pageId="21" pageNumber="22">Bellardithala dacica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="75A8203CFFE0FFC88D91FC59FE7B0411" box="[404,498,1015,1041]" pageId="21" pageNumber="22" rank="species">nov. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="14F512DEFFE0FFC88C92FBB5FE4F043B" box="[151,454,1051,1083]" pageId="21" pageNumber="22" type="description">
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<paragraph id="5C504155FFE0FFC88C92FBB5FE4F043B" blockId="21.[151,498,1014,1083]" box="[151,454,1051,1083]" pageId="21" pageNumber="22">
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<figureCitation id="C4D45DD0FFE0FFC88C92FBB5FF780435" box="[151,241,1051,1077]" captionStart="FIGURE 8" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[151,1436,669,1777]" captionTargetId="figure-225@22.[151,1436,669,1777]" captionTargetPageId="22" captionText="FIGURE 8 A1–A2. Bellardithala dacica nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0105. B1–B2. Bellardithala dacica nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0061/0001. C1–C2. Bellardithala dacica nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0061/0002. D1–D2. Bellardithala kostejana (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360340. E1–E2. Bellardithala kostejana (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0064/0001. F. Bellardithala kostejana (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0064/0002. G. Bellardithala sp., Devínska Nová Ves (Slovakia), Z23408-P2003/27." figureDoi="http://doi.org/10.5281/zenodo.4921858" httpUri="https://zenodo.org/record/4921858/files/figure.png" pageId="21" pageNumber="22">Figs 8A</figureCitation>
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<figureCitation id="C4D45DD0FFE0FFC88CF4FB83FE920435" box="[241,283,1051,1083]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="21" pageNumber="22">
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<subScript id="C06B4310FFE0FFC88CF4FB83FF70043B" attach="left" box="[241,249,1069,1083]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
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–A
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</figureCitation>
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<figureCitation id="C4D45DD0FFE0FFC88D1EFB83FEAA043B" box="[283,291,1069,1083]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="21" pageNumber="22">
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<subScript id="C06B4310FFE0FFC88D1EFB83FEAA043B" attach="left" box="[283,291,1069,1083]" fontSize="6" pageId="21" pageNumber="22">2</subScript>
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</figureCitation>
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, B
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<figureCitation id="C4D45DD0FFE0FFC88D41FB83FEE50435" box="[324,364,1051,1083]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="21" pageNumber="22">
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<subScript id="C06B4310FFE0FFC88D41FB83FEC5043B" attach="left" box="[324,332,1069,1083]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
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–B
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</figureCitation>
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<figureCitation id="C4D45DD0FFE0FFC88D68FB83FEFC043B" box="[365,373,1069,1083]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="21" pageNumber="22">
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<subScript id="C06B4310FFE0FFC88D68FB83FEFC043B" attach="right" box="[365,373,1069,1083]" fontSize="6" pageId="21" pageNumber="22">2</subScript>
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</figureCitation>
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, C
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||
<figureCitation id="C4D45DD0FFE0FFC88D93FB83FE370435" box="[406,446,1051,1083]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="21" pageNumber="22">
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<subScript id="C06B4310FFE0FFC88D93FB83FE17043B" attach="left" box="[406,414,1069,1083]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
|
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–C
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||
</figureCitation>
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||
<figureCitation id="C4D45DD0FFE0FFC88DBBFB83FE4F043B" box="[446,454,1069,1083]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="21" pageNumber="22">
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<subScript id="C06B4310FFE0FFC88DBBFB83FE4F043B" attach="left" box="[446,454,1069,1083]" fontSize="6" pageId="21" pageNumber="22">2</subScript>
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</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="14F512DEFFE0FFC88C92FBCEFCA404B5" pageId="21" pageNumber="22" type="reference_group">
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<paragraph id="5C504155FFE0FFC88C92FBCEFC2D0496" blockId="21.[151,1344,1119,1205]" pageId="21" pageNumber="22">
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<taxonomicName id="9BEF3AD6FFE0FFC88C92FBCEFE940477" ID-CoL="7P94F" authority="Laubei" authorityName="Laubei" box="[151,285,1119,1143]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFE0FFC88C92FBCEFE940477" box="[151,285,1119,1143]" italics="true" pageId="21" pageNumber="22">Mitra Laubei</emphasis>
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</taxonomicName>
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nov. form.—
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<bibRefCitation id="387E3CA4FFE0FFC88DA0FBF1FD450477" author="Hoernes, R. & Auinger, M." box="[421,716,1119,1143]" pageId="21" pageNumber="22" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes & Auinger 1880: 89</bibRefCitation>
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(pars) [
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<emphasis id="6E9B9D47FFE0FFC88F19FBCEFCCB0477" box="[796,834,1120,1143]" italics="true" pageId="21" pageNumber="22">non</emphasis>
|
||
<taxonomicName id="9BEF3AD6FFE0FFC88F4DFBF1FABF0477" ID-CoL="8KRR7" authority="(Hoernes & Auinger, 1880)" authorityName="Harzhauser & Landau" authorityYear="2021" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" box="[840,1334,1119,1143]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="laubei">
|
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<emphasis id="6E9B9D47FFE0FFC88F4DFBF1FB9D0477" box="[840,1044,1119,1143]" italics="true" pageId="21" pageNumber="22">Bellardithala laubei</emphasis>
|
||
(
|
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<bibRefCitation id="387E3CA4FFE0FFC88824FBF1FAA40477" author="Hoernes, R. & Auinger, M." box="[1057,1325,1119,1143]" pageId="21" pageNumber="22" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes & Auinger, 1880</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
].
