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<mods:title id="C8AE6E8E91501B3033D62C2E1E0DA40E">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="68D916BE136D7126F7BE773633F32AAE">Harzhauser, Mathias</mods:namePart>
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<treatment id="D446F043FFD8FFF38C92F95AFB660410" ID-DOI="http://doi.org/10.5281/zenodo.5043404" ID-GBIF-Taxon="183225104" ID-Zenodo-Dep="5043404" LSID="urn:lsid:plazi:treatment:D446F043FFD8FFF38C92F95AFB660410" httpUri="http://treatment.plazi.org/id/D446F043FFD8FFF38C92F95AFB660410" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<paragraph id="5C504155FFD8FFF08C92F95AFD2E070E" blockId="45.[151,679,1779,1848]" box="[151,679,1779,1806]" pageId="45" pageNumber="46">
<heading id="0718F639FFD8FFF08C92F95AFD2E070E" bold="true" box="[151,679,1779,1806]" fontSize="11" level="2" pageId="45" pageNumber="46" reason="4">
<emphasis id="6E9B9D47FFD8FFF08C92F95AFD2E070E" bold="true" box="[151,679,1779,1806]" pageId="45" pageNumber="46">
<taxonomicName id="9BEF3AD6FFD8FFF08C92F95AFD90070E" authority="(Friedberg, 1911)" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Friedberg" baseAuthorityYear="1911" box="[151,537,1779,1806]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="brevior" status="comb. nov.">
<emphasis id="6E9B9D47FFD8FFF08C92F95AFEB0070E" bold="true" box="[151,313,1779,1806]" italics="true" pageId="45" pageNumber="46">Pusia brevior</emphasis>
(
<bibRefCitation id="387E3CA4FFD8FFF08D4CF95AFD9B070E" author="Friedberg, W." box="[329,530,1780,1806]" pageId="45" pageNumber="46" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg, 1911</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFD8FFF08E24F95AFD2E070E" box="[545,679,1780,1806]" pageId="45" pageNumber="46" rank="species">nov. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="14F512DEFFD8FFF08C92F8B6FEA70738" box="[151,302,1816,1848]" pageId="45" pageNumber="46" type="description">
<paragraph id="5C504155FFD8FFF08C92F8B6FEA70738" blockId="45.[151,679,1779,1848]" box="[151,302,1816,1848]" pageId="45" pageNumber="46">
<figureCitation id="C4D45DD0FFD8FFF08C92F8B6FF740732" box="[151,253,1816,1842]" captionStart="FIGURE 18" captionStartId="52.[151,250,1435,1459]" captionTargetBox="[208,1379,776,1412]" captionTargetId="figure-253@52.[208,1379,776,1412]" captionTargetPageId="52" captionText="FIGURE 18 A. Tosapusia neudorfensis (Schaffer, 1898), reproduction of illustration of syntype by Schaffer (1898, fig. 5). B." figureDoi="http://doi.org/10.5281/zenodo.4921878" httpUri="https://zenodo.org/record/4921878/files/figure.png" pageId="45" pageNumber="46">Figs 18C</figureCitation>
<figureCitation id="C4D45DD0FFD8FFF08CF8F884FEAF0732" box="[253,294,1816,1848]" captionStart="FIGURE 1" captionStartId="5.[151,250,1852,1876]" captionTargetBox="[192,1396,181,1828]" captionTargetId="figure-17@5.[192,1396,181,1828]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, ESopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.4921844" httpUri="https://zenodo.org/record/4921844/files/figure.png" pageId="45" pageNumber="46">
