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<mods:title id="36EF6449B11D4DD3643332547517BD5E">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="50577F9B1EEC8D5977E3B6DFC6373370">Harzhauser, Mathias</mods:namePart>
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<treatment id="D446F043FFD4FFFC8C92FD78FB700770" ID-DOI="http://doi.org/10.5281/zenodo.5043364" ID-GBIF-Taxon="183225138" ID-Zenodo-Dep="5043364" LSID="urn:lsid:plazi:treatment:D446F043FFD4FFFC8C92FD78FB700770" httpUri="http://treatment.plazi.org/id/D446F043FFD4FFFC8C92FD78FB700770" lastPageNumber="34" pageId="33" pageNumber="34">
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<paragraph id="5C504155FFD4FFFC8C92FD78FE5602F0" blockId="33.[151,479,726,753]" box="[151,479,726,753]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8C92FD78FE5602F0" bold="true" box="[151,479,726,753]" pageId="33" pageNumber="34">
Genus
<taxonomicName id="9BEF3AD6FFD4FFFC8CEBFD78FEE302F1" ID-CoL="8KS5R" authority="Harzhauser &amp; Landau, 2021" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[238,362,726,753]" class="Gastropoda" family="Costellariidae" genus="Fedosovia" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="6E9B9D47FFD4FFFC8CEBFD78FEE302F1" bold="true" box="[238,362,726,753]" italics="true" pageId="33" pageNumber="34">Fedosovia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="75A8203CFFD4FFFC8D77FD78FE5602F0" box="[370,479,726,752]" pageId="33" pageNumber="34" rank="genus">nov. gen.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="14F512DEFFD4FFFC8C92FCB0FAD70338" box="[151,1374,798,825]" pageId="33" pageNumber="34" type="type_taxon">
<paragraph id="5C504155FFD4FFFC8C92FCB0FAD70338" blockId="33.[151,1437,798,1905]" box="[151,1374,798,825]" pageId="33" pageNumber="34">
<materialsCitation id="EC874B08FFD4FFFC8C92FCB0FAD30338" ID-GBIF-Occurrence="3311814426" box="[151,1370,798,825]" country="Romania" location="Miocene" municipality="Paratethys Sea" pageId="33" pageNumber="34" specimenCount="1" typeStatus="holotype">
<emphasis id="6E9B9D47FFD4FFFC8C92FCB0FEBB0338" bold="true" box="[151,306,798,824]" pageId="33" pageNumber="34">
<typeStatus id="8354FFF7FFD4FFFC8C92FCB0FF5B0338" box="[151,210,798,824]" pageId="33" pageNumber="34">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="9BEF3AD6FFD4FFFC8D3CFCB1FCE60338" authority="Hoernes &amp; Auinger, 1880" authorityName="Hoernes &amp; Auinger" authorityYear="1880" box="[313,879,798,825]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="fuchsi" subGenus="Callithea">
<emphasis id="6E9B9D47FFD4FFFC8D3CFCB1FEF00338" box="[313,377,799,824]" italics="true" pageId="33" pageNumber="34">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFD4FFFC8D8CFCB0FE780338" box="[393,497,798,824]" italics="true" pageId="33" pageNumber="34">Callithea</emphasis>
)
<emphasis id="6E9B9D47FFD4FFFC8E07FCB0FDCC0338" box="[514,581,798,824]" italics="true" pageId="33" pageNumber="34">fuchsi</emphasis>
<bibRefCitation id="387E3CA4FFD4FFFC8E49FCB1FCE60338" author="Hoernes, R. &amp; Auinger, M." box="[588,879,798,825]" pageId="33" pageNumber="34" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
</taxonomicName>
, middle
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,
<collectingMunicipality id="BC34DB2FFFD4FFFC883AFCB0FB6C0338" box="[1087,1253,798,824]" pageId="33" pageNumber="34">Paratethys Sea</collectingMunicipality>
,
<collectingCountry id="24F801C5FFD4FFFC88F4FCB0FAD30338" box="[1265,1370,798,824]" name="Romania" pageId="33" pageNumber="34">Romania</collectingCountry>
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.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD4FFFC8CC2FCECFACC0434" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="5C504155FFD4FFFC8CC2FCECFACC0434" blockId="33.[151,1437,798,1905]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8CC2FCECFECB035C" bold="true" box="[199,322,834,860]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
Large, moderately broad fusiform shell, with beaded sculpture, differing from