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<taxonomicName id="9BEF3AD6FFE0FFC88C92FBD0FF7E0496" ID-CoL="7PKC6" authorityName="H. Adams & A. Adams" authorityYear="1853" box="[151,247,1150,1174]" class="Gastropoda" family="Clavatulidae" genus="Turricula" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="22" phylum="Mollusca" rank="genus">
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||
<emphasis id="6E9B9D47FFE0FFC88C92FBD0FF7E0496" box="[151,247,1150,1174]" italics="true" pageId="21" pageNumber="22">Turricula</emphasis>
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</taxonomicName>
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||
cf.
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<emphasis id="6E9B9D47FFE0FFC88D1BFBD0FEED0496" box="[286,356,1150,1174]" italics="true" pageId="21" pageNumber="22">Laubei</emphasis>
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R. Hoern. u. Auing.—
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<bibRefCitation id="387E3CA4FFE0FFC88E4CFBD0FCAB0496" author="Friedberg, W." box="[585,802,1150,1174]" pageId="21" pageNumber="22" pagination="1 - 164" refId="ref56467" refString="Friedberg, W. (1938) Katalog mego zbioru mieczakow miocenskich Polski (Katalog meiner Sammlung der Miozanmollusken Polens). Memoires de l'Academie polonaise des sciences et des lettres, Classe des sciences mathematiques et naturelles. Serie B, Sciences naturelles, 12, 1 - 164." type="journal article" year="1938">Friedberg, 1938: 133</bibRefCitation>
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, text-fig. 42.
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||
</paragraph>
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||
<paragraph id="5C504155FFE0FFC88C92FB33FCA404B5" blockId="21.[151,1344,1119,1205]" box="[151,813,1180,1205]" pageId="21" pageNumber="22">
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||
<treatmentCitationGroup id="7CFF667BFFE0FFC88C92FB33FCA404B5" box="[151,813,1180,1205]" pageId="21" pageNumber="22">
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||
?
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||
<taxonomicName id="9BEF3AD6FFE0FFC88CACFB32FE6E04B4" authority="(Hoern. et Auing.)" baseAuthorityName="Hoern. et Auing." box="[169,487,1180,1205]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="laubei">
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<emphasis id="6E9B9D47FFE0FFC88CACFB32FEA004B4" box="[169,297,1180,1204]" italics="true" pageId="21" pageNumber="22">Thala laubei</emphasis>
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(Hoern. et Auing.)