<subScript id="C06B4310FFD8FFF08CF8F884FE8C0738" attach="left" box="[253,261,1834,1848]" fontSize="6" pageId="45" pageNumber="46">1</subScript>
C
</figureCitation>
<figureCitation id="C4D45DD0FFD8FFF08D23F884FEA70738" box="[294,302,1834,1848]" captionStart="FIGURE 2" captionStartId="6.[151,250,1599,1623]" captionTargetBox="[389,1196,417,1576]" captionTargetId="figure-124@6.[389,1196,417,1576]" captionTargetPageId="6" captionText="FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.4921846" httpUri="https://zenodo.org/record/4921846/files/figure.png" pageId="45" pageNumber="46">
<subScript id="C06B4310FFD8FFF08D23F884FEA70738" attach="left" box="[294,302,1834,1848]" fontSize="6" pageId="45" pageNumber="46">2</subScript>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD8FFF08C92F8F3FCD90793" pageId="45" pageNumber="46" type="reference_group">
<paragraph id="5C504155FFD8FFF08C92F8F3FC430775" blockId="45.[151,970,1884,1939]" box="[151,970,1884,1909]" pageId="45" pageNumber="46">
<treatmentCitationGroup id="7CFF667BFFD8FFF08C92F8F3FC430775" box="[151,970,1884,1909]" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08C92F8F3FF2D0774" box="[151,164,1885,1908]" italics="true" pageId="45" pageNumber="46">*</emphasis>
<taxonomicName id="9BEF3AD6FFD8FFF08CAFF8F2FDAF0774" authority="Bell" box="[170,550,1884,1908]" class="Gastropoda" family="Clavatulidae" genus="Turricula" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="variety" species="recticosta" variety="brevior">
<emphasis id="6E9B9D47FFD8FFF08CAFF8F2FEFA0774" box="[170,371,1884,1908]" italics="true" pageId="45" pageNumber="46">Turricula recticosta</emphasis>
Bell. var.
<emphasis id="6E9B9D47FFD8FFF08DD9F8F2FDAF0774" box="[476,550,1884,1908]" italics="true" pageId="45" pageNumber="46">brevior</emphasis>
</taxonomicName>
mihi.—
<treatmentCitation id="DD4E6744FFD8FFF08E7DF8F2FCB70775" author="Friedberg, W." box="[632,830,1884,1909]" page="24" pageId="45" pageNumber="46" year="1911">
<bibRefCitation id="387E3CA4FFD8FFF08E7DF8F2FCB70775" author="Friedberg, W." box="[632,830,1884,1909]" pageId="45" pageNumber="46" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg 1911: 24</bibRefCitation>
</treatmentCitation>
, pl. 1, fig. 17.
</treatmentCitationGroup>
</paragraph>
<paragraph id="5C504155FFD8FFF08C92F8D5FCD90793" blockId="45.[151,970,1884,1939]" box="[151,848,1915,1939]" pageId="45" pageNumber="46">
<treatmentCitationGroup id="7CFF667BFFD8FFF08C92F8D5FCD90793" box="[151,848,1915,1939]" pageId="45" pageNumber="46">
<taxonomicName id="9BEF3AD6FFD8FFF08C92F8D5FDE90793" authority="Friedb." authorityName="Friedb." box="[151,608,1915,1939]" class="Gastropoda" family="Clavatulidae" genus="Turricula" kingdom="Animalia" order="Neogastropoda" pageId="45" pageNumber="46" phylum="Mollusca" rank="variety" species="recticosta" variety="brevior">
<emphasis id="6E9B9D47FFD8FFF08C92F8D5FEE90793" box="[151,352,1915,1939]" italics="true" pageId="45" pageNumber="46">Turricula recticosta</emphasis>
Bell. var.
<emphasis id="6E9B9D47FFD8FFF08DCCF8D5FD9A0793" box="[457,531,1915,1939]" italics="true" pageId="45" pageNumber="46">brevior</emphasis>
Friedb.
