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<document id="917FF84D23636462BE8D7086D35ACC54" ID-CLB-Dataset="5925" ID-DOI="10.11646/zootaxa.4982.1.1" ID-GBIF-Dataset="f68b381d-d0dd-44e2-b9fe-6cf3c1aab1fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921842" ID-ZooBank="58388FB8-128A-4381-83D1-3C508D0D3873" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623285106829" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2021" docId="D446F043FFDDFFF58C92FF39FE450430" docLanguage="en" docName="zootaxa.4982.1.1.pdf" docOrigin="Zootaxa 4982 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pusia moravica Harzhauser &amp; Landau 2021, nov. comb." docType="treatment" docVersion="10" lastPageNumber="41" masterDocId="287F883BFFF5FFDD8C05FFAEFF890000" masterDocTitle="An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea" masterLastPageNumber="70" masterPageNumber="1" pageNumber="41" updateTime="1699009793899" updateUser="ExternalLinkService">
<mods:mods id="4DA16CBC504E59EFD64D20E3DF29496A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="514CEAD5DEB1B4D2BA2FD73C0B1A48CC">
<mods:title id="A497ED21F43EDEF684C19D2D43F0FBD6">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
</mods:titleInfo>
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<mods:namePart id="2486E37DDA982F6926745258F07FCA2E">Harzhauser, Mathias</mods:namePart>
<mods:nameIdentifier id="9555BDDA40C6701D9350D51B9C114015" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
<mods:nameIdentifier id="DD885B00439467C72CDBE4FD87823FF8" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="59A56D6402FB68FBA5CA4A6A60234CDA">Landau, Bernard</mods:namePart>
<mods:nameIdentifier id="420D5B2F43C11A91728E95672E88A2AB" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
<mods:nameIdentifier id="633DA5933486258D8EDE40E2EBD6609D" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:title id="ED6ED68425079CF50CDD3E5E1D223093">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="71031E5B57DA1B2ABF61893C4BAA4C2E">
<mods:date id="8B2739836EA0C1140D1FB5DDB76BF103">2021</mods:date>
<mods:detail id="0D84E3DFC0244D9ED03C2DFCDB3800F3" type="pubDate">
<mods:number id="DF98020B619E263B59ACE86DB15BB16A">2021-06-09</mods:number>
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<mods:detail id="F7A9C6F27E0DD8D128CBB038CF7A28D2" type="volume">
<mods:number id="02586EFEFDBB03AC5F79766E2446834C">4982</mods:number>
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<mods:classification id="512AD5D56ADFBE4DD1E9E85DE7761D96">journal article</mods:classification>
<mods:identifier id="324BBBB24B8A210494DF99F4028B10F2" type="CLB-Dataset">5925</mods:identifier>
<mods:identifier id="FE0BEDD4D4C25C24C3282CB9DF020A68" type="DOI">10.11646/zootaxa.4982.1.1</mods:identifier>
<mods:identifier id="F126DDA4E7AE63EB76588F03D9E7103E" type="GBIF-Dataset">f68b381d-d0dd-44e2-b9fe-6cf3c1aab1fa</mods:identifier>
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<treatment id="D446F043FFDDFFF58C92FF39FE450430" ID-DOI="http://doi.org/10.5281/zenodo.5044026" ID-GBIF-Taxon="183225099" ID-Zenodo-Dep="5044026" LSID="urn:lsid:plazi:treatment:D446F043FFDDFFF58C92FF39FE450430" httpUri="http://treatment.plazi.org/id/D446F043FFDDFFF58C92FF39FE450430" lastPageNumber="41" pageId="40" pageNumber="41">
<subSubSection id="14F512DEFFDDFFF58C92FF39FCBE00B1" box="[151,823,151,177]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="5C504155FFDDFFF58C92FF39FCBE00B1" blockId="40.[151,823,151,220]" box="[151,823,151,177]" pageId="40" pageNumber="41">
<heading id="0718F639FFDDFFF58C92FF39FCBE00B1" bold="true" box="[151,823,151,177]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="4">