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</taxonomicName>
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—
|
||
<treatmentCitation id="DD4E6744FFE0FFC88E04FB32FD2404B4" author="Kokay, J." box="[513,685,1180,1204]" page="61" pageId="21" pageNumber="22" year="1966">
|
||
<bibRefCitation id="387E3CA4FFE0FFC88E04FB32FD2404B4" author="Kokay, J." box="[513,685,1180,1204]" pageId="21" pageNumber="22" pagination="1 - 149" refId="ref58442" refString="Kokay, J. (1966) Geologische und palaontologische Untersuchung des Braunkohlengebietes von Herend-Marko (Bakony-Gebirge, Ungarn). Geologica Hungarica, series Palaeontologica, 36, 1 - 149. [https: // epa. oszk. hu / 02900 / 02989 / 00029 / pdf /]" type="journal article" year="1966">Kókay, 1966: 61</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 9, fig. 4.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE0FFC88C92FB4DFD4905B2" pageId="21" pageNumber="22" type="materials_examined">
|
||
<paragraph id="5C504155FFE0FFC88C92FB4DFB3C054B" blockId="21.[151,1437,1250,2033]" pageId="21" pageNumber="22">
|
||
<emphasis id="6E9B9D47FFE0FFC88C92FB4DFECE04FC" bold="true" box="[151,327,1250,1277]" pageId="21" pageNumber="22">Type material.</emphasis>
|
||
<materialsCitation id="EC874B08FFE0FFC88D4AFB4DFF650527" ID-GBIF-Occurrence="3311814322" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="21" pageNumber="22" specimenCode="NHMW 1854" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="8354FFF7FFE0FFC88D4AFB4DFE3504FD" box="[335,444,1251,1277]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCode id="0C49E92EFFE0FFC88DCDFB4DFDEA04FD" box="[456,611,1250,1277]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22" type="Museum">NHMW 1854</specimenCode>
|
||
/0035/0105, SL:
|
||
<quantity id="9B17ECB0FFE0FFC88F20FB4DFCF704FD" box="[805,894,1251,1277]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="21" pageNumber="22" unit="mm" value="6.9">6.9 mm</quantity>
|
||
, MD:
|
||
<quantity id="9B17ECB0FFE0FFC88FCCFB4DFBAA04FD" box="[969,1059,1251,1277]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="mm" value="2.4">2.4 mm</quantity>
|
||
,
|
||
<location id="5930178EFFE0FFC88835FB4DFB6804FD" LSID="urn:lsid:plazi:treatment:D446F043FFE0FFCB8C92FC58FBAD0286:5930178EFFE0FFC88835FB4DFB6804FD" box="[1072,1249,1251,1277]" country="Romania" name="Lapugiu de Sus" pageId="21" pageNumber="22">
|
||
Lăpugiu
|
||
<collectorName id="F11A2483FFE0FFC88893FB4DFB6804FD" box="[1174,1249,1251,1277]" pageId="21" pageNumber="22">de Sus</collectorName>
|
||
</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE0FFC888F7FB4DFAD404FE" box="[1266,1373,1251,1278]" name="Romania" pageId="21" pageNumber="22">Romania</collectingCountry>
|
||
), figs 8A
|
||
<subScript id="C06B4310FFE0FFC88CBCFAB7FF480527" attach="left" box="[185,193,1305,1319]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
|
||
–A
|
||
<subScript id="C06B4310FFE0FFC88CE1FAB7FF650527" attach="left" box="[228,236,1305,1319]" fontSize="6" pageId="21" pageNumber="22">2</subScript>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="EC874B08FFE0FFC88CFFFAA9FA1D0527" ID-GBIF-Occurrence="3311814480" box="[250,1428,1287,1319]" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="21" pageNumber="22" specimenCode="NHMW 2020" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="8354FFF7FFE0FFC88CFFFAA9FEE20521" box="[250,363,1287,1313]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCode id="0C49E92EFFE0FFC88D73FAA9FD9A0521" box="[374,531,1287,1313]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22" type="Museum">NHMW 2020</specimenCode>
|
||
/0061/0001, SL:
|
||
<quantity id="9B17ECB0FFE0FFC88ECAFAA9FCA10521" box="[719,808,1287,1313]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="21" pageNumber="22" unit="mm" value="7.3">7.3 mm</quantity>
|
||
, MD:
|
||
<quantity id="9B17ECB0FFE0FFC88F75FAA9FC410521" box="[880,968,1287,1313]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="mm" value="2.4">2.4 mm</quantity>
|
||
,
|
||
<location id="5930178EFFE0FFC88FD1FAA9FB0A0521" LSID="urn:lsid:plazi:treatment:D446F043FFE0FFCB8C92FC58FBAD0286:5930178EFFE0FFC88FD1FAA9FB0A0521" box="[980,1155,1287,1313]" country="Romania" name="Lapugiu de Sus" pageId="21" pageNumber="22">