</taxonomicName>
<treatmentCitation id="DD4E6744FFD8FFF08E7CF8D5FCC50793" author="Friedberg, W." box="[633,844,1915,1939]" page="132" pageId="45" pageNumber="46" year="1938">
<bibRefCitation id="387E3CA4FFD8FFF08E7CF8D5FCC50793" author="Friedberg, W." box="[633,844,1915,1939]" pageId="45" pageNumber="46" pagination="1 - 164" refId="ref56467" refString="Friedberg, W. (1938) Katalog mego zbioru mieczakow miocenskich Polski (Katalog meiner Sammlung der Miozanmollusken Polens). Memoires de l'Academie polonaise des sciences et des lettres, Classe des sciences mathematiques et naturelles. Serie B, Sciences naturelles, 12, 1 - 164." type="journal article" year="1938">Friedberg 1938: 132</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD8FFF38C92F86FFBA70284" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="5C504155FFD8FFF38C92F86FFB9500F9" blockId="45.[151,1436,1985,2012]" lastBlockId="46.[151,1437,150,1041]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<emphasis id="6E9B9D47FFD8FFF08C92F86FFECC07DB" bold="true" box="[151,325,1985,2011]" pageId="45" pageNumber="46">Type material.</emphasis>
<typeStatus id="8354FFF7FFD8FFF08D49F86CFE3507DC" box="[332,444,1986,2012]" pageId="45" pageNumber="46" type="lectotype">Lectotype</typeStatus>
(designated herein): SL. 12.0 mm, MD: 5.0 mm, Hołubica (
<collectingCountry id="24F801C5FFD8FFF0885AF86FFB3607DC" box="[1119,1215,1985,2012]" name="Ukraine" pageId="45" pageNumber="46">Ukraine</collectingCountry>
), stored in the State Museum of Natural History of the National Academy of Sciences of
<collectingCountry id="24F801C5FFDBFFF38F89FF38FC6F00B0" box="[908,998,150,176]" name="Ukraine" pageId="46" pageNumber="47">Ukraine</collectingCountry>
in
<collectingRegion id="9E2B8FB7FFDBFFF3880DFF38FBB400B0" box="[1032,1085,150,176]" country="Ukraine" name="L'viv" pageId="46" pageNumber="47">Lviv</collectingRegion>
(formerly Muzeum Dzieduszyckich), illustrated in
<bibRefCitation id="387E3CA4FFDBFFF38D85FF14FDE800D4" author="Friedberg, W." box="[384,609,186,212]" pageId="46" pageNumber="47" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg (1911: 24</bibRefCitation>
, pl. 1, fig. 17). Unfortunately, it was not possible to obtain pictures of the
<typeStatus id="8354FFF7FFDBFFF38C92FF70FF7700F8" box="[151,254,222,248]" pageId="46" pageNumber="47" type="lectotype">lectotype</typeStatus>
and we reproduce the illustration from
<bibRefCitation id="387E3CA4FFDBFFF38EBFFF70FCF000F8" author="Friedberg, W." box="[698,889,222,248]" pageId="46" pageNumber="47" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg (1911)</bibRefCitation>
, figs 18C
<subScript id="C06B4310FFDBFFF38FE1FF5FFC6500FF" attach="left" box="[996,1004,241,255]" fontSize="6" pageId="46" pageNumber="47">1</subScript>
C
<subScript id="C06B4310FFDBFFF38808FF5FFB9C00FF" attach="left" box="[1037,1045,241,255]" fontSize="6" pageId="46" pageNumber="47">2</subScript>
.
</paragraph>
<paragraph id="5C504155FFDBFFF38CC2FEACFECA0260" blockId="46.[151,1437,150,1041]" pageId="46" pageNumber="47">
<emphasis id="6E9B9D47FFDBFFF38CC2FEACFE3A011C" bold="true" box="[199,435,258,284]" pageId="46" pageNumber="47">Revised description</emphasis>
(based on description and illustration in
<bibRefCitation id="387E3CA4FFDBFFF38F8CFEACFBB7011C" author="Friedberg, W." box="[905,1086,258,284]" pageId="46" pageNumber="47" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg, 1911</bibRefCitation>
). Shell moderately large, stout fusiform to biconical. Protoconch multispiral, high conical, mammillate, of three smooth, weakly convex, whorls. Teleoconch boundary marked by beginning of axial sculpture. Teleoconch of five convex whorls, with periphery slightly below mid-whorl, separated by incised suture. Axial sculpture of prominent, close-set axial ribs, separated by narrower interspaces (1620 axial ribs on penultimate whorl), crossed by delicate spiral threads, most prominent in axial interspaces. Slightly deeper spiral groove separates nodulose subsutural spiral cord. Last whorl evenly convex, strongly constricted. Aperture narrow, elongate. Columellar callus narrow, indistinct. Columella with four folds, abapically decreasing in strength. Outer lip thin, prominent lirae inside aperture. Four prominent spiral cords on fasciole, partly from extensions of columellar folds. Siphonal canal moderately long, narrow, straight, with deep siphonal notch.