<emphasis id="6E9B9D47FFDDFFF58C92FF39FCBE00B1" bold="true" box="[151,823,151,177]" pageId="40" pageNumber="41">
<taxonomicName id="9BEF3AD6FFDDFFF58C92FF39FD2000B1" authority="(Hoernes &amp; Auinger, 1880)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[151,681,151,177]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="moravica" status="comb. nov.">
<emphasis id="6E9B9D47FFDDFFF58C92FF39FEDA00B1" bold="true" box="[151,339,151,177]" italics="true" pageId="40" pageNumber="41">Pusia moravica</emphasis>
(
<bibRefCitation id="387E3CA4FFDDFFF58D66FF39FD2800B1" author="Hoernes, R. &amp; Auinger, M." box="[355,673,151,177]" pageId="40" pageNumber="41" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFDDFFF58EB4FF39FCBE00B1" box="[689,823,151,177]" pageId="40" pageNumber="41" rank="species">nov. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDDFFF58C92FF15FEBD00D6" box="[151,308,187,220]" pageId="40" pageNumber="41" type="description">
<paragraph id="5C504155FFDDFFF58C92FF15FEBD00D6" blockId="40.[151,823,151,220]" box="[151,308,187,220]" pageId="40" pageNumber="41">
<figureCitation id="C4D45DD0FFDDFFF58C92FF15FF7D00D5" box="[151,244,187,213]" captionStart="FIGURE 13" captionStartId="39.[151,250,1651,1675]" captionTargetBox="[151,1435,181,1627]" captionTargetId="figure-17@39.[151,1436,181,1627]" captionTargetPageId="39" captionText="FIGURE 13 A1–A2. Pusia avellanella (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360343. B1–B2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0002. C1–C2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. D1–D2. Pusia avellanella (Boettger, 1906), Coşteiu de Sus (Romania), SMF 360344. E1–E2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0004. F1–F2. Pusia avellanella (Boettger, 1906), Steinebrunn (Austria), NHMW 1847/0037/0048. G1–G2. Pusia falsitranssylvanica nov. nom., holotype, Coşteiu de Sus (Roma- nia), SMF 360346. H1–H2. Pusia confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. I1–I2. Pusia moravica (Hoernes &amp; Auinger, 1880), lectotype, Rudice (Czech Republic), NHMW 1862/0029/0006. J. Pusia moravica (Hoernes &amp; Auinger, 1880), paralectotype, Rudice (Czech Republic), NHMW 2020/0097/0001." figureDoi="http://doi.org/10.5281/zenodo.4921868" httpUri="https://zenodo.org/record/4921868/files/figure.png" pageId="40" pageNumber="41">Figs 13I</figureCitation>
<figureCitation id="C4D45DD0FFDDFFF58CF1FF60FE9D00D6" box="[244,276,188,220]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="40" pageNumber="41">
<subScript id="C06B4310FFDDFFF58CF1FF60FF7500DC" attach="left" box="[244,252,206,220]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
–I
</figureCitation>
<figureCitation id="C4D45DD0FFDDFFF58D16FF60FE9200DC" box="[275,283,206,220]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="40" pageNumber="41">
<subScript id="C06B4310FFDDFFF58D16FF60FE9200DC" attach="left" box="[275,283,206,220]" fontSize="6" pageId="40" pageNumber="41">2</subScript>
</figureCitation>
, J
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDDFFF58C92FEAEFCB90118" box="[151,816,256,280]" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="5C504155FFDDFFF58C92FEAEFCB90118" blockId="40.[151,816,256,280]" box="[151,816,256,280]" pageId="40" pageNumber="41">
<treatmentCitationGroup id="7CFF667BFFDDFFF58C92FEAEFCB90118" box="[151,816,256,280]" pageId="40" pageNumber="41">
*
<taxonomicName id="9BEF3AD6FFDDFFF58CAFFEAEFEFA0117" authority="Var." authorityName="Var." box="[170,371,256,280]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="plicatula">
<emphasis id="6E9B9D47FFDDFFF58CAFFEAEFECD0118" box="[170,324,256,280]" italics="true" pageId="40" pageNumber="41">Mitra plicatula</emphasis>
Var.
