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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1070.73484" ID-GBIF-Dataset="457674ab-9403-4b5b-86ae-d60b98a58efc" ID-PMC="PMC8599384" ID-Pensoft-Pub="1313-2970-1070-41" ID-Pensoft-UUID="DB30D8A82FC3569AA28D0765F36139AE" ID-PubMed="34819770" ID-ZooBank="C6532DEC44A24150BBA311BF711C068B" ModsDocID="1313-2970-1070-41" checkinTime="1636588170873" checkinUser="pensoft" docAuthor="Sartori, Michel" docDate="2021" docId="D29523E862E85AD68AB1E22F8950AB8D" docLanguage="en" docName="ZooKeys 1070: 41-50" docOrigin="ZooKeys 1070" docPubDate="2021-11-10" docSource="http://dx.doi.org/10.3897/zookeys.1070.73484" docTitle="Neohagenulus hodeceki Sartori 2021, sp. nov." docType="treatment" docUuid="FDBF727E-87FD-4120-9E62-BDBCECBA8659" docUuidSource="ZooBank" docVersion="4" id="DB30D8A82FC3569AA28D0765F36139AE" lastPageNumber="41" masterDocId="DB30D8A82FC3569AA28D0765F36139AE" masterDocTitle="A new species of Neohagenulus Traver, 1938 from Hispaniola (Ephemeroptera, Leptophlebiidae, Hagenulinae, Hagenulini)" masterLastPageNumber="50" masterPageNumber="41" pageNumber="41" updateTime="1668151114791" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Neohagenulus Traver, 1938 from Hispaniola (Ephemeroptera, Leptophlebiidae, Hagenulinae, Hagenulini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sartori, Michel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3397-3397</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1014, Lausanne, Switzerland &amp; Department of Ecology and Evolution, Biophore, University of Lausanne, CH- 1015, Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">michel.sartori@vd.ch</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-11-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1070</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>50</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1070.73484</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1070.73484</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1070-41</mods:identifier>
<mods:identifier type="ZooBank">C6532DEC44A24150BBA311BF711C068B</mods:identifier>
<mods:identifier type="Pensoft-UUID">DB30D8A82FC3569AA28D0765F36139AE</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="190304161" LSID="urn:lsid:zoobank.org:act:FDBF727E-87FD-4120-9E62-BDBCECBA8659" httpUri="http://treatment.plazi.org/id/D29523E862E85AD68AB1E22F8950AB8D" lastPageNumber="41" pageId="0" pageNumber="41">
<subSubSection pageId="0" pageNumber="41" type="nomenclature">
<paragraph pageId="0" pageNumber="41">
<taxonomicName LSID="http://zoobank.org/FDBF727E-87FD-4120-9E62-BDBCECBA8659" authority="Sartori, 2021" authorityName="Sartori" authorityYear="2021" class="Insecta" family="Leptophlebiidae" genus="Neohagenulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neohagenulus hodeceki" order="Ephemeroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="hodeceki" status="sp. nov.">Neohagenulus hodeceki</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="41">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="materials_examined">
<paragraph pageId="0" pageNumber="41">Material.</paragraph>
<paragraph pageId="0" pageNumber="41">
<materialsCitation collectingDate="2021-07-12" collectorName="J. Hodecek" country="Dominican Republic" elevation="900" latitude="18.866945" location="Valle Nuevo National Park" longLatPrecision="21" longitude="-70.57889" specimenCount="1" stateProvince="La Vega Province" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
<collectingCountry name="Dominican Republic">Dominican Republic</collectingCountry>
male nymph in ethanol,
<collectingRegion country="Dominican Republic" name="La Vega">La Vega Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D29523E862E85AD68AB1E22F8950AB8D:41C437B3D23762C38A9077F65BA5D376" country="Dominican Republic" latitude="18.866945" longLatPrecision="21" longitude="-70.57889" name="Valle Nuevo National Park" stateProvince="La Vega Province">Valle Nuevo National Park</location>
,
<geoCoordinate degrees="18" direction="north" minutes="52" orientation="latitude" precision="15" seconds="01" value="18.866945">18°52'01&quot;N</geoCoordinate>
,
<geoCoordinate degrees="70" direction="west" minutes="34" orientation="longitude" precision="15" seconds="44" value="-70.57889">70°34'44&quot;W</geoCoordinate>
,
<collectingDate value="2021-07-12">12 Jul. 2021</collectingDate>
, ca
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</quantity>
a.s.l.