|
||
Lăpugiu
|
||
<collectorName id="F11A2483FFE0FFC8883CFAA9FB0A0521" box="[1081,1155,1287,1313]" pageId="21" pageNumber="22">de Sus</collectorName>
|
||
</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE0FFC88896FAA9FB770522" box="[1171,1278,1287,1314]" name="Romania" pageId="21" pageNumber="22">Romania</collectingCountry>
|
||
), figs 8B
|
||
<subScript id="C06B4310FFE0FFC88966FAB7FAE20527" attach="left" box="[1379,1387,1305,1319]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
|
||
–B
|
||
<subScript id="C06B4310FFE0FFC88989FAB7FA1D0527" attach="left" box="[1420,1428,1305,1319]" fontSize="6" pageId="21" pageNumber="22">2</subScript>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="EC874B08FFE0FFC88C92FA85FB38054B" ID-GBIF-Occurrence="3311814377" box="[151,1201,1323,1355]" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="21" pageNumber="22" specimenCode="NHMW 2020" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="0C49E92EFFE0FFC88C92FA85FEBB0545" box="[151,306,1323,1349]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22" type="Museum">NHMW 2020</specimenCode>
|
||
/0061/0002, SL:
|
||
<quantity id="9B17ECB0FFE0FFC88DEBFA85FDCF0545" box="[494,582,1323,1349]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="21" pageNumber="22" unit="mm" value="6.9">6.9 mm</quantity>
|
||
, MD:
|
||
<quantity id="9B17ECB0FFE0FFC88E8BFA85FD6F0545" box="[654,742,1323,1349]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="mm" value="2.4">2.4 mm</quantity>
|
||
,
|
||
<location id="5930178EFFE0FFC88EF4FA85FC160545" LSID="urn:lsid:plazi:treatment:D446F043FFE0FFCB8C92FC58FBAD0286:5930178EFFE0FFC88EF4FA85FC160545" box="[753,927,1323,1349]" country="Romania" name="Lapugiu de Sus" pageId="21" pageNumber="22">
|
||
Lăpugiu
|
||
<collectorName id="F11A2483FFE0FFC88F53FA85FC160545" box="[854,927,1323,1349]" pageId="21" pageNumber="22">de Sus</collectorName>
|
||
</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE0FFC88FABFA85FB900546" box="[942,1049,1323,1350]" name="Romania" pageId="21" pageNumber="22">Romania</collectingCountry>
|
||
), figs 8C
|
||
<subScript id="C06B4310FFE0FFC8887BFA93FB0F054B" attach="left" box="[1150,1158,1341,1355]" fontSize="6" pageId="21" pageNumber="22">1</subScript>
|
||
–C
|
||
<subScript id="C06B4310FFE0FFC888A2FA93FB38054B" attach="left" box="[1191,1201,1341,1355]" fontSize="6" pageId="21" pageNumber="22">2</subScript>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE0FFC88CC2FAE1FCBD0569" blockId="21.[151,1437,1250,2033]" box="[199,820,1358,1386]" pageId="21" pageNumber="22">
|
||
<materialsCitation id="EC874B08FFE0FFC88CC2FAE1FCB90569" ID-GBIF-Occurrence="3311814453" box="[199,816,1358,1386]" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="6E9B9D47FFE0FFC88CC2FAE1FEEF0568" bold="true" box="[199,358,1358,1385]" pageId="21" pageNumber="22">
|
||
<typeStatus id="8354FFF7FFE0FFC88CC2FAE1FE8B0569" box="[199,258,1359,1385]" pageId="21" pageNumber="22">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="5930178EFFE0FFC88D69FAE1FD930569" LSID="urn:lsid:plazi:treatment:D446F043FFE0FFCB8C92FC58FBAD0286:5930178EFFE0FFC88D69FAE1FD930569" box="[364,538,1359,1385]" country="Romania" name="Lapugiu de Sus" pageId="21" pageNumber="22">
|
||
Lăpugiu
|
||
<collectorName id="F11A2483FFE0FFC88DD4FAE1FD930569" box="[465,538,1359,1385]" pageId="21" pageNumber="22">de Sus</collectorName>
|
||
</location>
|
||
(
|
||
<collectingCountry id="24F801C5FFE0FFC88E2CFAE1FD1D056A" box="[553,660,1359,1386]" name="Romania" pageId="21" pageNumber="22">Romania</collectingCountry>
|
||
), Făget Basin
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE0FFC88CC2FADDFD75058D" blockId="21.[151,1437,1250,2033]" box="[199,764,1395,1421]" pageId="21" pageNumber="22">
|
||
<emphasis id="6E9B9D47FFE0FFC88CC2FADDFEE6058D" bold="true" box="[199,367,1395,1421]" pageId="21" pageNumber="22">
|
||
<typeStatus id="8354FFF7FFE0FFC88CC2FADDFE8B058D" box="[199,258,1395,1421]" pageId="21" pageNumber="22">Type</typeStatus>
|
||
stratum.
|
||
</emphasis>
|
||
Silt and clay of the Dej Formation.