<taxonomicName id="9BEF3AD6FFD4FFFC88ABFCECFB66035C" authorityName="H. Adams &amp; A. Adams" authorityYear="1853" box="[1198,1263,834,860]" class="Gastropoda" family="Costellariidae" genus="Thala" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC88ABFCECFB66035C" box="[1198,1263,834,860]" italics="true" pageId="33" pageNumber="34">Thala</emphasis>
</taxonomicName>
in the development of a parietal denticle and the less cyrtoconoid spire. Protoconch paucispiral, of weakly convex, smooth whorl. Teleoconch whorls with dense pattern of axial ribs and spiral cords forming densely spaced pillow-like, subquadrate nodes at intersections. Axial ribs of early teleoconch whorls of
<typeStatus id="8354FFF7FFD4FFFC8F8AFC01FC3603C9" box="[911,959,943,969]" pageId="33" pageNumber="34">type</typeStatus>
species bearing three spiral rows of nodes with tiny spines on middle spiral row [not preserved in
<taxonomicName id="9BEF3AD6FFD4FFFC8EFAFC7DFCFA03EC" baseAuthorityName="Toldo" baseAuthorityYear="1889" box="[767,883,978,1004]" class="Gastropoda" family="Costellariidae" genus="Fedosovia" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="zibinica">
<emphasis id="6E9B9D47FFD4FFFC8EFAFC7DFCFA03EC" box="[767,883,978,1004]" italics="true" pageId="33" pageNumber="34">F. zibinica</emphasis>
</taxonomicName>
]. Aperture moderately narrow to wide, with three columellar folds; adapical two folds very prominent. Columellar callus narrow, abapically contracting, no lirae inside aperture. Siphonal canal moderately long, straight or slightly bent to left. No signs of episodic growth.
</paragraph>
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<paragraph id="5C504155FFD4FFFC8CC2FB90FDB50458" blockId="33.[151,1437,798,1905]" box="[199,572,1086,1113]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8CC2FB90FED00458" bold="true" box="[199,345,1086,1112]" pageId="33" pageNumber="34">Description.</emphasis>
As for
<typeStatus id="8354FFF7FFD4FFFC8DA9FB91FE550459" box="[428,476,1087,1113]" pageId="33" pageNumber="34">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD4FFFC8CC2FBCCFDB204E8" pageId="33" pageNumber="34" type="etymology">
<paragraph id="5C504155FFD4FFFC8CC2FBCCFDE604A0" blockId="33.[151,1437,798,1905]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8CC2FBCCFEC7047C" bold="true" box="[199,334,1122,1148]" pageId="33" pageNumber="34">Etymology.</emphasis>
In honour of Alexander Fedosov (Russian Academy of Sciences,
<collectingRegion id="9E2B8FB7FFD4FFFC884CFBCDFB27047D" box="[1097,1198,1123,1149]" country="Russia" name="Moscow" pageId="33" pageNumber="34">Moscow</collectingRegion>
), in gratitude for the discussions on Paratethyan
<taxonomicName id="9BEF3AD6FFD4FFFC8DC9FB28FDE204A0" authorityName="MacDonald" authorityYear="1860" box="[460,619,1158,1184]" class="Gastropoda" family="Costellariidae" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="family">Costellariidae</taxonomicName>
.
</paragraph>
<paragraph id="5C504155FFD4FFFC8CC2FB04FDB204E8" blockId="33.[151,1437,798,1905]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8CC2FB04FE1A04C4" bold="true" box="[199,403,1194,1220]" pageId="33" pageNumber="34">Included species.</emphasis>
<taxonomicName id="9BEF3AD6FFD4FFFC8D9EFB05FC5004C4" authority="Hoernes &amp; Auinger, 1880" authorityName="Hoernes &amp; Auinger" authorityYear="1880" box="[411,985,1194,1221]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="fuchsi" subGenus="Callithea">
<emphasis id="6E9B9D47FFD4FFFC8D9EFB05FE5204C4" box="[411,475,1195,1220]" italics="true" pageId="33" pageNumber="34">Mitra</emphasis>
(
<emphasis id="6E9B9D47FFD4FFFC8DE8FB04FDDC04C4" box="[493,597,1194,1220]" italics="true" pageId="33" pageNumber="34">Callithea</emphasis>
)
<emphasis id="6E9B9D47FFD4FFFC8E62FB04FD2304C4" box="[615,682,1194,1220]" italics="true" pageId="33" pageNumber="34">fuchsi</emphasis>
<bibRefCitation id="387E3CA4FFD4FFFC8EB7FB05FC5004C4" author="Hoernes, R. &amp; Auinger, M." box="[690,985,1194,1221]" pageId="33" pageNumber="34" pagination="53 - 112" refId="ref57816" refString="Hoernes, R. &amp; Auinger, M. (1880) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der k. k Geologischen Reichsanstalt, 12, 53 - 112, pls. 7 - 12. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf] https: // doi. org / 10.5962 / bhl. title. 151405" type="journal article" year="1880">Hoernes &amp; Auinger, 1880</bibRefCitation>
