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<mods:title id="3DE9197A7329CB8E7F0BB5AA52E22275">One new genus and five new nematode species (Monhysterida, Xyalidae) from Tonga and Kermadec Trenches, Southwest Pacific</mods:title>
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<mods:namePart id="3A0419BCE6031FFEDA5ECC0BB9EF91A5">Leduc, Daniel</mods:namePart>
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<treatment id="C619C521BA7F9B49D8CCFE5130A6FB94" ID-DOI="http://doi.org/10.5281/zenodo.5661406" ID-GBIF-Taxon="119617968" ID-Zenodo-Dep="5661406" LSID="urn:lsid:plazi:treatment:C619C521BA7F9B49D8CCFE5130A6FB94" httpUri="http://treatment.plazi.org/id/C619C521BA7F9B49D8CCFE5130A6FB94" lastPageNumber="509" pageId="8" pageNumber="509">
<subSubSection id="06AA27BCBA7F9B49D8CCFE513382FE3D" box="[151,440,439,466]" pageId="8" pageNumber="509" type="nomenclature">
<paragraph id="4E0F7437BA7F9B49D8CCFE513382FE3D" blockId="8.[151,440,439,466]" box="[151,440,439,466]" pageId="8" pageNumber="509">
<heading id="1547C35BBA7F9B49D8CCFE513382FE3D" bold="true" box="[151,440,439,466]" fontSize="11" level="1" pageId="8" pageNumber="509" reason="1">
<taxonomicName id="89B00FB4BA7F9B49D8CCFE513382FE3D" ID-CoL="8FT28" authority="Cobb, 1920" authorityName="Cobb" authorityYear="1920" box="[151,440,439,466]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49D8CCFE513382FE3D" bold="true" box="[151,440,439,466]" pageId="8" pageNumber="509">
<emphasis id="7CC4A825BA7F9B49D8CCFE513319FE3E" bold="true" box="[151,291,439,465]" italics="true" pageId="8" pageNumber="509">Daptonema</emphasis>
Cobb, 1920
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="06AA27BCBA7F9B49D8CCFE18304DFD25" pageId="8" pageNumber="509" type="diagnosis">
<paragraph id="4E0F7437BA7F9B49D8CCFE18304DFD25" blockId="8.[151,1436,510,1147]" pageId="8" pageNumber="509">
<emphasis id="7CC4A825BA7F9B49D8CCFE183330FDF8" bold="true" box="[151,266,510,535]" pageId="8" pageNumber="509">Diagnosis</emphasis>
(modified from
<bibRefCitation id="2A2109C6BA7F9B49D990FE19313DFDF8" author="Fonseca" box="[459,775,511,535]" pageId="8" pageNumber="509" refString="Fonseca, G. &amp; Bezerra, T. N. (2014) Order Monhysterida. In: Shmidt-Rhaesa, A. (Ed.), Handbook of Zoology Gastrotricha, Cyclioneura and Gnathifera. Vol. 2. Nematoda, De Gruyter, Hamburg, pp. 435 - 465." type="book" year="2014">Fonseca &amp; Bezerra (2014))</bibRefCitation>
: Cuticle striated. Somatic setae generally shorter than 1.5 times the corresponding body diameter. Inner labial sensillae papilliform; outer labial and cephalic setae in one circle. Amphideal fovea circular, of varying size and position relative to anterior end. Buccal cavity conicalshaped. Post- and/or pre-vulval sac may be present. Spicules usually short (12 cbd), often strongly curved or Lshaped. Gubernaculum usually present, often with lateral crurae and sometimes with dorso-caudal apophyses. Tail conico-cylindrical with 24 terminal setae.