</elevation>
,
<collectorName>
J.
<normalizedToken originalValue="Hodeček">Hodecek</normalizedToken>
</collectorName>
leg. (GBIFCH00834690) [MZL]
</materialsCitation>
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Hodecek" country="Dominican Republic" elevation="900" latitude="18.866945" location="Valle Nuevo National Park" longLatPrecision="21" longitude="-70.57889" specimenCount="2" specimenCount-female="2" stateProvince="La Vega" typeStatus="Paratypes">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">
<typeStatus>Paratypes</typeStatus>
</emphasis>
. 5 nymphs in ethanol [MNHNSD] (MNHNSD 11.05 - MNHNSD 11.09), 12 nymphs in ethanol (GBIFCH00834691),
<specimenCount type="female">2 female</specimenCount>
nymphs on slide (GBIFCH00604114-GBIFCH00604115), same data as holotype. [MZL]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="materials_examined">
<paragraph pageId="0" pageNumber="41">Other material.</paragraph>
<paragraph pageId="0" pageNumber="41">
<materialsCitation collectingDate="2021-07-15" collectorName="J. Hodecek" country="Dominican Republic" elevation="1100" latitude="19.067223" location="Armando Bermudez National Park" longLatPrecision="15" longitude="-70.86408" specimenCount="1" stateProvince="La Vega Province">
<collectingCountry name="Dominican Republic">Dominican Republic</collectingCountry>
1 nymph,
<collectingRegion country="Dominican Republic" name="La Vega">La Vega Province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:D29523E862E85AD68AB1E22F8950AB8D:DCC26DC4E70B946AB8C3226B7B903A97" country="Dominican Republic" latitude="19.067223" longLatPrecision="15" longitude="-70.86408" name="Armando Bermudez National Park" stateProvince="La Vega Province">
Armando
<normalizedToken originalValue="Bermúdez">Bermudez</normalizedToken>
National Park
</location>
,
<geoCoordinate degrees="19" direction="north" minutes="04" orientation="latitude" precision="15" seconds="02" value="19.067223">19°04'02&quot;N</geoCoordinate>
<geoCoordinate degrees="70" direction="west" minutes="51" orientation="longitude" precision="1" seconds="50.7" value="-70.86408">70°51'50.7&quot;W</geoCoordinate>
, ca
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
,
<collectingDate value="2021-07-15">15 Jul. 2021</collectingDate>
,
<collectorName>
J.
<normalizedToken originalValue="Hodeček">Hodecek</normalizedToken>
</collectorName>
leg. (GBIFCH00834694) [MZL]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="etymology">
<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="41">
The new species is named after its collector, Dr
<normalizedToken originalValue="Jiří">Jiri</normalizedToken>
<normalizedToken originalValue="Hodeček">Hodecek</normalizedToken>
(CHUV, Lausanne), forensic entomologist.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="41" type="description">
<paragraph pageId="0" pageNumber="41">Description.</paragraph>
<paragraph pageId="0" pageNumber="41">
<taxonomicName genus="Nymph" lsidName="Nymph" pageId="0" pageNumber="41" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Nymph</emphasis>
</taxonomicName>
(not mature): body length up to 7 mm, cerci slightly longer than body length, paracercus longer than cerci.
</paragraph>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Coloration.</emphasis>
Cuticular coloration evenly light brown on whole body; hypodermal coloration as in Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Neohagenulus hodeceki sp. nov., nymphal habitus A dorsal view B lateral view C ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1070.73484.figure1" httpUri="https://binary.pensoft.net/fig/609353" pageId="0" pageNumber="41">1</figureCitation>
: head washed with grey, darker between ocelli, convex band between antennae, upper portion of male eyes black; prothorax greyish laterally and on posterior margin, mesothorax with blackish tracheation laterally, with maculae posteriorly; legs light brown, apex of femora with blackish dots, tarsi darker than tibiae; abdominal tergites I-V light brown with posterior black band larger laterally and with two antero-submedian maculae, tergites VI-VIII greyish brown, posterior margin and sagittal line light brown, tergite IX paler than previous ones, tergite X medium brown; sternites uniformly light brown, last two darker; cerci uniformly light brown.