</taxonomicName>
(
<taxonomicName id="9BEF3AD6FFDDFFF58D84FEAEFE680117" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[385,481,256,279]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="moravica">
<emphasis id="6E9B9D47FFDDFFF58D84FEAEFE680117" box="[385,481,256,279]" italics="true" pageId="40" pageNumber="41">moravica</emphasis>
</taxonomicName>
)—
<treatmentCitation id="DD4E6744FFDDFFF58E07FEAEFCA20118" author="Hoernes, R. &amp; Auinger, M." box="[514,811,256,280]" page="84" pageId="40" pageNumber="41" year="1880">
<bibRefCitation id="387E3CA4FFDDFFF58E07FEAEFCA20118" author="Hoernes, R. &amp; Auinger, M." box="[514,811,256,280]" pageId="40" pageNumber="41" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger 1880: 84</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDDFFF58C92FEE8FD6302EC" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="5C504155FFDDFFF58C92FEE8FEC501AE" blockId="40.[151,1437,325,1072]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58C92FEE8FECD015F" bold="true" box="[151,324,325,352]" pageId="40" pageNumber="41">Type material.</emphasis>
<materialsCitation id="EC874B08FFDDFFF58D4FFEE8FE1C018A" ID-GBIF-Occurrence="3311814301" collectionCode="NHMW" country="Czech Republic" location="Rudice" pageId="40" pageNumber="41" specimenCode="NHMW 1862" specimenCount="1" typeStatus="lectotype">
<typeStatus id="8354FFF7FFDDFFF58D4FFEE8FE330160" box="[330,442,326,352]" pageId="40" pageNumber="41" type="lectotype">Lectotype</typeStatus>
(designated herein):
<specimenCode id="0C49E92EFFDDFFF58EA1FEE8FCB40160" box="[676,829,326,352]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="41" type="Museum">NHMW 1862</specimenCode>
/0029/0006, SL:
<quantity id="9B17ECB0FFDDFFF58FFCFEE8FBD90160" box="[1017,1104,326,352]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="40" pageNumber="41" unit="mm" value="9.8">9.8 mm</quantity>
, MD:
<quantity id="9B17ECB0FFDDFFF58893FEE8FB640160" box="[1174,1261,326,352]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="40" pageNumber="41" unit="mm" value="4.7">4.7 mm</quantity>
,
<location id="5930178EFFDDFFF588FDFEE8FACE0160" LSID="urn:lsid:plazi:treatment:D446F043FFDDFFF58C92FF39FE450430:5930178EFFDDFFF588FDFEE8FACE0160" box="[1272,1351,326,352]" country="Czech Republic" name="Rudice" pageId="40" pageNumber="41">Rudice</location>
(
<collectingCountry id="24F801C5FFDDFFF58951FEE8FE880185" name="Czech Republic" pageId="40" pageNumber="41">Czech Republic</collectingCountry>
), figs 13I
<subScript id="C06B4310FFDDFFF58D68FED2FEFC018A" attach="left" box="[365,373,380,394]" fontSize="6" pageId="40" pageNumber="41">1</subScript>
–I
<subScript id="C06B4310FFDDFFF58D88FED2FE1C018A" attach="left" box="[397,405,380,394]" fontSize="6" pageId="40" pageNumber="41">2</subScript>
</materialsCitation>
.
<materialsCitation id="EC874B08FFDDFFF58DA0FEC4FEC001A8" ID-GBIF-Occurrence="3311814457" collectionCode="NHMW" country="Czech Republic" location="Rudice" pageId="40" pageNumber="41" specimenCode="NHMW 2020" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="8354FFF7FFDDFFF58DA0FEC4FDCB0184" box="[421,578,362,388]" pageId="40" pageNumber="41" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCode id="0C49E92EFFDDFFF58E4BFEC4FD620184" box="[590,747,362,388]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="41" type="Museum">NHMW 2020</specimenCode>
/0097/0001, SL:
<quantity id="9B17ECB0FFDDFFF58FAFFEC4FB9B0184" box="[938,1042,362,388]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="40" pageNumber="41" unit="mm" value="12.6">12.6 mm</quantity>
, MD:
<quantity id="9B17ECB0FFDDFFF58858FEC4FB3F0184" box="[1117,1206,362,389]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="40" pageNumber="41" unit="mm" value="6.5">6.5 mm</quantity>
,
<location id="5930178EFFDDFFF588C1FEC4FA9A0184" LSID="urn:lsid:plazi:treatment:D446F043FFDDFFF58C92FF39FE450430:5930178EFFDDFFF588C1FEC4FA9A0184" box="[1220,1299,362,388]" country="Czech Republic" name="Rudice" pageId="40" pageNumber="41">Rudice</location>
(
<collectingCountry id="24F801C5FFDDFFF58926FEC4FF6801A9" name="Czech Republic" pageId="40" pageNumber="41">Czech Republic</collectingCountry>
), fig. 13J
</materialsCitation>
<subScript id="C06B4310FFDDFFF58D4DFE0EFEC501AE" attach="left" box="[328,332,416,430]" fontSize="6" pageId="40" pageNumber="41">.</subScript>
</paragraph>
<paragraph id="5C504155FFDDFFF58CC2FE1FFBFD02A4" blockId="40.[151,1437,325,1072]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2FE1FFE3001CB" bold="true" box="[199,441,433,459]" pageId="40" pageNumber="41">Revised description.</emphasis>
Shell moderately large, stout fusiform, with slightly gradate spire. Protoconch and early teleoconch unknown. Teleoconch whorls low, weakly convex, with periphery in abapical third; suture impressed. Sculpture of sharp crested, orthocline to weakly prosocline ribs, separated by interspaces of about same width (about 15 axial ribs on penultimate whorl). Adapical weak subsutural spiral groove cuts ribs, forming narrow subsutural spiral cord of weak nodes. Further delicate spirals placed below subsutural cord, abapically rapidly weakening. Last whorl convex with short, moderately constricted base. Axial ribs becoming subobsolete on last whorl. Columella with four prominent, oblique columellar folds. Siphonal canal moderately long, straight.
</paragraph>
<paragraph id="5C504155FFDDFFF58CC2FD03FD6302EC" blockId="40.[151,1437,325,1072]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2FD03FDB302C7" bold="true" box="[199,570,685,712]" pageId="40" pageNumber="41">Shell measurements and ratios.</emphasis>
SL:
<quantity id="9B17ECB0FFDDFFF58E6AFD00FC8C02C8" box="[623,773,686,712]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.26" metricValueMin="0.98" pageId="40" pageNumber="41" unit="mm" value="11.2" valueMax="12.6" valueMin="9.8">9.8–12.6 mm</quantity>
, MD:
<quantity id="9B17ECB0FFDDFFF58F4EFD00FC5B02C8" box="[843,978,686,713]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="6.5" metricValueMin="4.7" pageId="40" pageNumber="41" unit="mm" value="5.6" valueMax="6.5" valueMin="4.7">4.7–6.5 mm</quantity>
(fragments; restored height:
<quantity id="9B17ECB0FFDDFFF58916FD00FA1902C8" box="[1299,1424,686,712]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="40" pageNumber="41" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13–16 mm</quantity>
); assumed SL of complete specimen:
<quantity id="9B17ECB0FFDDFFF58E2FFD7CFDF202EC" box="[554,635,722,748]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="40" pageNumber="41" unit="mm" value="18.0">18 mm</quantity>
, AA: 33°.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDDFFF58CC2FD5BFF5E03E8" pageId="40" pageNumber="41" type="discussion">
<paragraph id="5C504155FFDDFFF58CC2FD5BFD8E03A0" blockId="40.[151,1437,325,1072]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2FD5BFEC5030F" bold="true" box="[199,332,757,783]" pageId="40" pageNumber="41">Discussion.</emphasis>
Only two fragments are available, which
