treatments-xml/data/D2/FE/FB/D2FEFB8B6DD7503198692CF249A17299.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.994.56810" ID-GBIF-Dataset="8639369d-0b63-4725-aa0c-aed78224bbe5" ID-PMC="PMC7686225" ID-Pensoft-Pub="1313-2970-994-1" ID-Pensoft-UUID="11A23FB601685F7ABFCC123875393447" ID-PubMed="33273881" ID-ZooBank="DB1879C919044320BE674D9F863F6775" ModsDocID="1313-2970-994-1" checkinTime="1605693934925" checkinUser="pensoft" docAuthor="Ng, Peter K. L." docDate="2020" docId="D2FEFB8B6DD7503198692CF249A17299" docLanguage="en" docName="ZooKeys 994: 1-34" docOrigin="ZooKeys 994" docSource="http://dx.doi.org/10.3897/zookeys.994.56810" docTitle="Johora michaeli Ng 2020, sp. nov." docType="treatment" docUuid="4CA305BB-71CF-49E5-9559-24BB4019660C" docUuidSource="ZooBank" docVersion="4" id="11A23FB601685F7ABFCC123875393447" lastPageNumber="1" masterDocId="11A23FB601685F7ABFCC123875393447" masterDocTitle="Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus" masterLastPageNumber="34" masterPageNumber="1" pageNumber="1" updateTime="1668169916828" updateUser="ExternalLinkService">
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<mods:title>Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ng, Peter K. L.</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5946-0608</mods:nameIdentifier>
<mods:nameIdentifier type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:date>2020</mods:date>
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<mods:number>994</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.994.56810</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.994.56810</mods:identifier>
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<treatment ID-GBIF-Taxon="169094340" LSID="urn:lsid:zoobank.org:act:4CA305BB-71CF-49E5-9559-24BB4019660C" httpUri="http://treatment.plazi.org/id/D2FEFB8B6DD7503198692CF249A17299" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="D2FEFB8B-6DD7-5031-9869-2CF249A17299" authority="Ng, 2020" authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora michaeli" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="michaeli" status="sp. nov.">Johora michaeli</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">Figures 1H</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">, 2H</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">, 3H</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">, 4H</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">, 5H</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">, 9I-L</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">, 12H</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">, 13H</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">, 14H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: male (22.7 
<normalizedToken originalValue="×">x</normalizedToken>
19.2 mm) (ZRC 2020.0361), waterfalls at rock pools, ca. 5 minutes walk upstream from chalets, Sekayu Waterfall, 
<geoCoordinate degrees="4" direction="north" minutes="59" orientation="latitude" precision="15" seconds="35" value="4.993056">4°59'35&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="102" direction="east" minutes="56" orientation="longitude" precision="15" seconds="50" value="102.94722">102°56'50&quot;E</geoCoordinate>
, Terengganu, coll. Tan HH, 21 October 1998. Paratype: 1 female (29.9 
<normalizedToken originalValue="×">x</normalizedToken>
25.0 mm) (ZRC 2010.0047), same data as holotype. Others: 1 female (30.8 
<normalizedToken originalValue="×">x</normalizedToken>
25.2 mm) (ZRC 1984.6794), Gunung Padang, Terengganu, 
<geoCoordinate degrees="4" direction="north" minutes="50" orientation="latitude" precision="1" seconds="55.7" value="4.848806">4°50'55.7&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="102" direction="east" minutes="52" orientation="longitude" precision="1" seconds="1.9" value="102.867195">102°52'1.9&quot;E</geoCoordinate>
, coll. Hislep JSA, 1952. All locations in Peninsular Malaysia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult carapace width to length ratio 1.20-1.22 (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1H</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2H</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12H</figureCitation>
); dorsal surface gently convex in frontal view, not inflated (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3H</figureCitation>
); frontal margin almost straight (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2H</figureCitation>
); suborbital, pterygostomial and sub-branchial regions rugose, pterygostomial region covered with dense setae (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3H</figureCitation>
); epigastric cristae distinct, distinctly anterior to sharp postorbital cristae, postorbital cristae with lateral edges low, not joining lateral margin (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2H</figureCitation>
); external orbital tooth separated from epibranchial tooth by distinct cleft, epibranchial tooth sharp, distinct (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2H</figureCitation>
); anterolateral margin distinctly convex (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2H</figureCitation>
); posterolateral margin with median concavity or sinuous, distinctly converging towards gently convex to almost straight, entire posterior carapace margin (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2H</figureCitation>
); posterior margin of epistome with triangular median triangle, lateral margin obliquely sloping (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3H</figureCitation>
); outer surfaces of third maxillipeds with long stiff setae; ischium subrectangular, with shallow median oblique groove (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3H</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4H</figureCitation>
); ambulatory legs not elongate, length to width ratio of merus of fourth ambulatory leg 2.7-2.8 (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1H</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12H</figureCitation>
); G1 subterminal segment gradually tapering from broad proximal part to slender distal part, without distinct shelf-like structure along gently concave outer margin; terminal segment almost straight, slightly curved outwards (from median part of sternum), ca. two-thirds length of subterminal segment, surfaces with scattered short setae (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">9I-K</figureCitation>
); G2 subequal in length to G1, distal segment long, about one-third length of basal segment (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">9L</figureCitation>
). Female pleon ovate; somites 3-6 progressively narrower; telson subtriangular (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13H</figureCitation>
). Vulvae large, on anterior half of sternite 6, adjacent to suture with sternite 5, lateral sternal vulvar cover subtruncate (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14H</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 14.</emphasis>
Sternopleonal cavity and vulvae 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName baseAuthorityName="Ng" baseAuthorityYear="1986" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora intermedia" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="1">Johora intermedia</emphasis>