</paragraph>
<caption doi="10.3897/zookeys.1070.73484.figure1" httpUri="https://binary.pensoft.net/fig/609353" pageId="0" pageNumber="41" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 1.</emphasis>
<taxonomicName authorityName="Sartori" authorityYear="2021" class="Insecta" family="Leptophlebiidae" genus="Neohagenulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neohagenulus hodeceki" order="Ephemeroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="hodeceki">
<emphasis italics="true" pageId="0" pageNumber="41">Neohagenulus hodeceki</emphasis>
</taxonomicName>
sp. nov., nymphal habitus
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
ventral view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Mouthparts.</emphasis>
Labrum (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neohagenulus hodeceki sp. nov. mouthparts A labrum in dorsal view B antero-median emargination of the labrum C left mandible D maxilla. Scale bars: 0.2 mm (A, C, D); 0.05 mm (B)." figureDoi="10.3897/zookeys.1070.73484.figure2" httpUri="https://binary.pensoft.net/fig/609354" pageId="0" pageNumber="41">2A</figureCitation>
) larger than clypeus, about two times broader than long, dorsally with two rows of long and thin setae, proximal row very close to distal margin, tuft of long setae laterally, more abundant at anterolateral corner, ventrally with two submedian fields of long and stout setae, antero-median emargination smooth, with four equally sized denticles (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neohagenulus hodeceki sp. nov. mouthparts A labrum in dorsal view B antero-median emargination of the labrum C left mandible D maxilla. Scale bars: 0.2 mm (A, C, D); 0.05 mm (B)." figureDoi="10.3897/zookeys.1070.73484.figure2" httpUri="https://binary.pensoft.net/fig/609354" pageId="0" pageNumber="41">2B</figureCitation>
). Mandibles outer margin regularly convex, with tuft of small and thin setae in middle, with outer and inner incisors composed of three teeth, outer margins slightly serrated, prostheca with stout and long process and well-developed tuft of thin setae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neohagenulus hodeceki sp. nov. mouthparts A labrum in dorsal view B antero-median emargination of the labrum C left mandible D maxilla. Scale bars: 0.2 mm (A, C, D); 0.05 mm (B)." figureDoi="10.3897/zookeys.1070.73484.figure2" httpUri="https://binary.pensoft.net/fig/609354" pageId="0" pageNumber="41">2C</figureCitation>
). Maxillary palp three-segmented (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neohagenulus hodeceki sp. nov. mouthparts A labrum in dorsal view B antero-median emargination of the labrum C left mandible D maxilla. Scale bars: 0.2 mm (A, C, D); 0.05 mm (B)." figureDoi="10.3897/zookeys.1070.73484.figure2" httpUri="https://binary.pensoft.net/fig/609354" pageId="0" pageNumber="41">2D</figureCitation>
), second segment ca 1.25
<normalizedToken originalValue="×">x</normalizedToken>
length of segment 1, segment 3 conical, outer margin concave near apex, ca 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide and 0.50-0.60
<normalizedToken originalValue="×">x</normalizedToken>
length of segment two, crown of the galea-lacinia with subapical setae arranged in two rows of 8-9 laterally and 11-12 centrally. Hypopharynx with lingua convex, with deep incision distally, lateral processes well developed, but shorter than lingua, slightly curved inwards; superlinguae laterally expanded, distal margin covered with long setae up to the tip (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neohagenulus hodeceki sp. nov. mouthparts and legs A labium B hypopharynx C claw D detail of hind tibia. Scale bars: 0.2 mm (A-C); 0.1 mm (D)" figureDoi="10.3897/zookeys.1070.73484.figure3" httpUri="https://binary.pensoft.net/fig/609355" pageId="0" pageNumber="41">3B</figureCitation>
). Labium (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neohagenulus hodeceki sp. nov. mouthparts and legs A labium B hypopharynx C claw D detail of hind tibia. Scale bars: 0.2 mm (A-C); 0.1 mm (D)" figureDoi="10.3897/zookeys.1070.73484.figure3" httpUri="https://binary.pensoft.net/fig/609355" pageId="0" pageNumber="41">3A</figureCitation>
) with narrow glossae, with stout setae at apex, paraglossae rhomboid, outer margin almost straight, with numerous long setae laterally and apically; labial palp three-segmented, first and second segments subequal in length, or segment two slightly longer, third segment ca 0.35-0.40
<normalizedToken originalValue="×">x</normalizedToken>
length of segment two, 2.5-2.6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide.