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE0FFC88CC2FA38FD4905B2" blockId="21.[151,1437,1250,2033]" box="[199,704,1430,1458]" pageId="21" pageNumber="22">
|
||
<emphasis id="6E9B9D47FFE0FFC88CC2FA38FF7405B0" bold="true" box="[199,253,1430,1456]" pageId="21" pageNumber="22">Age.</emphasis>
|
||
Middle Miocene, Badenian (Langhian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE0FFC88CC2FA15FE9F05F9" pageId="21" pageNumber="22" type="etymology">
|
||
<paragraph id="5C504155FFE0FFC88CC2FA15FE9F05F9" blockId="21.[151,1437,1250,2033]" pageId="21" pageNumber="22">
|
||
<emphasis id="6E9B9D47FFE0FFC88CC2FA15FEC705D5" bold="true" box="[199,334,1467,1493]" pageId="21" pageNumber="22">Etymology.</emphasis>
|
||
Referring to Dacia, a province of the Roman Empire, covering also the territory of the Transylvanian Basin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE0FFC88CC2F9ACFDFB0689" pageId="21" pageNumber="22" type="diagnosis">
|
||
<paragraph id="5C504155FFE0FFC88CC2F9ACFDFB0689" blockId="21.[151,1437,1250,2033]" pageId="21" pageNumber="22">
|
||
<emphasis id="6E9B9D47FFE0FFC88CC2F9ACFECB061C" bold="true" box="[199,322,1538,1564]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
|
||
<taxonomicName id="9BEF3AD6FFE0FFC88D4FF9ADFDBE061D" box="[330,567,1539,1565]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="6E9B9D47FFE0FFC88D4FF9ADFE57061D" box="[330,478,1539,1565]" italics="true" pageId="21" pageNumber="22">Bellardithala</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of medium size, moderately slender fusiform shell, with faintly cyrtoconoid slightly gradate spire, differing from congeneric species by its sculpture, which changes from prominent axial ribs and spiral cords with distinct nodes on early teleoconch whorls to densely spaced, cancellate pattern, with blurred nodes on later spire whorls and last whorl.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE0FFCB8CC2F93CFEF80141" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
|
||
<paragraph id="5C504155FFE0FFCB8CC2F93CFE5300F9" blockId="21.[151,1437,1250,2033]" lastBlockId="22.[151,1436,151,646]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<emphasis id="6E9B9D47FFE0FFC88CC2F93CFED006AC" bold="true" box="[199,345,1682,1708]" pageId="21" pageNumber="22">Description.</emphasis>
|
||
Shell medium-sized, slender fusiform with faintly cyrtoconoid and slightly gradate spire. Protoconch largely damaged; last two protoconch whorls suggesting high conical shape. Teleoconch of 4.5 weakly convex whorl, separated by impressed suture, with periphery shifting from abapical suture to mid-whorl during growth. First teleoconch whorl with axial ribs separated by interspaces of about same width, crossed by five close-set spiral cords, forming spirally elongated nodes at intersections with ribs, persisting into interspaces. Nodes on adapical spiral row most prominent. Axial ribs broader and less prominent on later teleoconch whorls. Five to six spiral cords on later spire whorls, forming cancellate pattern with axial ribs. Spiral cord at adapical suture broader than others, separated by wider and deeper spiral groove from next spiral cord. Last whorl high, weakly convex, slowly contracting, with up to 25 densely spaced spiral cords. Nodes on spiral cords somewhat blurred above periphery and largely reduced below. Aperture elongate, narrow to moderately narrow, with indistinct anal notch. Columellar callus narrow, indistinct. Columella with four prominent columellar folds, weakening only slightly abapically. Outer lip weakly thickened, with about 11 delicate denticles within. Siphonal canal moderately long, bent slightly to left, with shallow siphonal notch.