</taxonomicName>
, middle Miocene,
<collectingCountry id="24F801C5FFD4FFFC88A8FB04FA9F04C4" box="[1197,1302,1194,1220]" name="Romania" pageId="33" pageNumber="34">Romania</collectingCountry>
;
<taxonomicName id="9BEF3AD6FFD4FFFC8927FB05FED104E8" authority="Toldo, 1889" authorityName="Toldo" authorityYear="1889" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="zibinica">
<emphasis id="6E9B9D47FFD4FFFC8927FB05FF4E04E8" italics="true" pageId="33" pageNumber="34">Mitra zibinica</emphasis>
<bibRefCitation id="387E3CA4FFD4FFFC8CCBFB60FED104E8" author="Toldo, G." box="[206,344,1230,1256]" pageId="33" pageNumber="34" pagination="144 - 150" refId="ref62853" refString="Toldo, G. (1889) Mitridae del Miocene superiore di Montegibbio. Bollettino della Societa Malacologica Italiana, 14, 144 - 150. [https: // www. biodiversitylibrary. org / item / 121430 # page / 148 / mode / 1 up]" type="journal article" year="1889">Toldo, 1889</bibRefCitation>
</taxonomicName>
, late Miocene,
<collectingCountry id="24F801C5FFD4FFFC8E07FB60FDBE04E8" box="[514,567,1230,1256]" name="Italy" pageId="33" pageNumber="34">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD4FFFC8CC2FB5CFDF50579" pageId="33" pageNumber="34" type="description">
<paragraph id="5C504155FFD4FFFC8CC2FB5CFD1F0530" blockId="33.[151,1437,798,1905]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8CC2FB5CFDF0050C" bold="true" box="[199,633,1266,1292]" pageId="33" pageNumber="34">Stratigraphic and geographic range.</emphasis>
Middle Miocene (Badenian) deposits of the Central Paratethys Sea and late Miocene of the proto-Mediterranean Sea.
</paragraph>
<paragraph id="5C504155FFD4FFFC8CC2FA94FDF50579" blockId="33.[151,1437,798,1905]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8CC2FA94FE3F0554" bold="true" box="[199,438,1338,1364]" pageId="33" pageNumber="34">Palaeoenvironment.</emphasis>
Unknown; most probably from middle neritic depositional environments with pelitic bottom (based on sediment from the aperture).
</paragraph>
</subSubSection>
<subSubSection id="14F512DEFFD4FFFC8CC2FA2CFB700770" pageId="33" pageNumber="34" type="discussion">
<paragraph id="5C504155FFD4FFFC8CC2FA2CFB490651" blockId="33.[151,1437,798,1905]" pageId="33" pageNumber="34">
<emphasis id="6E9B9D47FFD4FFFC8CC2FA2CFEC5059C" bold="true" box="[199,332,1410,1436]" pageId="33" pageNumber="34">Discussion.</emphasis>
<taxonomicName id="9BEF3AD6FFD4FFFC8D52FA2CFE45059C" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[343,460,1410,1436]" class="Gastropoda" family="Costellariidae" genus="Fedosovia" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8D52FA2CFE45059C" box="[343,460,1410,1436]" italics="true" pageId="33" pageNumber="34">Fedosovia</emphasis>
</taxonomicName>
differs from other costellariid genera by the combination of a peculiar pillow-like sculpture and a paucispiral protoconch (as opposed to
<taxonomicName id="9BEF3AD6FFD4FFFC8F06FA08FC0E05C0" authorityName="Monterosato" authorityYear="1917" box="[771,903,1446,1472]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8F06FA08FC0E05C0" box="[771,903,1446,1472]" italics="true" pageId="33" pageNumber="34">Ebenomitra</emphasis>
</taxonomicName>
, which has a paucispiral protoconch, but lacks the peculiar sculpture). Protoconch of paucispiral or multispiral
<typeStatus id="8354FFF7FFD4FFFC8F59FA65FC0505E5" box="[860,908,1483,1509]" pageId="33" pageNumber="34">type</typeStatus>
is not a useful distinguishing character in many gastropod groups. However, amongst the
<taxonomicName id="9BEF3AD6FFD4FFFC8E6AFA40FB840608" authority=", Fedosov et al. (2017)" authorityName="Fedosov" authorityYear="2017" box="[623,1037,1518,1545]" class="Gastropoda" family="Costellariidae" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="family">
Costellariidae,
<bibRefCitation id="387E3CA4FFD4FFFC8F1FFA40FB840608" author="Fedosov, A. &amp; Puillandre, N. &amp; Herrmann, M. &amp; Dgebuadze, P. &amp; Bouchet, P." box="[794,1037,1518,1545]" pageId="33" pageNumber="34" pagination="541 - 626" refId="ref56065" refString="Fedosov, A. E, Puillandre, N., Herrmann, M., Dgebuadze, P. &amp; Bouchet, P. (2017) Phylogeny, systematics, and evolution of the family Costellariidae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 179, 541 - 626. https: // doi. org / 10.1111 / zoj. 12431" type="journal article" year="2017">