</paragraph>
<caption doi="10.3897/zookeys.1070.73484.figure2" httpUri="https://binary.pensoft.net/fig/609354" pageId="0" pageNumber="41" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 2.</emphasis>
<taxonomicName authorityName="Sartori" authorityYear="2021" class="Insecta" family="Leptophlebiidae" genus="Neohagenulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neohagenulus hodeceki" order="Ephemeroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="hodeceki">
<emphasis italics="true" pageId="0" pageNumber="41">Neohagenulus hodeceki</emphasis>
</taxonomicName>
sp. nov. mouthparts
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
labrum in dorsal view
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
antero-median emargination of the labrum
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
left mandible
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
maxilla. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
); 0.05 mm (
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Thorax.</emphasis>
Forelegs with femora ca 2.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, outer margin with row of very long and thin setae, together with apical and subapical rows of long, stout and pointed setae, inner margin with subapical row of long and stout setae (shorter than those on the outer margin), with small stout setae in proximal part, dorsal surface with scattered small to medium-sized stout setae; tibiae shorter than femora, with long and thin setae on outer margin, inner margin with several rows of stout small to medium-sized setae; tarsi only with thin setae, claw moderately hooked, with a row of 12-14 teeth, increasing in size distally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neohagenulus hodeceki sp. nov. mouthparts and legs A labium B hypopharynx C claw D detail of hind tibia. Scale bars: 0.2 mm (A-C); 0.1 mm (D)" figureDoi="10.3897/zookeys.1070.73484.figure3" httpUri="https://binary.pensoft.net/fig/609355" pageId="0" pageNumber="41">3C</figureCitation>
). Midlegs similar to forelegs, except inner margin of tibiae with fewer stout setae. Hindlegs with femora almost 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, similar to forelegs in ornamentation except outer margin with shorter and less numerous thin setae; hind tibia shorter than femora, with outer margin covered with long and thin setae, together with short and medium-sized stout and pointed setae, inner margin with marginal row of short and submarginal row of longer stout setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neohagenulus hodeceki sp. nov. mouthparts and legs A labium B hypopharynx C claw D detail of hind tibia. Scale bars: 0.2 mm (A-C); 0.1 mm (D)" figureDoi="10.3897/zookeys.1070.73484.figure3" httpUri="https://binary.pensoft.net/fig/609355" pageId="0" pageNumber="41">3D</figureCitation>
). Fore wingpad markedly different between male and female nymphs: in males, evenly brown with veins hardly visible (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Neohagenulus hodeceki sp. nov., nymphal habitus A dorsal view B lateral view C ventral view. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1070.73484.figure1" httpUri="https://binary.pensoft.net/fig/609353" pageId="0" pageNumber="41">1A</figureCitation>
), in females with longitudinal veins well marked, crossveins flanked with dark brown bands (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neohagenulus hodeceki sp. nov. thorax and abdomen. A female fore wingpad B female hind wingpad (same specimen) C abdomen in ventral view (arrows indicate posterolateral projections on segments II-IX) D gill IV. Scale bars: 0.5 mm (A, D), 0.2 mm (B), 1 mm (C)." figureDoi="10.3897/zookeys.1070.73484.figure4" httpUri="https://binary.pensoft.net/fig/609356" pageId="0" pageNumber="41">4A</figureCitation>
); in both sexes, base of wingpads tinted with brown in costal, subcostal and anal fields. Hind wingpad very small, tinted with greyish brown at base, with costal process large and slightly pointed, almost as long as the rounded apex of wing (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neohagenulus hodeceki sp. nov. thorax and abdomen. A female fore wingpad B female hind wingpad (same specimen) C abdomen in ventral view (arrows indicate posterolateral projections on segments II-IX) D gill IV. Scale bars: 0.5 mm (A, D), 0.2 mm (B), 1 mm (C)." figureDoi="10.3897/zookeys.1070.73484.figure4" httpUri="https://binary.pensoft.net/fig/609356" pageId="0" pageNumber="41">4B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1070.73484.figure3" httpUri="https://binary.pensoft.net/fig/609355" pageId="0" pageNumber="41" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 3.</emphasis>