</paragraph>
<paragraph id="5C504155FFDBFFF38CC2FDC4FBA70284" blockId="46.[151,1437,150,1041]" box="[199,1070,618,645]" pageId="46" pageNumber="47">
<emphasis id="6E9B9D47FFDBFFF38CC2FDC4FDB50284" bold="true" box="[199,572,618,644]" pageId="46" pageNumber="47">Shell measurements and ratios.</emphasis>
SL:
<quantity id="9B17ECB0FFDBFFF38E77FDC4FD640285" box="[626,749,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="46" pageNumber="47" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
, MD: 4.25.0 mm; AA: 40°.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDBFFF38CC2FD20FDBA03C8" pageId="46" pageNumber="47" type="discussion">
<paragraph id="5C504155FFDBFFF38CC2FD20FEEA03A4" blockId="46.[151,1437,150,1041]" pageId="46" pageNumber="47">
<emphasis id="6E9B9D47FFDBFFF38CC2FD20FEC502A8" bold="true" box="[199,332,654,680]" pageId="46" pageNumber="47">Discussion.</emphasis>
This species is characterised by its dense axial/spiral sculpture. At first sight, the illustration in
<bibRefCitation id="387E3CA4FFDBFFF38C92FD1CFEDD02CC" author="Friedberg, W." box="[151,340,690,716]" pageId="46" pageNumber="47" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg (1911)</bibRefCitation>
is reminiscent of an
<taxonomicName id="9BEF3AD6FFDBFFF38E3AFD1CFC8E02CC" box="[575,775,690,716]" class="Gastropoda" family="Costellariidae" genus="Antithala" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="6E9B9D47FFDBFFF38E3AFD1CFD2E02CC" box="[575,679,690,716]" italics="true" pageId="46" pageNumber="47">Antithala</emphasis>
species.
</taxonomicName>
The thin outer lip, the wider aperture with lirae inside, and the presence of delicate spiral cords in the axial interspaces exclude a closer relationship with that genus.
<taxonomicName id="9BEF3AD6FFDBFFF38959FD79FF610314" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Friedberg" baseAuthorityYear="1911" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="brevior">
<emphasis id="6E9B9D47FFDBFFF38959FD79FF610314" italics="true" pageId="46" pageNumber="47">Pusia brevior</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="387E3CA4FFDBFFF38DD3FD54FD1F0314" author="Friedberg, W." box="[470,662,762,788]" pageId="46" pageNumber="47" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg (1911)</bibRefCitation>
as variety of
<taxonomicName id="9BEF3AD6FFDBFFF38F39FD55FB2D0315" authority="Bellardi, 1850" authorityName="Bellardi" authorityYear="1850" box="[828,1188,762,789]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="recticosta">
<emphasis id="6E9B9D47FFDBFFF38F39FD55FC7A0314" box="[828,1011,763,788]" italics="true" pageId="46" pageNumber="47">Mitra recticosta</emphasis>
<bibRefCitation id="387E3CA4FFDBFFF38FF8FD54FB2D0315" author="Bellardi, L." box="[1021,1188,762,789]" pageId="46" pageNumber="47" pagination="357 - 390" refId="ref54310" refString="Bellardi, L. (1850) Monografia delle mitre fossili del Piemonte. Memorie della Reale Accademia delle Scienze di Torino, 2 / 11, 357 - 390. [https: // www. biodiversitylibrary. org / item / 32672 # page / 441 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 59630" type="journal article" year="1850">Bellardi, 1850</bibRefCitation>
</taxonomicName>
, but differs from this Pliocene Italian species distinctly by its squatter shape, the presence of spiral sculpture, the smaller size, and wider apical angle. The absence of spiral sculpture distinguishes also
<taxonomicName id="9BEF3AD6FFDBFFF38F76FCEDFBAC035C" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[883,1061,834,860]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="confunda" status="sp. nov.">
<emphasis id="6E9B9D47FFDBFFF38F76FCEDFBAC035C" box="[883,1061,834,860]" italics="true" pageId="46" pageNumber="47">Pusia confunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFDBFFF3882BFCEDFB00035D" box="[1070,1161,835,861]" pageId="46" pageNumber="47" rank="species">nov. sp.</taxonomicNameLabel>
, from
<taxonomicName id="9BEF3AD6FFDBFFF388DDFCEDFACE035C" authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Friedberg" baseAuthorityYear="1911" box="[1240,1351,834,860]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="brevior">
<emphasis id="6E9B9D47FFDBFFF388DDFCEDFACE035C" box="[1240,1351,834,860]" italics="true" pageId="46" pageNumber="47">P. brevior</emphasis>
</taxonomicName>
, which develops a similar shape.