</paragraph>
</subSubSection>
<subSubSection id="06AA27BCBA7F9B49D89CFD3030A6FB94" pageId="8" pageNumber="509" type="discussion">
<paragraph id="4E0F7437BA7F9B49D89CFD3030A6FB94" blockId="8.[151,1436,510,1147]" pageId="8" pageNumber="509">
<emphasis id="7CC4A825BA7F9B49D89CFD303301FD00" bold="true" box="[199,315,726,751]" pageId="8" pageNumber="509">Remarks.</emphasis>
The genus was revised by
<bibRefCitation id="2A2109C6BA7F9B49DA36FD313111FD00" author="Lorenzen" box="[621,811,727,751]" pageId="8" pageNumber="509" refString="Lorenzen, S. (1977) Revision det Xyalidae (freilebende nematoden) auf der grundlage einer kritischen analyse von 56 Aarten aus Nord- und Ostsee. Veroffentlichungen des Instituts fur Meeresforschung in Bremerhaven, 16, 197 - 261" type="journal article" year="1977">Lorenzen (1977)</bibRefCitation>
. Molecular analyses by
<bibRefCitation id="2A2109C6BA7F9B49DC60FD313735FD00" box="[1083,1295,726,751]" pageId="8" pageNumber="509" refString="Neres, P. F., da Fonseca-Genevois, V. G., Torres, R. A., Cavalcanti, M. F., de Castro, F. J. V., da Silva, N. R. R., Rieger, T. T. &amp; Decraemer, W. (2010) Morphological and molecular taxonomy of a new Daptonema (Nematoda, Xyalidae) with comments on the systematics of some related taxa. Zoological Journal of the Linnean Society, 158, 1 - 15. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00528. x" type="journal article">
Neres
<emphasis id="7CC4A825BA7F9B49DCDFFD313687FD00" box="[1156,1213,726,751]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
(2010)
</bibRefCitation>
suggest that
<taxonomicName id="89B00FB4BA7F9B49D8CCFD1A3320FCFC" box="[151,282,764,787]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49D8CCFD1A3320FCFC" box="[151,282,764,787]" italics="true" pageId="8" pageNumber="509">Daptonema</emphasis>
</taxonomicName>
is paraphyletic, with some species of the genus more closely related to
<taxonomicName id="89B00FB4BA7F9B49DC22FD1C36DBFCFC" box="[1145,1249,762,787]" class="Adenophorea" family="Monhysteridae" genus="Theristus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49DC22FD1C36DBFCFC" box="[1145,1249,762,787]" italics="true" pageId="8" pageNumber="509">Theristus</emphasis>
</taxonomicName>
, and species of
<taxonomicName id="89B00FB4BA7F9B49D8CCFCF83359FCD8" box="[151,355,798,823]" class="Adenophorea" family="Monhysteridae" genus="Metadesmolaimus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49D8CCFCF83359FCD8" box="[151,355,798,823]" italics="true" pageId="8" pageNumber="509">Metadesmolaimus</emphasis>
</taxonomicName>
grouped with
<taxonomicName id="89B00FB4BA7F9B49DA7BFCF93099FCD9" box="[544,675,799,822]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49DA7BFCF93099FCD9" box="[544,675,799,822]" italics="true" pageId="8" pageNumber="509">Daptonema</emphasis>
</taxonomicName>
. These three genera are similar morphologically;
<taxonomicName id="89B00FB4BA7F9B49DD4AFCF83743FCD8" box="[1297,1401,798,823]" class="Adenophorea" family="Monhysteridae" genus="Theristus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49DD4AFCF83743FCD8" box="[1297,1401,798,823]" italics="true" pageId="8" pageNumber="509">Theristus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="89B00FB4BA7F9B49D92FFCA233CCFCB4" box="[372,502,836,859]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49D92FFCA233CCFCB4" box="[372,502,836,859]" italics="true" pageId="8" pageNumber="509">Daptonema</emphasis>
</taxonomicName>
based solely on tail shape (conical without terminal setae) (
<bibRefCitation id="2A2109C6BA7F9B49DCFDFCA437AAFCB4" author="Venekey" box="[1190,1424,834,859]" pageId="8" pageNumber="509" refString="Venekey, V., Gheller, P. F., Maria, T. F., Brustolin, M. C., Kandratavicius, N., Vieira, D. C., Brito, S., Souza, G. S. &amp; Fonseca, G. (2014) The state of the art of Xyalidae (Nematoda, Monhysterida) with reference to the Brazilian records. Marine Biodiversity, 44, 367 - 390. http: // dx. doi. org / 10.1007 / s 12526 - 014 - 0226 - 3" type="journal article" year="2014">
Venekey
<emphasis id="7CC4A825BA7F9B49DD4AFCA23771FCB4" box="[1297,1355,834,859]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
2014
</bibRefCitation>
), whereas
<taxonomicName id="89B00FB4BA7F9B49D8A1FC8033FCFC90" box="[250,454,870,895]" class="Adenophorea" family="Monhysteridae" genus="Metadesmolaimus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49D8A1FC8033FCFC90" box="[250,454,870,895]" italics="true" pageId="8" pageNumber="509">Metadesmolaimus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="89B00FB4BA7F9B49DA9BFC813179FC91" box="[704,835,871,894]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49DA9BFC813179FC91" box="[704,835,871,894]" italics="true" pageId="8" pageNumber="509">Daptonema</emphasis>