<taxonomicName authorityName="Sartori" authorityYear="2021" class="Insecta" family="Leptophlebiidae" genus="Neohagenulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neohagenulus hodeceki" order="Ephemeroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="hodeceki">
<emphasis italics="true" pageId="0" pageNumber="41">Neohagenulus hodeceki</emphasis>
</taxonomicName>
sp. nov. mouthparts and legs
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
labium
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
hypopharynx
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
claw
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
detail of hind tibia. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="41">A-C</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
)
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" italics="true" pageId="0" pageNumber="41">Abdomen</emphasis>
. Posterior margin of tergites I-III smooth, of segments IV-VIII with small needle-like denticles, slightly increasing in size posteriorly, tergites IX and X with triangular denticles; posterior margin of sternite IX concave in the middle (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neohagenulus hodeceki sp. nov. thorax and abdomen. A female fore wingpad B female hind wingpad (same specimen) C abdomen in ventral view (arrows indicate posterolateral projections on segments II-IX) D gill IV. Scale bars: 0.5 mm (A, D), 0.2 mm (B), 1 mm (C)." figureDoi="10.3897/zookeys.1070.73484.figure4" httpUri="https://binary.pensoft.net/fig/609356" pageId="0" pageNumber="41">4C</figureCitation>
); posterolateral projections on abdominal segments II-IX, increasing in size posteriorly (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neohagenulus hodeceki sp. nov. thorax and abdomen. A female fore wingpad B female hind wingpad (same specimen) C abdomen in ventral view (arrows indicate posterolateral projections on segments II-IX) D gill IV. Scale bars: 0.5 mm (A, D), 0.2 mm (B), 1 mm (C)." figureDoi="10.3897/zookeys.1070.73484.figure4" httpUri="https://binary.pensoft.net/fig/609356" pageId="0" pageNumber="41">4C</figureCitation>
); gills present on segments I-VII alike, each gill deeply forked almost at base (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neohagenulus hodeceki sp. nov. thorax and abdomen. A female fore wingpad B female hind wingpad (same specimen) C abdomen in ventral view (arrows indicate posterolateral projections on segments II-IX) D gill IV. Scale bars: 0.5 mm (A, D), 0.2 mm (B), 1 mm (C)." figureDoi="10.3897/zookeys.1070.73484.figure4" httpUri="https://binary.pensoft.net/fig/609356" pageId="0" pageNumber="41">4D</figureCitation>
) with purplish longitudinal and lateral tracheations, size of gills in decreasing order: II=V&gt;III&gt;IV=VI&gt;I&gt;VII. Cerci with whorls of small setae at the end of each segment.
</paragraph>
<caption doi="10.3897/zookeys.1070.73484.figure4" httpUri="https://binary.pensoft.net/fig/609356" pageId="0" pageNumber="41" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="41">
<emphasis bold="true" pageId="0" pageNumber="41">Figure 4.</emphasis>
<taxonomicName authorityName="Sartori" authorityYear="2021" class="Insecta" family="Leptophlebiidae" genus="Neohagenulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neohagenulus hodeceki" order="Ephemeroptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="hodeceki">
<emphasis italics="true" pageId="0" pageNumber="41">Neohagenulus hodeceki</emphasis>
</taxonomicName>
sp. nov. thorax and abdomen.
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
female fore wingpad
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
female hind wingpad (same specimen)
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
abdomen in ventral view (arrows indicate posterolateral projections on segments II-IX)
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
gill IV. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="41">A</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="41">D</emphasis>
), 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="41">B</emphasis>
), 1 mm (
<emphasis bold="true" pageId="0" pageNumber="41">C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="41">Male imago, female imago, eggs unknown.</paragraph>
</subSubSection>
</treatment>
</document>