<taxonomicName id="9BEF3AD6FFDBFFF38DB2FCC9FCA80380" authority="Meznerics, 1933" authorityName="Meznerics" authorityYear="1933" box="[439,801,870,896]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="schafferi">
<emphasis id="6E9B9D47FFDBFFF38DB2FCC9FDE90380" box="[439,608,870,896]" italics="true" pageId="46" pageNumber="47">Pusia schafferi</emphasis>
<bibRefCitation id="387E3CA4FFDBFFF38E62FCC8FCA80380" author="Meznerics, I." box="[615,801,870,896]" pageId="46" pageNumber="47" pagination="319 - 359" refId="ref59495" refString="Meznerics, I. (1933) Die Minutien der tortonischen Ablagerungen von Steinabrunn in Niederosterreich. Annalen des Naturhistorischen Museums Wien, 46, 319 - 359. [https: // www. zobodat. at / pdf / ANNA _ 46 _ 0319 - 0359. pdf]" type="journal article" year="1933">Meznerics, 1933</bibRefCitation>
</taxonomicName>
has comparable sculpture, but is much more slender and has a higher spire.
</paragraph>
<paragraph id="5C504155FFDBFFF38CC2FC00FDBA03C8" blockId="46.[151,1437,150,1041]" box="[199,563,942,968]" pageId="46" pageNumber="47">
<emphasis id="6E9B9D47FFDBFFF38CC2FC00FE3F03C8" bold="true" box="[199,438,942,968]" pageId="46" pageNumber="47">Palaeoenvironment.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFDBFFF38CC2FC7CFB660410" pageId="46" pageNumber="47" type="distribution">
<paragraph id="5C504155FFDBFFF38CC2FC7CFB660410" blockId="46.[151,1437,150,1041]" pageId="46" pageNumber="47">
<emphasis id="6E9B9D47FFDBFFF38CC2FC7CFDE103EC" bold="true" box="[199,616,978,1004]" pageId="46" pageNumber="47">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis id="6E9B9D47FFDBFFF38FB3FC7CFA8303EC" box="[950,1290,978,1004]" italics="true" pageId="46" pageNumber="47">Polish Fore-Carpathian Basin</emphasis>
: Borki Wielkie (
<collectingCountry id="24F801C5FFDBFFF38CCDFC58FE900411" box="[200,281,1014,1041]" name="Poland" pageId="46" pageNumber="47">Poland</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFDBFFF38D2AFC58FE560410" author="Friedberg, W." box="[303,479,1014,1040]" pageId="46" pageNumber="47" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg 1911</bibRefCitation>
);
<emphasis id="6E9B9D47FFDBFFF38DF1FC58FD2F0410" box="[500,678,1014,1040]" italics="true" pageId="46" pageNumber="47">Voronyaky Hills</emphasis>
: Hołubica (Holubytsia) (
<collectingCountry id="24F801C5FFDBFFF38FBAFC58FB940411" box="[959,1053,1014,1041]" name="Ukraine" pageId="46" pageNumber="47">Ukraine</collectingCountry>
) (
<bibRefCitation id="387E3CA4FFDBFFF38837FC58FB6A0410" author="Friedberg, W." box="[1074,1251,1014,1040]" pageId="46" pageNumber="47" pagination="1 - 112" refId="ref56356" refString="Friedberg, W. (1911, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie, 1 &amp; 5, 1 - 112, pls. 1 - 5 (1911) &amp; 441 - 631, pls. 27 - 38 (1928). [reprinted 1951 - 55 with slightly different title and pagination, Wydawnictwa Geologiczne, Warszawa, http: // rcin. org. pl / miiz / dlibra / publication / 4792 / edition / 29890 / content]" type="journal article" year="1911">Friedberg 1911</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>