</taxonomicName>
based on presence of setose inner labial sensillae and extended, cylindrical anterior buccal cavity (
<bibRefCitation id="2A2109C6BA7F9B49DAD0FC6C3139FC4C" author="Platt" box="[651,771,906,931]" pageId="8" pageNumber="509" refString="Platt, H. M. (1983) New species of Metadesmolaimus and Stephanolaimus (Nematoda: Chromadoria) from Northern Ireland with reviews of the genera. Zoological Journal of the Linnean Society, 78, 363 - 373. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1975. tb 02265. x" type="journal article" year="1983">Platt 1983</bibRefCitation>
). In some
<taxonomicName id="89B00FB4BA7F9B49DB2CFC6C3679FC4C" box="[887,1091,906,931]" class="Adenophorea" family="Monhysteridae" genus="Metadesmolaimus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49DB2CFC6C3679FC4C" box="[887,1091,906,931]" italics="true" pageId="8" pageNumber="509">Metadesmolaimus</emphasis>
</taxonomicName>
species, the cylindrical part of the anterior buccal cavity is very thin (e.g.,
<taxonomicName id="89B00FB4BA7F9B49DAC2FC4931D5FC28" authority="Gerlach, 1953" authorityName="Gerlach" authorityYear="1953" box="[665,1007,942,967]" class="Adenophorea" family="Monhysteridae" genus="Metadesmolaimus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="species" species="aversivulva">
<emphasis id="7CC4A825BA7F9B49DAC2FC493179FC28" box="[665,835,942,967]" italics="true" pageId="8" pageNumber="509">M. aversivulva</emphasis>
Gerlach, 1953
</taxonomicName>
and
<taxonomicName id="89B00FB4BA7F9B49DC77FC4932DEFC04" authority="Wieser &amp; Hopper, 1967" authorityName="Wieser &amp; Hopper" authorityYear="1967" class="Adenophorea" family="Monhysteridae" genus="Metadesmolaimus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="species" species="canicula">
<emphasis id="7CC4A825BA7F9B49DC77FC49368CFC28" box="[1068,1206,942,967]" italics="true" pageId="8" pageNumber="509">M. canicula</emphasis>
(Wieser &amp; Hopper, 1967))
</taxonomicName>
, and some
<taxonomicName id="89B00FB4BA7F9B49D932FC3233D6FC04" box="[361,492,980,1003]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49D932FC3233D6FC04" box="[361,492,980,1003]" italics="true" pageId="8" pageNumber="509">Daptonema</emphasis>
</taxonomicName>
species possess a buccal cavity with a small, cuticularised extension (cheilostom, e.g.,
<taxonomicName id="89B00FB4BA7F9B49D894FC1133D1FBE0" authority="Gerlach, 1965" authorityName="Gerlach" authorityYear="1965" box="[207,491,1014,1039]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="species" species="biggi">
<emphasis id="7CC4A825BA7F9B49D894FC113317FBE0" box="[207,301,1014,1039]" italics="true" pageId="8" pageNumber="509">D. biggi</emphasis>
(Gerlach, 1965)
</taxonomicName>
,
<taxonomicName id="89B00FB4BA7F9B49D9A2FC1130ACFBE0" box="[505,662,1014,1039]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="species" species="amphorum" status="sp. nov.">
<emphasis id="7CC4A825BA7F9B49D9A2FC1130ACFBE0" box="[505,662,1014,1039]" italics="true" pageId="8" pageNumber="509">D. amphorum</emphasis>
</taxonomicName>
<emphasis id="7CC4A825BA7F9B49DAC4FC1030E5FBE0" bold="true" box="[671,735,1014,1039]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="67F7155EBA7F9B49DAC4FC1030E5FBE0" box="[671,735,1014,1039]" pageId="8" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) so that the distinction between the two genera is sometimes based mainly on the presence of setose (
<taxonomicName id="89B00FB4BA7F9B49DA2EFBFC317BFBDC" box="[629,833,1050,1075]" class="Adenophorea" family="Monhysteridae" genus="Metadesmolaimus" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49DA2EFBFC317BFBDC" box="[629,833,1050,1075]" italics="true" pageId="8" pageNumber="509">Metadesmolaimus</emphasis>
</taxonomicName>
)
<emphasis id="7CC4A825BA7F9B49DB0EFBFA3156FBDC" box="[853,876,1052,1075]" italics="true" pageId="8" pageNumber="509">vs</emphasis>
papillose (
<taxonomicName id="89B00FB4BA7F9B49DBAAFBFA364EFBDC" box="[1009,1140,1052,1075]" class="Chromadorea" family="Xyalidae" genus="Daptonema" higherTaxonomySource="CoL" kingdom="Animalia" order="Monhysterida" pageId="8" pageNumber="509" phylum="Nematoda" rank="genus">
<emphasis id="7CC4A825BA7F9B49DBAAFBFA364EFBDC" box="[1009,1140,1052,1075]" italics="true" pageId="8" pageNumber="509">Daptonema</emphasis>
</taxonomicName>
) inner labial sensillae. A revision of these genera based on further morphological and molecular data is needed; such a revision would likely result in changes in the assignment of species.
</paragraph>
</subSubSection>
</treatment>
</document>