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE3FFCB8CC2FEACFEF80141" blockId="22.[151,1436,151,646]" pageId="22" pageNumber="23">
|
||
<emphasis id="6E9B9D47FFE3FFCB8CC2FEACFDBC011C" bold="true" box="[199,565,258,285]" pageId="22" pageNumber="23">Shell measurements and ratios</emphasis>
|
||
. SL:
|
||
<quantity id="9B17ECB0FFE3FFCB8E71FEADFD74011D" box="[628,765,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="7.3" metricValueMin="6.9" pageId="22" pageNumber="23" unit="mm" value="7.1" valueMax="7.3" valueMin="6.9">6.9–7.3 mm</quantity>
|
||
, MD:
|
||
<quantity id="9B17ECB0FFE3FFCB8F40FEADFC14011D" box="[837,925,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="22" pageNumber="23" unit="mm" value="2.4">2.4 mm</quantity>
|
||
, AA: 35–40°, SL/MD: 2.8–3.1, AL/AW: 5.0– 5.5, AH/S: 2.1–2.3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE3FFCB8CC2FEE4FB07023D" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="5C504155FFE3FFCB8CC2FEE4FAD601D1" blockId="22.[151,1436,151,646]" pageId="22" pageNumber="23">
|
||
<emphasis id="6E9B9D47FFE3FFCB8CC2FEE4FEC50164" bold="true" box="[199,332,330,356]" pageId="22" pageNumber="23">Discussion.</emphasis>
|
||
The
|
||
<typeStatus id="8354FFF7FFE3FFCB8D82FEE5FE600165" box="[391,489,331,357]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB8E14FEE5FD710165" authority="Harzhauser & Landau, 2021" authorityName="Harzhauser & Landau" authorityYear="2021" box="[529,760,331,357]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="dacica" status="sp. nov.">
|
||
<emphasis id="6E9B9D47FFE3FFCB8E14FEE5FD710165" box="[529,760,331,357]" italics="true" pageId="22" pageNumber="23">Bellardithala dacica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="75A8203CFFE3FFCB8F05FEE5FCDE0165" box="[768,855,331,357]" pageId="22" pageNumber="23" rank="species">nov. sp.</taxonomicNameLabel>
|
||
was mixed with
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB881FFEE5FF500189" authority="(Hoernes & Auinger, 1880)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE3FFCB881FFEE5FB210165" box="[1050,1192,331,357]" italics="true" pageId="22" pageNumber="23">Mitra laubei</emphasis>
|
||
(
|
||
<bibRefCitation id="387E3CA4FFE3FFCB88B2FEE5FF590189" author="Hoernes, R. & Auinger, M." pageId="22" pageNumber="23" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. & Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes & Auinger, 1880</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by those authors. Despite some superficial similarity due to the prominent subsutural spiral cord, the species are distinguished by the slender shape of
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB8E66FE3DFD4E01AD" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" box="[611,711,403,429]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE3FFCB8E66FE3DFD4E01AD" box="[611,711,403,429]" italics="true" pageId="22" pageNumber="23">B. laubei</emphasis>
|
||
</taxonomicName>
|
||
and the different sculpture of early spire whorls (broad axial ribs with subordinate spiral sculpture in
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB8E2CFE19FD0701D1" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1880" box="[553,654,439,465]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE3FFCB8E2CFE19FD0701D1" box="[553,654,439,465]" italics="true" pageId="22" pageNumber="23">B. laubei</emphasis>
|
||
</taxonomicName>
|
||
vs narrower axial ribs and prominent spiral cords in
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB88DAFE19FAC601D1" authorityName="Harzhauser & Landau" authorityYear="2021" box="[1247,1359,439,465]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="dacica">
|
||
<emphasis id="6E9B9D47FFE3FFCB88DAFE19FAC601D1" box="[1247,1359,439,465]" italics="true" pageId="22" pageNumber="23">M. dacica</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5C504155FFE3FFCB8CC2FE75FB07023D" blockId="22.[151,1436,151,646]" pageId="22" pageNumber="23">