<bibRefCitation id="387E3CA4FFDDFFF58F18FD58FBC10310" author="Hoernes, R. &amp; Auinger, M." box="[797,1096,758,784]" pageId="40" pageNumber="41" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger (1880)</bibRefCitation>
described as variety of “
<taxonomicName id="9BEF3AD6FFDDFFF58959FD59FF720334" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="plicatula">
<emphasis id="6E9B9D47FFDDFFF58959FD59FF720334" italics="true" pageId="40" pageNumber="41">Mitra plicatula</emphasis>
</taxonomicName>
” (=
<taxonomicName id="9BEF3AD6FFDDFFF58D34FCB4FDA60334" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[305,559,794,820]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="transalpinum" status="sp. nov.">
<emphasis id="6E9B9D47FFDDFFF58D34FCB4FDA60334" box="[305,559,794,820]" italics="true" pageId="40" pageNumber="41">Vexillum transalpinum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFDDFFF58E32FCB4FD180334" box="[567,657,794,820]" pageId="40" pageNumber="41" rank="species">nov. sp.</taxonomicNameLabel>
). A close relationship between
<taxonomicName id="9BEF3AD6FFDDFFF58FF0FCB5FF500358" authority="(Hoernes &amp; Auinger, 1880)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="moravica">
<emphasis id="6E9B9D47FFDDFFF58FF0FCB5FB2E0334" box="[1013,1191,795,820]" italics="true" pageId="40" pageNumber="41">Pusia moravica</emphasis>
(
<bibRefCitation id="387E3CA4FFDDFFF588B2FCB4FF590358" author="Hoernes, R. &amp; Auinger, M." pageId="40" pageNumber="41" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="9BEF3AD6FFDDFFF58D0BFC91FE370358" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[270,446,830,856]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="transalpinum">
<emphasis id="6E9B9D47FFDDFFF58D0BFC91FE370358" box="[270,446,830,856]" italics="true" pageId="40" pageNumber="41">V. transalpinum</emphasis>
</taxonomicName>
, however, is unlikely in respect to the short spire whorls and broad, convex last whorl of
<taxonomicName id="9BEF3AD6FFDDFFF58C92FCCDFE94037C" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Hoernes &amp; Auinger" baseAuthorityYear="1880" box="[151,285,867,892]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="moravica">
<emphasis id="6E9B9D47FFDDFFF58C92FCCDFE94037C" box="[151,285,867,892]" italics="true" pageId="40" pageNumber="41">P. moravica</emphasis>
</taxonomicName>
. In addition, the subsutural spiral row of weak nodes and the presence of spiral grooves allows a clear separation from
<taxonomicName id="9BEF3AD6FFDDFFF58D4AFC29FE7603A0" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[335,511,902,928]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="transalpinum">
<emphasis id="6E9B9D47FFDDFFF58D4AFC29FE7603A0" box="[335,511,902,928]" italics="true" pageId="40" pageNumber="41">V. transalpinum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5C504155FFDDFFF58CC2FC07FF5E03E8" blockId="40.[151,1437,325,1072]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2FC07FE3F03C3" bold="true" box="[199,438,937,963]" pageId="40" pageNumber="41">Palaeoenvironment.</emphasis>
The specimens derive from basinal clay, suggesting a middle neritic depositional environment.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDDFFF58CC2FC5FFE450430" pageId="40" pageNumber="41" type="distribution">
<paragraph id="5C504155FFDDFFF58CC2FC5FFE450430" blockId="40.[151,1437,325,1072]" pageId="40" pageNumber="41">
<emphasis id="6E9B9D47FFDDFFF58CC2FC5FFDFA040C" bold="true" box="[199,627,1009,1036]" pageId="40" pageNumber="41">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFDDFFF58FD5FC5DFBF9040C" box="[976,1136,1011,1036]" italics="true" pageId="40" pageNumber="41">Vienna Basin:</emphasis>
Rudice (
<collectingCountry id="24F801C5FFDDFFF588D9FC5CFA1C040D" box="[1244,1429,1010,1037]" name="Czech Republic" pageId="40" pageNumber="41">Czech Republic</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFDDFFF58C9AFBB8FE360430" author="Hoernes, R. &amp; Auinger, M." box="[159,447,1046,1072]" pageId="40" pageNumber="41" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger 1880</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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