</taxonomicName>
Ng, 1986, female (17.9 
<normalizedToken originalValue="×">x</normalizedToken>
14.7 mm) (ZRC 2001.1002), Selangor 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
(Bott, 1966), female (24.4 
<normalizedToken originalValue="×">x</normalizedToken>
21.0 mm) (ZRC 1984.6765), Pahang 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
Ng, 1990, female (31.7 
<normalizedToken originalValue="×">x</normalizedToken>
25.5 mm) (ZRC 2001.1167), Terengganu 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
Ng &amp; Takeda, 1992, paratype female (22.1 
<normalizedToken originalValue="×">x</normalizedToken>
19.0 mm) (ZRC 1984.6675), Kelantan 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 
<normalizedToken originalValue="×">x</normalizedToken>
17.4 mm) (ZRC 2006.0053), Thailand 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
sp. nov., paratype female (40.4 
<normalizedToken originalValue="×">x</normalizedToken>
34.4 mm) (ZRC 1995.270), Pahang 
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
sp. nov., paratype female (32.8 
<normalizedToken originalValue="×">x</normalizedToken>
26.9 mm) (ZRC 1995.268), Perak 
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
sp. nov., paratype female (29.9 
<normalizedToken originalValue="×">x</normalizedToken>
25.0 mm) (ZRC 2010.0047), Terengganu.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species is named after the last director of the Raffles Museum (1946-1967), the late Michael Tweedie, an intrepid collector of interesting animals from Malaysia. The author had the pleasure of knowing him, finally meeting him when the refreshed museum opened as the ZRC in 1988; and even after many years since retiring, he retained his great passion for his crabs, snakes, and fish.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The holotype male of 
<taxonomicName authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora michaeli" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">Johora michaeli</emphasis>
</taxonomicName>
sp. nov. is not fully adult and it is clear that it can grow larger, with the adult females reaching 30 mm in carapace width. The G1, however, remains diagnostic, with the terminal segment elongate and almost straight (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">9I-K</figureCitation>
). While this somewhat resembles that of 
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
which is known from the nearby island of Pulau Redang, that of 
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
is distinctly less elongate and slender, being only about two-thirds the length of subterminal segment. Specimens of 
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
even smaller than the type of 
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
remain the same G1 terminal segment shape and proportions of adults (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">9E-G</figureCitation>
) so the differences observed here are independent of size. The vulva of 
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
is diagnostic, the lateral sternal vulvar cover being triangular in shape (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14C</figureCitation>
); it is subtruncate in 
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14H</figureCitation>
). The general shape of the G1 terminal segment somewhat resembles that of 
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
s. str. (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8A-C</figureCitation>
), it is clearly straighter and slenderer in 
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">9I-K</figureCitation>
). It is also unlike that of 
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
which has a proportionately and even straighter G1 terminal segment (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8E-G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The G1 of 
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
also resembles that of 
<taxonomicName lsidName="J. singaporensis" pageId="0" pageNumber="1" rank="species" species="singaporensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. singaporensis</emphasis>
</taxonomicName>
but the terminal segment in this species is longer (
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
: fig. 8A, B). In any case, the carapace of 
<taxonomicName lsidName="J. singaporensis" pageId="0" pageNumber="1" rank="species" species="singaporensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. singaporensis</emphasis>
</taxonomicName>
is quite different from that of 
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
as it is a member of the 
<taxonomicName lsidName="J. johorensis" pageId="0" pageNumber="1" rank="species" species="johorensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. johorensis</emphasis>
</taxonomicName>
species group (see remarks for the species group under remarks for the genus).
</paragraph>
<paragraph pageId="0" pageNumber="1">One large female (ZRC 1984.6794) had been collected from Gunung Padang, which is relatively close to the type locality in Sekayu Falls. Both are part of the same mountain system east of Lake Kenyir in Terengganu.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora michaeli" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">Johora michaeli</emphasis>
</taxonomicName>
sp. nov. is known so far only from highland streams in central Terengganu (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of species of the Johora tahanensis species group 1 J. tahanensis (Bott, 1966) 2 J. thoi Ng, 1990 3 J. hoiseni Ng &amp; Takeda, 1992 4 J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005 5 J. booliati sp. nov. 6 J. erici sp. nov. 7 J. michaeli sp. nov. Map data 2020 Google." figureDoi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1">15</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 15.</emphasis>
Distribution of species of the 
<taxonomicName authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">Johora tahanensis</emphasis>
</taxonomicName>
species group 
<emphasis bold="true" pageId="0" pageNumber="1">1</emphasis>
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
(Bott, 1966) 
<emphasis bold="true" pageId="0" pageNumber="1">2</emphasis>
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
Ng, 1990 
<emphasis bold="true" pageId="0" pageNumber="1">3</emphasis>
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
Ng &amp; Takeda, 1992 
<emphasis bold="true" pageId="0" pageNumber="1">4</emphasis>
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
Leelawathanagoon, Lheknim &amp; Ng, 2005 
<emphasis bold="true" pageId="0" pageNumber="1">5</emphasis>
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">6</emphasis>
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">7</emphasis>
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
sp. nov. Map data 2020 Google.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation">
<paragraph pageId="0" pageNumber="1">Conservation.</paragraph>
<paragraph pageId="0" pageNumber="1">
The conservation status for 
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
should be regarded as vulnerable as it is known from a relatively well protected area that is not subject to development (see 
<bibRefCitation DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>