Fedosov
<emphasis id="6E9B9D47FFD4FFFC8F84FA41FC3A0608" box="[897,947,1518,1544]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
. (2017)
</bibRefCitation>
</taxonomicName>
placed importance at supraspecific level to protoconch
<typeStatus id="8354FFF7FFD4FFFC8D72F9BDFE2E062D" box="[375,423,1555,1581]" pageId="33" pageNumber="34">type</typeStatus>
in many of their genera, and specified the protoconch
<typeStatus id="8354FFF7FFD4FFFC8808F9BDFBB4062D" box="[1037,1085,1555,1581]" pageId="33" pageNumber="34">type</typeStatus>
in his generic descriptions. We follow this lead here and consider the protoconch
<typeStatus id="8354FFF7FFD4FFFC8EC2F999FD7E0651" box="[711,759,1591,1617]" pageId="33" pageNumber="34">type</typeStatus>
a genus-specific character in
<taxonomicName id="9BEF3AD6FFD4FFFC8840F998FB330650" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[1093,1210,1590,1616]" class="Gastropoda" family="Costellariidae" genus="Fedosovia" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8840F998FB330650" box="[1093,1210,1590,1616]" italics="true" pageId="33" pageNumber="34">Fedosovia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5C504155FFD4FFFC8CC2F9F4FB700770" blockId="33.[151,1437,798,1905]" pageId="33" pageNumber="34">
The paucispiral protoconch excludes a relation with
<taxonomicName id="9BEF3AD6FFD4FFFC8F2EF9F4FC020674" authorityName="Roding" authorityYear="1798" box="[811,907,1626,1652]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8F2EF9F4FC020674" box="[811,907,1626,1652]" italics="true" pageId="33" pageNumber="34">Vexillum</emphasis>
</taxonomicName>
, although the extant
<taxonomicName id="9BEF3AD6FFD4FFFC8883F9F4FF520699" authority="(Pease, 1860)" baseAuthorityName="Pease" baseAuthorityYear="1860" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="piceum">
<emphasis id="6E9B9D47FFD4FFFC8883F9F4FACB0674" box="[1158,1346,1626,1652]" italics="true" pageId="33" pageNumber="34">Vexillum piceum</emphasis>
(
<bibRefCitation id="387E3CA4FFD4FFFC8951F9F4FF5A0699" author="Pease, W. H." pageId="33" pageNumber="34" pagination="141 - 148" refId="ref60273" refString="Pease, W. H. (1860) Descriptions of new species of Mollusca from the Sandwich Islands, part II. Proceedings of the Zoological Society of London, 28, 141 - 148. [https: // www. biodiversitylibrary. org / item / 46225 # page / 197 / mode / 1 up]" type="journal article" year="1860">Pease, 1860</bibRefCitation>
)
</taxonomicName>
, from
<collectingCountry id="24F801C5FFD4FFFC8D2CF9D0FE010698" box="[297,392,1662,1688]" name="Réunion" pageId="33" pageNumber="34">Réunion</collectingCountry>
and Hawaii, develops a comparable sculpture (but is much smaller, SH &lt;
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). A paucispiral protoconch occurs in the extant
<taxonomicName id="9BEF3AD6FFD4FFFC8E5FF90CFCE806BD" authority="Huang, 2015" authorityName="Huang" authorityYear="2015" box="[602,865,1698,1725]" class="Gastropoda" family="Costellariidae" genus="Atlantilux" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8E5FF90CFD4106BC" box="[602,712,1698,1724]" italics="true" pageId="33" pageNumber="34">Atlantilux</emphasis>
<bibRefCitation id="387E3CA4FFD4FFFC8ED5F90DFCE806BD" author="Huang, S. I." box="[720,865,1698,1725]" pageId="33" pageNumber="34" pagination="43 - 53" refId="ref58305" refString="Huang, S. I. (2015) New Costellariidae (Mollusca: Gastropoda) from Taiwan and the North Atlantic Ocean. Visaya, 4, 43 - 53." type="journal article" year="2015">Huang, 2015</bibRefCitation>
</taxonomicName>
but
<taxonomicName id="9BEF3AD6FFD4FFFC8F91F90CFBD206BC" box="[916,1115,1698,1724]" class="Gastropoda" family="Costellariidae" genus="Atlantilux" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="6E9B9D47FFD4FFFC8F91F90CFB8B06BC" box="[916,1026,1698,1724]" italics="true" pageId="33" pageNumber="34">Atlantilux</emphasis>
species
</taxonomicName>
are characterised by a wide, slightly concave subsutural ramp and convex periphery close to the abapical suture.
<taxonomicName id="9BEF3AD6FFD4FFFC8841F968FB3B06E0" authorityName="Huang" authorityYear="2015" box="[1092,1202,1734,1760]" class="Gastropoda" family="Costellariidae" genus="Atlantilux" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8841F968FB3B06E0" box="[1092,1202,1734,1760]" italics="true" pageId="33" pageNumber="34">Atlantilux</emphasis>
</taxonomicName>
is also distinguished from
<taxonomicName id="9BEF3AD6FFD4FFFC8CD3F944FEC20704" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[214,331,1770,1796]" class="Gastropoda" family="Costellariidae" genus="Fedosovia" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8CD3F944FEC20704" box="[214,331,1770,1796]" italics="true" pageId="33" pageNumber="34">Fedosovia</emphasis>
</taxonomicName>
by the sculpture of dense axial riblets in the subsutural area opposed by wide fold-like axial ribs along the periphery (see
<bibRefCitation id="387E3CA4FFD4FFFC8DB7F8A0FD290728" author="Fedosov, A. &amp; Puillandre, N. &amp; Herrmann, M. &amp; Dgebuadze, P. &amp; Bouchet, P." box="[434,672,1806,1832]" pageId="33" pageNumber="34" pagination="541 - 626" refId="ref56065" refString="Fedosov, A. E, Puillandre, N., Herrmann, M., Dgebuadze, P. &amp; Bouchet, P. (2017) Phylogeny, systematics, and evolution of the family Costellariidae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 179, 541 - 626. https: // doi. org / 10.1111 / zoj. 12431" type="journal article" year="2017">
Fedosov
<emphasis id="6E9B9D47FFD4FFFC8E1FF8A1FDD30728" box="[538,602,1806,1832]" italics="true" pageId="33" pageNumber="34">et al.,</emphasis>
2017
</bibRefCitation>
).
<taxonomicName id="9BEF3AD6FFD4FFFC8EB1F8A0FCA00728" authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[692,809,1806,1832]" class="Gastropoda" family="Costellariidae" genus="Fedosovia" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8EB1F8A0FCA00728" box="[692,809,1806,1832]" italics="true" pageId="33" pageNumber="34">Fedosovia</emphasis>
</taxonomicName>
is superficially reminiscent of
<taxonomicName id="9BEF3AD6FFD4FFFC8889F8A1FE2D074C" authority="Fedosov, Herrmann &amp; Bouchet, 2017" authorityName="Fedosov, Herrmann &amp; Bouchet" authorityYear="2017" class="Gastropoda" family="Costellariidae" genus="Costapex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="6E9B9D47FFD4FFFC8889F8A1FB7D0729" box="[1164,1268,1807,1833]" italics="true" pageId="33" pageNumber="34">Costapex</emphasis>
<bibRefCitation id="387E3CA4FFD4FFFC88F8F8A0FE2D074C" author="Fedosov, A. &amp; Puillandre, N. &amp; Herrmann, M. &amp; Dgebuadze, P. &amp; Bouchet, P." pageId="33" pageNumber="34" pagination="541 - 626" refId="ref56065" refString="Fedosov, A. E, Puillandre, N., Herrmann, M., Dgebuadze, P. &amp; Bouchet, P. (2017) Phylogeny, systematics, and evolution of the family Costellariidae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 179, 541 - 626. https: // doi. org / 10.1111 / zoj. 12431" type="journal article" year="2017">
Fedosov, Herrmann &amp; Bouchet,
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in shape and sculpture. A closer relationship with this extant genus, however, is unlikely in respect to the paucispiral protoconch and the presence of only three columellar folds instead of four.
</paragraph>
</subSubSection>
</treatment>
</document>