|
||
A specimen described by
|
||
<bibRefCitation id="387E3CA4FFE3FFCB8DECFE75FD0F01F6" author="Kokay, J." box="[489,646,475,502]" pageId="22" pageNumber="23" pagination="1 - 149" refId="ref58442" refString="Kokay, J. (1966) Geologische und palaontologische Untersuchung des Braunkohlengebietes von Herend-Marko (Bakony-Gebirge, Ungarn). Geologica Hungarica, series Palaeontologica, 36, 1 - 149. [https: // epa. oszk. hu / 02900 / 02989 / 00029 / pdf /]" type="journal article" year="1966">Kókay (1966)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB8EA9FE75FCB201F5" box="[684,827,475,501]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE3FFCB8EA9FE75FCB201F5" box="[684,827,475,501]" italics="true" pageId="22" pageNumber="23">Thala laubei</emphasis>
|
||
</taxonomicName>
|
||
from Herend (
|
||
<collectingCountry id="24F801C5FFE3FFCB8FE7FE75FBCE01F5" box="[994,1095,475,501]" name="Hungary" pageId="22" pageNumber="23">Hungary</collectingCountry>
|
||
) differs from
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB88E0FE75FAC301F5" box="[1253,1354,475,501]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="laubei">
|
||
<emphasis id="6E9B9D47FFE3FFCB88E0FE75FAC301F5" box="[1253,1354,475,501]" italics="true" pageId="22" pageNumber="23">B. laubei</emphasis>
|
||
</taxonomicName>
|
||
also in its broader shape. It is reminiscent of
|
||
<taxonomicName id="9BEF3AD6FFE3FFCB8E39FE51FD430219" authorityName="Harzhauser & Landau" authorityYear="2021" box="[572,714,511,537]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="dacica">
|
||
<emphasis id="6E9B9D47FFE3FFCB8E39FE51FDDC0218" box="[572,597,511,536]" italics="true" pageId="22" pageNumber="23">B.</emphasis>
|
||
cf.
|
||
<emphasis id="6E9B9D47FFE3FFCB8E7AFE51FD430219" box="[639,714,511,537]" italics="true" pageId="22" pageNumber="23">dacica</emphasis>
|
||
</taxonomicName>
|
||
but differs in its higher number of axial ribs on the spire and the distinct subsutural spiral cord. This specimen might indicate a further undescribed species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="14F512DEFFE3FFCB8CC2FDE8FBAD0286" pageId="22" pageNumber="23" type="distribution">
|
||
<paragraph id="5C504155FFE3FFCB8CC2FDE8FBAD0286" blockId="22.[151,1436,151,646]" pageId="22" pageNumber="23">
|
||
<emphasis id="6E9B9D47FFE3FFCB8CC2FDE8FDEA0261" bold="true" box="[199,611,582,609]" pageId="22" pageNumber="23">Distribution in Central Paratethys.</emphasis>
|
||
Badenian (middle Miocene):
|
||
<emphasis id="6E9B9D47FFE3FFCB8FAEFDE9FAA10260" box="[939,1320,583,609]" italics="true" pageId="22" pageNumber="23">Ukrainian Fore-Carpathian Basin</emphasis>
|
||
: Żukowce (
|
||
<emphasis id="6E9B9D47FFE3FFCB8CA4FDC5FE810285" box="[161,264,619,645]" italics="true" pageId="22" pageNumber="23">Zhukivtsi</emphasis>
|
||
) (
|
||
<collectingCountry id="24F801C5FFE3FFCB8D1AFDC5FEF40286" box="[287,381,619,646]" name="Ukraine" pageId="22" pageNumber="23">Ukraine</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="387E3CA4FFE3FFCB8D96FDC5FDCD0285" author="Friedberg, W." box="[403,580,619,645]" pageId="22" pageNumber="23" pagination="1 - 164" refId="ref56467" refString="Friedberg, W. (1938) Katalog mego zbioru mieczakow miocenskich Polski (Katalog meiner Sammlung der Miozanmollusken Polens). Memoires de l'Academie polonaise des sciences et des lettres, Classe des sciences mathematiques et naturelles. Serie B, Sciences naturelles, 12, 1 - 164." type="journal article" year="1938">Friedberg 1938</bibRefCitation>
|
||
);
|
||
<emphasis id="6E9B9D47FFE3FFCB8E5DFDC5FD690284" box="[600,736,619,645]" italics="true" pageId="22" pageNumber="23">Făget Basin</emphasis>
|
||
: Lăpugiu de Sus (
|
||
<collectingCountry id="24F801C5FFE3FFCB8FA9FDC5FB910286" box="[940,1048,619,646]" name="Romania" pageId="22" pageNumber="23">Romania</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |