230 lines
26 KiB
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230 lines
26 KiB
XML
<document id="868A3E668EC95B47C6D2A32FDCC39959" ID-CLB-Dataset="5925" ID-DOI="10.11646/zootaxa.4982.1.1" ID-GBIF-Dataset="f68b381d-d0dd-44e2-b9fe-6cf3c1aab1fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921842" ID-ZooBank="58388FB8-128A-4381-83D1-3C508D0D3873" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623285106829" checkinUser="plazi" docAuthor="Harzhauser, Mathias & Landau, Bernard" docDate="2021" docId="D446F043FFCDFFE58C92FF39FA1404FE" docLanguage="en" docName="zootaxa.4982.1.1.pdf" docOrigin="Zootaxa 4982 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tosapusia Habe 1964" docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="287F883BFFF5FFDD8C05FFAEFF890000" masterDocTitle="An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea" masterLastPageNumber="70" masterPageNumber="1" pageNumber="57" updateTime="1699009793899" updateUser="ExternalLinkService">
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<mods:mods id="B08D64FAF28F6B0B7E3DA3737F666D19" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="08248608BF64D76D1CB31874A5BCDC5E">
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<mods:title id="07C77F6E34B74487FF5CE66B93272A7F">An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:name id="0BF504F9B32C6807EDD915244277B044" type="personal">
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<mods:roleTerm id="A7B0D84077EA643ED3E42DD1FD1F6534">Author</mods:roleTerm>
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<mods:namePart id="38ADCF96EFDBDCAAB152F3BB5A6723DB">Harzhauser, Mathias</mods:namePart>
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<mods:nameIdentifier id="E63872352FE198F8272EC3B9BEDBB544" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
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<mods:nameIdentifier id="BA74A4F57FD7343D1D64C1B2950795BE" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:name id="583FC37CBB83F0DE9540076D6FE3A00E" type="personal">
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<mods:namePart id="3034ACBCCE15A339A0B8F1591539016B">Landau, Bernard</mods:namePart>
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<mods:nameIdentifier id="8BE30EF23E021AF944BB7E04426EE099" type="ORCID">0000-0002-4471-6655</mods:nameIdentifier>
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<mods:nameIdentifier id="693FB0BAFBC312524CD3D1AA18B7C44B" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:typeOfResource id="1884FCBB5294260E61AB084C9053E3DB">text</mods:typeOfResource>
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<mods:titleInfo id="E95CF5FCAEEFFF6F818843425475372C">
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<mods:title id="61D95ACD79A2C882B091D103C22B1B6F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="52D1A686F469D37AAF9ED3FAB781E96D">
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<mods:date id="1911EFF6C1386FD440C7E932766AF7FB">2021</mods:date>
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<mods:number id="0EF3A9A08E61A59CAA901747BA996378">2021-06-09</mods:number>
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<mods:number id="65322177D6C7AA03A5CBA652FD941511">4982</mods:number>
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<mods:detail id="F381CD61BC4B3C762EF8FE4DD3C0D74D" type="issue">
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<mods:number id="6AF6DF8A9960EEC54978E98B31F4A5F4">1</mods:number>
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<mods:start id="32D1D984C1243750EE5D029938FD8174">1</mods:start>
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<mods:end id="C978BF6F3F3356810620AD834A8CD4A7">70</mods:end>
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<mods:identifier id="85A7FC8515F7D2F68850CBC33D923AA1" type="DOI">10.11646/zootaxa.4982.1.1</mods:identifier>
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<treatment id="D446F043FFCDFFE58C92FF39FA1404FE" ID-DOI="http://doi.org/10.5281/zenodo.5043416" ID-GBIF-Taxon="183225140" ID-Zenodo-Dep="5043416" LSID="urn:lsid:plazi:treatment:D446F043FFCDFFE58C92FF39FA1404FE" httpUri="http://treatment.plazi.org/id/D446F043FFCDFFE58C92FF39FA1404FE" lastPageNumber="57" pageId="56" pageNumber="57">
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<subSubSection id="14F512DEFFCDFFE58C92FF39FE7200B1" box="[151,507,151,177]" pageId="56" pageNumber="57" type="nomenclature">
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<paragraph id="5C504155FFCDFFE58C92FF39FE7200B1" blockId="56.[151,507,151,177]" box="[151,507,151,177]" pageId="56" pageNumber="57">
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<emphasis id="6E9B9D47FFCDFFE58C92FF39FE7200B1" bold="true" box="[151,507,151,177]" pageId="56" pageNumber="57">
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Genus
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<taxonomicName id="9BEF3AD6FFCDFFE58CEBFF39FE7200B1" ID-CoL="99G38" authority="Habe, 1964" authorityName="Habe" authorityYear="1964" box="[238,507,151,177]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFCDFFE58CEBFF39FEEE00B1" bold="true" box="[238,359,151,177]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
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<bibRefCitation id="387E3CA4FFCDFFE58D6AFF39FE7200B1" author="Habe, T." box="[367,507,151,177]" pageId="56" pageNumber="57" refId="ref56846" refString="Habe, T. (1964) Shells of the Western Pacific in color. Vol. II. Osaka, Hoikusha, 233 pp." type="book" year="1964">Habe, 1964</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="14F512DEFFCDFFE58C92FF71FA0400F9" box="[151,1421,223,250]" pageId="56" pageNumber="57" type="type_taxon">
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<paragraph id="5C504155FFCDFFE58C92FF71FA0400F9" blockId="56.[151,1437,223,1078]" box="[151,1421,223,250]" pageId="56" pageNumber="57">
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<emphasis id="6E9B9D47FFCDFFE58C92FF71FEBB00F9" bold="true" box="[151,306,223,249]" pageId="56" pageNumber="57">
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<typeStatus id="8354FFF7FFCDFFE58C92FF71FF5B00F9" box="[151,210,223,249]" pageId="56" pageNumber="57">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="9BEF3AD6FFCDFFE58D3CFF71FD4800F9" ID-CoL="43RKW" authority="Kuroda & Sakurai" authorityName="Kuroda & Sakurai" authorityYear="1959" box="[313,705,223,249]" class="Gastropoda" family="Costellariidae" genus="Mitropifex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="isaoi">
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<emphasis id="6E9B9D47FFCDFFE58D3CFF71FE6200F9" box="[313,491,223,249]" italics="true" pageId="56" pageNumber="57">Mitropifex isaoi</emphasis>
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Kuroda & Sakurai
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</taxonomicName>
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in
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<bibRefCitation id="387E3CA4FFCDFFE58EE1FF71FCF700FA" author="Kuroda, T." box="[740,894,223,250]" pageId="56" pageNumber="57" pagination="317 - 335" refId="ref58557" refString="Kuroda, T. (1959) Description of new species of marine shells from Japan. Venus, 20, 317 - 335." type="journal article" year="1959">Kuroda, 1959</bibRefCitation>
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, by monotypy, Present-day, Indo-West Pacific.
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</paragraph>
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</subSubSection>
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<subSubSection id="14F512DEFFCDFFE58CC2FEADFAE101F6" pageId="56" pageNumber="57" type="description">
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<paragraph id="5C504155FFCDFFE58CC2FEADFAE101F6" blockId="56.[151,1437,223,1078]" pageId="56" pageNumber="57">
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<emphasis id="6E9B9D47FFCDFFE58CC2FEADFED0011D" bold="true" box="[199,345,259,285]" pageId="56" pageNumber="57">Description.</emphasis>
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“
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<emphasis id="6E9B9D47FFCDFFE58D6BFEADFB9701F5" italics="true" pageId="56" pageNumber="57">Shell elongate fusiform, sometimes widely fusiform or turriform, typically with high spire and long to very long siphonal canal. Suture typically canaliculated or impressed, often resulting in telescopic appearance of spire. Whorls outline varying from subcylindric to flattened or evenly convex. Axial sculpture of straight ribs strong and widely set to fine and closely set. Spiral sculpture varying from fine grooves, restricted to siphonal canal or pronounced in interspaces between axials only, to strong cords overriding axial ribs to form beads or strong gemmae at intersections. Aperture elongate, its outer lip smooth or bearing fine lirae on its inner surface. Inner lip with between three and five columellar folds, with the upmost fold being the strongest</emphasis>
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” (
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<bibRefCitation id="387E3CA4FFCDFFE5883DFE75FAD201F6" author="Fedosov, A. & Puillandre, N. & Herrmann, M. & Dgebuadze, P. & Bouchet, P." box="[1080,1371,475,502]" pageId="56" pageNumber="57" pagination="541 - 626" refId="ref56065" refString="Fedosov, A. E, Puillandre, N., Herrmann, M., Dgebuadze, P. & Bouchet, P. (2017) Phylogeny, systematics, and evolution of the family Costellariidae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 179, 541 - 626. https: // doi. org / 10.1111 / zoj. 12431" type="journal article" year="2017">
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Fedosov
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<emphasis id="6E9B9D47FFCDFFE588A5FE72FB5801F5" box="[1184,1233,475,501]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
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., 2017: 564
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="14F512DEFFCDFFE58CC2FE51FD0A0435" pageId="56" pageNumber="57" type="discussion">
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<paragraph id="5C504155FFCDFFE58CC2FE51FC110339" blockId="56.[151,1437,223,1078]" pageId="56" pageNumber="57">
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<emphasis id="6E9B9D47FFCDFFE58CC2FE51FEC50219" bold="true" box="[199,332,511,537]" pageId="56" pageNumber="57">Discussion.</emphasis>
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The large and slender fusiform
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<taxonomicName id="9BEF3AD6FFCDFFE58EAFFDAEFBAE0219" authority="Schaffer, 1898" authorityName="Schaffer" authorityYear="1898" box="[682,1063,511,537]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="neudorfensis">
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<emphasis id="6E9B9D47FFCDFFE58EAFFDAEFCF60219" box="[682,895,511,537]" italics="true" pageId="56" pageNumber="57">Mitra neudorfensis</emphasis>
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<bibRefCitation id="387E3CA4FFCDFFE58F80FE51FBAE0219" author="Schaffer F." box="[901,1063,511,537]" pageId="56" pageNumber="57" pagination="533 - 548" refId="ref61664" refString="Schaffer F. (1898) Der marine Tegel von Theben - Neudorf in Ungarn. Jahrbuch der k. k. Geologischen Reichsanstalt, 47, 533 - 548. [https: // opac. geologie. ac. at / ais 312 / dokumente / JB 0473 _ 533 _ A. pdf]" type="journal article" year="1898">Schaffer, 1898</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="9BEF3AD6FFCDFFE5885FFE51FEA8023D" authority="Baluk, 1997" authorityName="Baluk" authorityYear="1997" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="pseudocupressinum">
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<emphasis id="6E9B9D47FFCDFFE5885FFE51FA150219" box="[1114,1436,511,537]" italics="true" pageId="56" pageNumber="57">Vexillum pseudocupressinum</emphasis>
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<bibRefCitation id="387E3CA4FFCDFFE58C92FD8DFEA8023D" author="Baluk, W." box="[151,289,547,573]" pageId="56" pageNumber="57" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk, 1997</bibRefCitation>
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</taxonomicName>
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have been placed so far in
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<taxonomicName id="9BEF3AD6FFCDFFE58E5EFD8DFD32023D" authorityName="Roding" authorityYear="1798" box="[603,699,547,573]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFCDFFE58E5EFD8DFD32023D" box="[603,699,547,573]" italics="true" pageId="56" pageNumber="57">Vexillum</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="387E3CA4FFCDFFE58EE2FD8DFC17023D" author="Biskupic, R." box="[743,926,547,573]" pageId="56" pageNumber="57" pagination="75 - 88" refId="ref54576" refString="Biskupic, R. (2020) A new evidence of Vexillum (Gastropoda: Costellariidae) from the Middle Miocene (Serravallian) of Vienna Basin (Slovakia). Acta Geologica Slovaca, 12 / 2, 75 - 88." type="journal article" year="2020">Biskupič (2020)</bibRefCitation>
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and
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<bibRefCitation id="387E3CA4FFCDFFE58FD3FD8DFBE6023D" author="Baluk, W." box="[982,1135,547,573]" pageId="56" pageNumber="57" pagination="1 - 75" refId="ref54188" refString="Baluk, W. (1997) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part III. Acta Geologica Polonica, 47, 1 - 75. [https: // geojournals. pgi. gov. pl / agp / article / view / 13688 / 12126]" type="journal article" year="1997">Bałuk (1997)</bibRefCitation>
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. Alternatively, we propose a placement in
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<taxonomicName id="9BEF3AD6FFCDFFE58D41FDE6FDB70261" authority="Habe, 1964" authorityName="Habe" authorityYear="1964" box="[324,574,583,609]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFCDFFE58D41FDE6FE3C0261" box="[324,437,584,609]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
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<bibRefCitation id="387E3CA4FFCDFFE58DB9FDE9FDB70261" author="Habe, T." box="[444,574,583,609]" pageId="56" pageNumber="57" refId="ref56846" refString="Habe, T. (1964) Shells of the Western Pacific in color. Vol. II. Osaka, Hoikusha, 233 pp." type="book" year="1964">Habe, 1964</bibRefCitation>
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</taxonomicName>
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. The genus
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<taxonomicName id="9BEF3AD6FFCDFFE58ECDFDE6FCB00261" authorityName="Habe" authorityYear="1964" box="[712,825,584,609]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFCDFFE58ECDFDE6FCB00261" box="[712,825,584,609]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="9BEF3AD6FFCDFFE58FCAFDE9FB2E0261" box="[975,1191,583,609]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="especially">
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<emphasis id="6E9B9D47FFCDFFE58FCAFDE9FB2E0261" box="[975,1191,583,609]" italics="true" pageId="56" pageNumber="57">Vexillum especially</emphasis>
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</taxonomicName>
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in its longer siphonal canal, which lacks a deep notch. In addition, axial ribs in
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<taxonomicName id="9BEF3AD6FFCDFFE58F35FDC2FC280285" authorityName="Habe" authorityYear="1964" box="[816,929,620,645]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFCDFFE58F35FDC2FC280285" box="[816,929,620,645]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
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</taxonomicName>
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are typically straight, but tend to be arcuate and undulating in
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<taxonomicName id="9BEF3AD6FFCDFFE58D6CFD21FD4402A9" authority="(Fedosov et al. 2017)" baseAuthorityName="Fedosov" baseAuthorityYear="2017" box="[361,717,655,681]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
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<emphasis id="6E9B9D47FFCDFFE58D6CFD21FE4002A9" box="[361,457,655,681]" italics="true" pageId="56" pageNumber="57">Vexillum</emphasis>
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(
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<bibRefCitation id="387E3CA4FFCDFFE58DDEFD21FD4C02A9" author="Fedosov, A. & Puillandre, N. & Herrmann, M. & Dgebuadze, P. & Bouchet, P." box="[475,709,655,681]" pageId="56" pageNumber="57" pagination="541 - 626" refId="ref56065" refString="Fedosov, A. E, Puillandre, N., Herrmann, M., Dgebuadze, P. & Bouchet, P. (2017) Phylogeny, systematics, and evolution of the family Costellariidae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 179, 541 - 626. https: // doi. org / 10.1111 / zoj. 12431" type="journal article" year="2017">
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Fedosov
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<emphasis id="6E9B9D47FFCDFFE58E40FD3EFDF602A9" box="[581,639,655,681]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
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2017
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</bibRefCitation>
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)
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</taxonomicName>
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.
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<taxonomicName id="9BEF3AD6FFCDFFE58EDFFD3EFB9502A9" authorityName="Harzhauser & Landau" authorityYear="2021" baseAuthorityName="Baluk" baseAuthorityYear="1997" box="[730,1052,655,681]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="pseudocupressina">
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<emphasis id="6E9B9D47FFCDFFE58EDFFD3EFB9502A9" box="[730,1052,655,681]" italics="true" pageId="56" pageNumber="57">Tosapusia pseudocupressina</emphasis>
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</taxonomicName>
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is strikingly similar to the extant Indo-West Pacific
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<taxonomicName id="9BEF3AD6FFCDFFE58D60FD1AFCF702CD" authority="(Guillot de Suduiraut, 2007)" baseAuthorityName="Guillot de Suduiraut" baseAuthorityYear="2007" box="[357,894,691,717]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="evelynae">
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<emphasis id="6E9B9D47FFCDFFE58D60FD1AFDB402CD" box="[357,573,691,717]" italics="true" pageId="56" pageNumber="57">Tosapusia evelynae</emphasis>
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|
(
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<bibRefCitation id="387E3CA4FFCDFFE58E4EFD1DFCFF02CD" author="Guillot de Suduiraut E." box="[587,886,691,717]" pageId="56" pageNumber="57" pagination="96 - 100" refId="ref56815" refString="Guillot de Suduiraut E. (2007) Description of three new species of Costellariidae from the Indian Ocean and the Pacific. Visaya, 2, 96 - 100." type="journal article" year="2007">Guillot de Suduiraut, 2007</bibRefCitation>
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)
|
|
</taxonomicName>
|
|
in outline and both species display a comparable variability in sculpture.
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|
<taxonomicName id="9BEF3AD6FFCDFFE58D9AFD76FC9702F1" authority="Kuroda & Sakurai" authorityName="Kuroda & Sakurai" box="[415,798,727,753]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="isaoi">
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<emphasis id="6E9B9D47FFCDFFE58D9AFD76FDC502F1" box="[415,588,728,753]" italics="true" pageId="56" pageNumber="57">Tosapusia isaoi</emphasis>
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|
Kuroda & Sakurai
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</taxonomicName>
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|
in
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|
<bibRefCitation id="387E3CA4FFCDFFE58F3AFD79FC5102F2" author="Kuroda, T." box="[831,984,727,754]" pageId="56" pageNumber="57" pagination="317 - 335" refId="ref58557" refString="Kuroda, T. (1959) Description of new species of marine shells from Japan. Venus, 20, 317 - 335." type="journal article" year="1959">Kuroda, 1959</bibRefCitation>
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|
is a modern analogue of
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|
<taxonomicName id="9BEF3AD6FFCDFFE588E9FD76FA1C02F1" authorityName="Schaffer" authorityYear="1898" box="[1260,1429,727,753]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="neudorfensis">
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|
<emphasis id="6E9B9D47FFCDFFE588E9FD76FA1C02F1" box="[1260,1429,727,753]" italics="true" pageId="56" pageNumber="57">T. neudorfensis</emphasis>
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|
</taxonomicName>
|
|
, but lacks a subsutural groove. Moreover, the preference for offshore settings of the two discussed Miocene species is consistent with the deep water habitat of extant
|
|
<taxonomicName id="9BEF3AD6FFCDFFE58ECDFC8EFC110339" box="[712,920,799,825]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="undefined-a">
|
|
<emphasis id="6E9B9D47FFCDFFE58ECDFC8EFCB00339" box="[712,825,800,825]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
|
|
species.
|
|
</taxonomicName>
|
|
</paragraph>
|
|
<paragraph id="5C504155FFCDFFE58CC2FCEDFB530381" blockId="56.[151,1437,223,1078]" pageId="56" pageNumber="57">
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|
A further species from the European Neogene, which should be placed in
|
|
<taxonomicName id="9BEF3AD6FFCDFFE58804FCEAFBFB035D" authorityName="Habe" authorityYear="1964" box="[1025,1138,836,861]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
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|
<emphasis id="6E9B9D47FFCDFFE58804FCEAFBFB035D" box="[1025,1138,836,861]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
|
|
</taxonomicName>
|
|
, is
|
|
<taxonomicName id="9BEF3AD6FFCDFFE5889CFCEAFF500381" authority="(Brocchi, 1814)" baseAuthorityName="Brocchi" baseAuthorityYear="1814" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="cupressina">
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|
<emphasis id="6E9B9D47FFCDFFE5889CFCEAFAA5035D" box="[1177,1324,836,861]" italics="true" pageId="56" pageNumber="57">T. cupressina</emphasis>
|
|
(
|
|
<bibRefCitation id="387E3CA4FFCDFFE5893EFCEDFF590381" author="Brocchi, G." pageId="56" pageNumber="57" refId="ref54754" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Vols. 1 - 2. Stamperia Reale, Milano, 712 pp. [https: // www. biodiversitylibrary. org / item / 43873 # page / 9 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
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|
)
|
|
</taxonomicName>
|
|
from the middle Miocene to Pliocene of the proto-Mediterranean Sea (
|
|
<bibRefCitation id="387E3CA4FFCDFFE58FF0FCC9FB470381" author="Landau, B. M. & Harzhauser, M. & Yslamodlu, Y. & Marques da Silva, C." box="[1013,1230,871,897]" pageId="56" pageNumber="57" pagination="3 - 576" refId="ref58833" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. & Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576." type="journal article" year="2013">
|
|
Landau
|
|
<emphasis id="6E9B9D47FFCDFFE58857FCC6FB030381" box="[1106,1162,871,897]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
|
|
2013
|
|
</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="5C504155FFCDFFE58CC2FC22FD0A0435" blockId="56.[151,1437,223,1078]" pageId="56" pageNumber="57">
|
|
Extant
|
|
<taxonomicName id="9BEF3AD6FFCDFFE58D12FC22FE5603A5" box="[279,479,907,933]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="undetermined">
|
|
<emphasis id="6E9B9D47FFCDFFE58D12FC22FE0103A5" box="[279,392,908,933]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
have an Indo-West Pacific distribution (Fedosov
|
|
<emphasis id="6E9B9D47FFCDFFE58803FC22FBB403A5" box="[1030,1085,907,933]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
|
|
2007) but reached to the protoMediterranean Sea and the Central Paratethys during the Miocene. The genus persisted in the proto-Mediterranean Sea until the Pliocene, clearly postdating the Tethys closure during the early Miocene (Harzhauser
|
|
<emphasis id="6E9B9D47FFCDFFE58915FC7AFACA03ED" box="[1296,1347,979,1005]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
|
|
. 2007). Therefore, our data document a restriction of the former distribution area of
|
|
<taxonomicName id="9BEF3AD6FFCDFFE58FF4FC56FBEB0411" authorityName="Habe" authorityYear="1964" box="[1009,1122,1016,1041]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="genus">
|
|
<emphasis id="6E9B9D47FFCDFFE58FF4FC56FBEB0411" box="[1009,1122,1016,1041]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
|
|
</taxonomicName>
|
|
and should not be misinterpreted as immigration from the IWP region.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="14F512DEFFCDFFE58C92FB29FA1404FE" pageId="56" pageNumber="57" type="key">
|
|
<paragraph id="5C504155FFCDFFE58C92FB29FDFA04A1" blockId="56.[151,627,1159,1185]" box="[151,627,1159,1185]" pageId="56" pageNumber="57">
|
|
<emphasis id="6E9B9D47FFCDFFE58C92FB29FDFA04A1" bold="true" box="[151,627,1159,1185]" pageId="56" pageNumber="57">
|
|
Key to Paratethyan
|
|
<taxonomicName id="9BEF3AD6FFCDFFE58D96FB29FDE704A1" box="[403,622,1159,1185]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="undetermined">
|
|
<emphasis id="6E9B9D47FFCDFFE58D96FB29FD8504A1" bold="true" box="[403,524,1159,1185]" italics="true" pageId="56" pageNumber="57">Tosapusia</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
</paragraph>
|
|
<paragraph id="5C504155FFCDFFE58C92FB63FA1404FE" blockId="56.[151,1437,1228,1279]" pageId="56" pageNumber="57">
|
|
- Sculpture very fine, almost straight-sided whorls.............................................
|
|
<taxonomicName id="9BEF3AD6FFCDFFE588DFFB63FA1504E1" ID-CoL="8KSZG" authorityName="Harzhauser & Landau" authorityYear="2021" baseAuthorityName="Baluk" baseAuthorityYear="1997" box="[1242,1436,1228,1250]" class="Gastropoda" family="Costellariidae" genus="Tosapusia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="pseudocupressina">
|
|
<emphasis id="6E9B9D47FFCDFFE588DFFB63FA1504E1" box="[1242,1436,1228,1250]" italics="true" pageId="56" pageNumber="57">T. pseudocupressina</emphasis>
|
|
</taxonomicName>
|
|
- Prominent subsutural furrow.................................................................
|
|
<taxonomicName id="9BEF3AD6FFCDFFE5890FFB44FA1404FE" authorityName="Schaffer" authorityYear="1898" box="[1290,1437,1257,1279]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="neudorfensis">
|
|
<emphasis id="6E9B9D47FFCDFFE5890FFB44FA1404FE" box="[1290,1437,1257,1279]" italics="true" pageId="56" pageNumber="57">T. neudorfensis</emphasis>
|
|
</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |