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<document id="5A9ED085B1AC60198FCA38ABBB5EDDF0" ID-CLB-Dataset="46817" ID-DOI="10.5281/zenodo.202031" ID-GBIF-Dataset="ddc05270-1333-499e-8b62-45e157183cfb" ID-ISSN="1175-5326" ID-Zenodo-Dep="202031" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460360659916" checkinUser="plazi" docAuthor="Hekmatara, Maryam, Sari, Alireza &amp; Baladehi, Mohammad-Hadi Heidary" docDate="2011" docId="A17F87A0FFB4FFEEF8B8F89AFDA6D107" docLanguage="en" docName="zt02894p057.pdf" docOrigin="Zootaxa 2894" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gammarus hegmatanensis Hekmatara, Sari &amp; Baladehi, 2011, sp. n." docType="treatment" docVersion="11" lastPageNumber="49" masterDocId="5D46FFD8FFB5FFE4F82FFFD3FFF1D063" masterDocTitle="Two new Gammarus species (Crustacea: Amphipoda: Gammaridae) from Zagros Mountains, Iran" masterLastPageNumber="57" masterPageNumber="39" pageNumber="40" updateTime="1698294442888" updateUser="ExternalLinkService">
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<mods:title id="1E2C647A2ED97A6D15BDFA99A8FBD4C1">Two new Gammarus species (Crustacea: Amphipoda: Gammaridae) from Zagros Mountains, Iran</mods:title>
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<mods:namePart id="58F98A9992A36D2D0F0D88A5EC870CC5">Hekmatara, Maryam</mods:namePart>
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<mods:namePart id="6C8255D9FDC8BF2E87FA93873BA1F6C9">Sari, Alireza</mods:namePart>
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<mods:namePart id="C393E5E10EFB8416CA7A9DFCBE9DD89A">Baladehi, Mohammad-Hadi Heidary</mods:namePart>
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<mods:date id="5A46C1848BD02D1B3C83DA74DDDCBCD4">2011</mods:date>
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<treatment id="A17F87A0FFB4FFEEF8B8F89AFDA6D107" ID-DOI="http://doi.org/10.5281/zenodo.3501095" ID-GBIF-Taxon="119409622" ID-Zenodo-Dep="3501095" LSID="urn:lsid:plazi:treatment:A17F87A0FFB4FFEEF8B8F89AFDA6D107" httpUri="http://treatment.plazi.org/id/A17F87A0FFB4FFEEF8B8F89AFDA6D107" lastPageId="10" lastPageNumber="49" pageId="1" pageNumber="40">
<subSubSection id="61CC653DFFB4FFE5F8B8F89AFEE3D7E7" pageId="1" pageNumber="40" type="nomenclature">
<paragraph id="296936B6FFB4FFE5F8B8F89AFDD2D700" blockId="1.[151,547,1865,1924]" box="[151,547,1865,1891]" pageId="1" pageNumber="40">
<heading id="722181DAFFB4FFE5F8B8F89AFDD2D700" bold="true" box="[151,547,1865,1891]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
<emphasis id="1BA2EAA4FFB4FFE5F8B8F89AFDD2D700" bold="true" box="[151,547,1865,1891]" pageId="1" pageNumber="40">
<taxonomicName id="EED64D35FFB4FFE5F8B8F89AFE28D700" ID-CoL="3F8CV" box="[151,473,1865,1891]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="hegmatanensis" status="sp. nov.">
<emphasis id="1BA2EAA4FFB4FFE5F8B8F89AFE28D700" bold="true" box="[151,473,1865,1891]" italics="true" pageId="1" pageNumber="40">Gammarus hegmatanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="009157DFFFB4FFE5F9CEF89AFDD2D700" box="[481,547,1865,1891]" pageId="1" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="296936B6FFB4FFE5F8B8F8BFFEE3D7E7" blockId="1.[151,547,1865,1924]" box="[151,274,1900,1924]" pageId="1" pageNumber="40">
(
<figureCitation id="B1ED2A33FFB4FFE5F8B0F8BFFEF8D7E7" box="[159,265,1900,1924]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="2.[151,249,1977,2000]" captionStartId-1="4.[151,250,1961,1984]" captionStartId-2="5.[151,250,1949,1972]" captionStartId-3="6.[151,249,1803,1826]" captionStartId-4="7.[151,249,1875,1898]" captionStartId-5="8.[151,249,1825,1848]" captionTargetBox-0="[230,1281,448,1957]" captionTargetBox-1="[200,1374,208,1845]" captionTargetBox-2="[152,1425,198,1948]" captionTargetBox-3="[152,1432,225,1802]" captionTargetBox-4="[309,1242,459,1739]" captionTargetBox-5="[152,1428,204,1824]" captionTargetId-0="figure@2.[151,1436,427,1973]" captionTargetId-1="figure@4.[151,1436,111,1940]" captionTargetId-2="figure@5.[151,1436,109,2008]" captionTargetId-3="figure@6.[151,1436,61,1828]" captionTargetId-4="figure@7.[151,1436,391,1873]" captionTargetId-5="figure@8.[151,1436,121,1900]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 2. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv." captionText-1="FIGURE 3. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (GN 1) and C (GN 2) on scale i; B (PGN 1) and D (PGN 1) on scale ii; E (P 3) on scale iii; F (dactyl of P 3) on scale iv." captionText-2="FIGURE 4. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 4), B (P 5), C (P 6) and D (P 7) on scale i; E (US) on scale ii; F (dactyl of P 4), G (dactyl of P 5), H (dactyl of P 6) and I (dactyl of P 7) on scale iii." captionText-3="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." captionText-4="FIGURE 6. Gammarus hegmatanensis sp. n., Allotype, Ƥ, 15.1 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (P 3) and D (P 4) on scale ii; E (dactyl of P 3) and F (dactyl of P 4) on scale iii." captionText-5="FIGURE 7. Gammarus hegmatanensis sp. n., Allotype, Ƥ, 15.1 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 5), B (P 6), C (P 7) and D (O) on scale i; E (U 3) on scale ii; F (dactyl of P 5), G (dactyl of P 6) and H (dactyl of 7) on scale iii." httpUri-0="https://zenodo.org/record/202033/files/figure.png" httpUri-1="https://zenodo.org/record/202034/files/figure.png" httpUri-2="https://zenodo.org/record/202035/files/figure.png" httpUri-3="https://zenodo.org/record/202036/files/figure.png" httpUri-4="https://zenodo.org/record/202037/files/figure.png" httpUri-5="https://zenodo.org/record/202038/files/figure.png" pageId="1" pageNumber="40">Figs. 27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61CC653DFFB4FFE6F8B8F867FEC5D1E4" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40" type="materials_examined">
<paragraph id="296936B6FFB4FFE5F8B8F867FE5DD793" blockId="1.[151,1436,1971,2032]" pageId="1" pageNumber="40">
<emphasis id="1BA2EAA4FFB4FFE5F8B8F867FEC2D7AF" bold="true" box="[151,307,1971,1996]" pageId="1" pageNumber="40">
<typeStatus id="F66D8814FFB4FFE5F8B8F867FF21D7AF" box="[151,208,1972,1996]" pageId="1" pageNumber="40">Type</typeStatus>
locality.
</emphasis>
Gardaneh Asadabad spring, W of Hamadan city, Hamadan Province,
<collectingCountry id="51C17626FFB4FFE5FC6FF866FB9CD7AF" box="[1088,1133,1973,1996]" name="Iran" pageId="1" pageNumber="40">Iran</collectingCountry>
(N 34º 49ʹ, E 48º 0 4ʹ), altitude
<quantity id="EE2E9B53FFB4FFE5F8E1F80BFED2D793" box="[206,291,2008,2032]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="1" pageNumber="40" unit="m" value="1677.0">1677 m</quantity>
(locality 1).
</paragraph>
<paragraph id="296936B6FFB7FFE6F8EAFF44FA8ED15C" blockId="2.[151,1436,151,391]" pageId="2" pageNumber="41">
<emphasis id="1BA2EAA4FFB7FFE6F8EAFF44FE41D0D3" bold="true" box="[197,432,151,176]" pageId="2" pageNumber="41">Material examined.</emphasis>
One male (
<typeStatus id="F66D8814FFB7FFE6FA19FF44FD6FD0CC" box="[566,670,151,175]" pageId="2" pageNumber="41" type="holotype">holotype</typeStatus>
), body length
<quantity id="EE2E9B53FFB7FFE6FB6FFF44FC65D0CC" box="[832,916,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="41" unit="mm" value="18.0">18 mm</quantity>
, one female (allotype), body length
<quantity id="EE2E9B53FFB7FFE6FD1FFF44FA69D0CC" box="[1328,1432,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="2" pageNumber="41" unit="mm" value="15.1">15.1 mm</quantity>
, few
<typeStatus id="F66D8814FFB7FFE6F8E7FF6EFEC6D0B6" box="[200,311,189,213]" pageId="2" pageNumber="41" type="paratype">paratypes</typeStatus>
.
<collectionCode id="4FC7AE73FFB7FFE6F96DFF6FFE7FD0B7" box="[322,398,188,212]" pageId="2" pageNumber="41">ZUTC</collectionCode>
, Amph. 2249,
<date id="5D681076FFB7FFE6FA1BFF6FFD1ED0B7" box="[564,751,188,212]" pageId="2" pageNumber="41" value="2007-09">September 2007</date>
, collected by M. Hekmatara. Additional samples: Chavaneh- Lar qanat and spring, Hame-kasi village, 41 Km NW of Hamadan (N 34º 57ʹ, E 48º 0 9ʹ), altitude
<quantity id="EE2E9B53FFB7FFE6FCCDFF0CFAC6D094" box="[1250,1335,223,247]" metricMagnitude="3" metricUnit="m" metricValue="2.109" pageId="2" pageNumber="41" unit="m" value="2109.0">2109 m</quantity>
(locality 2, 3). River of Ali-Abad village, Hamadan-Ghorveh road, 30 Km NW of Hamadan, (N 34 º 56ʹ, E 48 º 15ʹ), altitude 1830 (locality 4). Descriptions were based on the
<typeStatus id="F66D8814FFB7FFE6FAEBFEF4FCD7D15C" box="[708,806,295,319]" pageId="2" pageNumber="41" type="holotype">holotype</typeStatus>
and were compared to some
<typeStatus id="F66D8814FFB7FFE6FC41FEFBFB28D123" box="[1134,1241,296,320]" pageId="2" pageNumber="41" type="paratype">paratypes</typeStatus>
for variations.
</paragraph>
<paragraph id="296936B6FFB7FFE6F8EAFE9FFEC5D1E4" blockId="2.[151,1436,151,391]" pageId="2" pageNumber="41">
All specimens including
<typeStatus id="F66D8814FFB7FFE6F9F4FE9FFDCCD107" box="[475,573,332,356]" pageId="2" pageNumber="41" type="holotype">holotype</typeStatus>
are deposited in the Zoological Museum of the University of Tehran (
<collectionCode id="4FC7AE73FFB7FFE6FD65FE9FFA66D107" box="[1354,1431,332,356]" pageId="2" pageNumber="41">ZUTC</collectionCode>
, Amph. 2249).
</paragraph>
</subSubSection>
<caption id="7DA9663EFFB7FFE6F8B8F86AFB59D78D" httpUri="https://zenodo.org/record/202033/files/figure.png" pageId="2" pageNumber="41" targetBox="[230,1281,448,1957]" targetPageId="2">
<paragraph id="296936B6FFB7FFE6F8B8F86AFB59D78D" blockId="2.[151,1436,1977,2031]" pageId="2" pageNumber="41">
<emphasis id="1BA2EAA4FFB7FFE6F8B8F86AFEE3D7AC" bold="true" box="[151,274,1977,2000]" pageId="2" pageNumber="41">FIGURE 2.</emphasis>
<taxonomicName id="EED64D35FFB7FFE6F936F869FDD6D7AC" box="[281,551,1977,1999]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="hegmatanensis" status="sp. nov.">
<emphasis id="1BA2EAA4FFB7FFE6F936F869FDD6D7AC" box="[281,551,1977,1999]" italics="true" pageId="2" pageNumber="41">Gammarus hegmatanensis</emphasis>
</taxonomicName>
<emphasis id="1BA2EAA4FFB7FFE6FA02F869FD96D7AC" bold="true" box="[557,615,1978,1999]" pageId="2" pageNumber="41">
<taxonomicNameLabel id="009157DFFFB7FFE6FA02F869FD96D7AC" box="[557,615,1978,1999]" pageId="2" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A1) and B (A2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv.
</paragraph>
</caption>
<subSubSection id="61CC653DFFB6FFE7F8EAFF44FA67D15C" pageId="3" pageNumber="42" type="diagnosis">
<paragraph id="296936B6FFB6FFE7F8EAFF44FA67D15C" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFF44FECED0D3" bold="true" box="[197,319,151,176]" pageId="3" pageNumber="42">Diagnosis.</emphasis>
A mediumlarge species; it differs from other similar taxa by following distinct characters: the most remarkable feature of this species is the shape of epimeral plate 3, with a very sharply pointed and extended posterior corner and a distinct lobate anterior corner; no protruding lobe at postero-distal corner of pereopod 7 basis; antenna 2 flagellum armed with numerous long setae, the fifth peduncular article longer than the fourth and highly setose; pereopods 57 posterior margin poorly setose; uropod 3 endopodite length about 80% of exopodite.
</paragraph>
</subSubSection>
<subSubSection id="61CC653DFFB6FFE7F8EAFE9FFBE6D1E4" pageId="3" pageNumber="42" type="etymology">
<paragraph id="296936B6FFB6FFE7F8EAFE9FFBE6D1E4" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFE9FFEBAD107" bold="true" box="[197,331,332,356]" pageId="3" pageNumber="42">Etymology.</emphasis>
The epithet of
<taxonomicName id="EED64D35FFB6FFE7F9DAFE9FFD6DD107" box="[501,668,332,356]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="hegmatanensis">
<emphasis id="1BA2EAA4FFB6FFE7F9DAFE9FFD6DD107" box="[501,668,332,356]" italics="true" pageId="3" pageNumber="42">hegmatanensis</emphasis>
</taxonomicName>
derived from “Hegmataneh”; the ancient city whose old remains are found within Hamadan city. The
<typeStatus id="F66D8814FFB6FFE7FA27FEA3FDC9D1EB" box="[520,568,368,392]" pageId="3" pageNumber="42">type</typeStatus>
locality is in the vicinity of Hamadan city.
</paragraph>
</subSubSection>
<subSubSection id="61CC653DFFB6FFEDF8EAFE40FD05D2E3" lastPageId="9" lastPageNumber="48" pageId="3" pageNumber="42" type="description">
<paragraph id="296936B6FFB6FFE7F8EAFE40FEE2D197" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFE40FE58D1CF" bold="true" box="[197,425,403,428]" pageId="3" pageNumber="42">Description. Male:</emphasis>
Body length
<quantity id="EE2E9B53FFB6FFE7FA7CFE47FD5DD1CF" box="[595,684,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="42" unit="mm" value="18.0">18 mm</quantity>
. Head: Lateral head lobe rounded, anterior lower part of head extended forward; eyes elliptical to slightly reniform, smaller than diameter of first peduncular article of antenna 1 (
<figureCitation id="B1ED2A33FFB6FFE7F8B0FE0FFF15D197" box="[159,228,476,500]" captionStart="FIGURE 2" captionStartId="2.[151,249,1977,2000]" captionTargetBox="[230,1281,448,1957]" captionTargetId="figure@2.[151,1436,427,1973]" captionTargetPageId="2" captionText="FIGURE 2. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv." httpUri="https://zenodo.org/record/202033/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
, C).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAFE2CFC5ED25F" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFE2CFEB0D27B" bold="true" box="[197,321,511,536]" pageId="3" pageNumber="42">Antenna 1</emphasis>
: Longer than antenna 2, flagellum with 2527 articles, armed with very short simple setae, peduncular articles 1&gt;2&gt;3, accessory flagellum with four articles (
<figureCitation id="B1ED2A33FFB6FFE7FB15FDF7FC8FD25F" box="[826,894,548,572]" captionStart="FIGURE 2" captionStartId="2.[151,249,1977,2000]" captionTargetBox="[230,1281,448,1957]" captionTargetId="figure@2.[151,1436,427,1973]" captionTargetPageId="2" captionText="FIGURE 2. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv." httpUri="https://zenodo.org/record/202033/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
, A).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAFD94FE1DD2AF" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFD94FEDDD203" bold="true" box="[197,300,583,608]" pageId="3" pageNumber="42">Antenna</emphasis>
2: Fifth peduncular article longer than the fourth, highly setose; flagellum with 810 articles, armed with long simple setae. Their size twice or longer than the diameter of the corresponding antennal articles; length of the first flagellar article two times the second; gland cone is pointed, shorter than the third peduncular article; calceoli not present (
<figureCitation id="B1ED2A33FFB6FFE7F9AFFD67FE4CD2AF" box="[384,445,692,716]" captionStart="FIGURE 2" captionStartId="2.[151,249,1977,2000]" captionTargetBox="[230,1281,448,1957]" captionTargetId="figure@2.[151,1436,427,1973]" captionTargetPageId="2" captionText="FIGURE 2. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv." httpUri="https://zenodo.org/record/202033/files/figure.png" pageId="3" pageNumber="42">Fig.2</figureCitation>
, B).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAFD04FE26D33F" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFD04FE85D293" bold="true" box="[197,372,727,752]" pageId="3" pageNumber="42">Mandible palp</emphasis>
: First article without setae; second segment with ventral setae; 67 proximal setae shorter than the width of the segment and 810 closely placed distal setae longer than the width of the segment; inferior margin of the third segment armed with a comb-like row of 3237 D-setae, 56 long E-setae, one group of B-setae and one group of A-setae (
<figureCitation id="B1ED2A33FFB6FFE7F94DFC97FE56D33F" box="[354,423,836,860]" captionStart="FIGURE 2" captionStartId="2.[151,249,1977,2000]" captionTargetBox="[230,1281,448,1957]" captionTargetId="figure@2.[151,1436,427,1973]" captionTargetPageId="2" captionText="FIGURE 2. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv." httpUri="https://zenodo.org/record/202033/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
, D).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAFCB4FEE0D38F" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFCB4FEC7D3E3" bold="true" box="[197,310,871,896]" pageId="3" pageNumber="42">Maxilla 1</emphasis>
: Palps asymmetric; left palp with seven median spines, one subapical spine on its outer margin and three short setae (
<figureCitation id="B1ED2A33FFB6FFE7F972FC5FFE50D3C7" box="[349,417,908,932]" captionStart="FIGURE 2" captionStartId="2.[151,249,1977,2000]" captionTargetBox="[230,1281,448,1957]" captionTargetId="figure@2.[151,1436,427,1973]" captionTargetPageId="2" captionText="FIGURE 2. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv." httpUri="https://zenodo.org/record/202033/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
, F); right palp with five robust tooth-like spines, one longer separate subapical spine on its outer margin with one short seta opposed by one smaller spine, anterodistally; lateral margin with two short setae (
<figureCitation id="B1ED2A33FFB6FFE7F8B0FC07FF15D38F" box="[159,228,980,1004]" captionStart="FIGURE 2" captionStartId="2.[151,249,1977,2000]" captionTargetBox="[230,1281,448,1957]" captionTargetId="figure@2.[151,1436,427,1973]" captionTargetPageId="2" captionText="FIGURE 2. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (H) on scale ii; D (MNDP) on scale iii; E (PRMX) and F (PLMX) on scale iv." httpUri="https://zenodo.org/record/202033/files/figure.png" pageId="3" pageNumber="42">Fig. 2</figureCitation>
, E).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAFC24FB9DD4A7" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFC24FE90D473" bold="true" box="[197,353,1015,1040]" pageId="3" pageNumber="42">Gnathopod 1</emphasis>
: Coxal plate distally not or slightly wider than proximally, rounded corners with a postero-distal seta and two to three antero-distal setae. Basis with long simple setae on anterior and posterior margins. Ischium with a postero-distal row of setae. Carpus triangular and anterior margin with two groups of simple setae, posteriorly with plumose rows of setae. Propodus poorly setose, armed with one mid-palmar and one palmar angle spine, six spines at posterior palmar margin, several spines on posterior sub-margin, two groups of setae on propodal surface, one group of setae on the propodal anterior margin; dactylus long (
<figureCitation id="B1ED2A33FFB6FFE7FBEEFB7FFBF9D4A7" box="[961,1032,1196,1220]" captionStart="FIGURE 3" captionStartId="4.[151,250,1961,1984]" captionTargetBox="[200,1374,208,1845]" captionTargetId="figure@4.[151,1436,111,1940]" captionTargetPageId="4" captionText="FIGURE 3. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (GN 1) and C (GN 2) on scale i; B (PGN 1) and D (PGN 1) on scale ii; E (P 3) on scale iii; F (dactyl of P 3) on scale iv." httpUri="https://zenodo.org/record/202034/files/figure.png" pageId="3" pageNumber="42">Figs.3</figureCitation>
, A &amp; B).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAFB01FA85D519" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFB01FE93D488" bold="true" box="[197,354,1234,1259]" pageId="3" pageNumber="42">Gnathopod 2</emphasis>
: Distal end of coxal plate finely wider than proximal, rounded corners with a seta postero-ventrally and three setae antero-ventrally. Basis with long setae on anterior and posterior margins. Ischium with postero-distal row of setae. Carpus triangular and anterior margin with two groups of setae, posteriorly with plumose rows of setae. Propodus densely setose, palm subrectangular with one mid-palmar spine, 23 palmar angle spines, five groups of setae on propodal surface, three groups of setae at the propodal anterior margin (
<figureCitation id="B1ED2A33FFB6FFE7FCEFFAB1FAE1D519" box="[1216,1296,1378,1402]" captionStart="FIGURE 3" captionStartId="4.[151,250,1961,1984]" captionTargetBox="[200,1374,208,1845]" captionTargetId="figure@4.[151,1436,111,1940]" captionTargetPageId="4" captionText="FIGURE 3. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (GN 1) and C (GN 2) on scale i; B (PGN 1) and D (PGN 1) on scale ii; E (P 3) on scale iii; F (dactyl of P 3) on scale iv." httpUri="https://zenodo.org/record/202034/files/figure.png" pageId="3" pageNumber="42">Figs. 3</figureCitation>
, C &amp; D).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAFA59FB46D636" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAFA59FEBCD5C1" bold="true" box="[197,333,1417,1442]" pageId="3" pageNumber="42">Pereopod 3</emphasis>
: Coxal plate rectangular and rounded distally; with four very short setae antero-distally and one postero-distally; basis with some long simple setae; basiscarpus bear groups of long setae at posterior margin; posterior margin of merus with several groups of dense setae and anterior margin with two groups of one (or none) short seta and one spine and a group of long setae and a spine antero-distally; carpus with three groups of spines and 56 groups of setae on posterior margin and a long spine and a group of longer setae implanted on both its antero and postero-distal; propodus with five groups of spines and some long setae (
<figureCitation id="B1ED2A33FFB6FFE7FC6AF9EEFB78D636" box="[1093,1161,1597,1621]" captionStart="FIGURE 4" captionStartId="5.[151,250,1949,1972]" captionTargetBox="[152,1425,198,1948]" captionTargetId="figure@5.[151,1436,109,2008]" captionTargetPageId="5" captionText="FIGURE 4. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 4), B (P 5), C (P 6) and D (P 7) on scale i; E (US) on scale ii; F (dactyl of P 4), G (dactyl of P 5), H (dactyl of P 6) and I (dactyl of P 7) on scale iii." httpUri="https://zenodo.org/record/202035/files/figure.png" pageId="3" pageNumber="42">Fig. 4</figureCitation>
, E).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAF9B1FDB1D76A" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAF9B1FEBBD619" bold="true" box="[197,330,1633,1658]" pageId="3" pageNumber="42">Pereopod 4</emphasis>
: Coxal plate wide and emarginated with three small setae on antero-distal margin and several setae on postero-distal margin; segments similar to pereopod 3, but setae are much shorter and the number of setae and groups is lower; anterior margin of merus with just one spine, one spine implanted antero-distally among a group of simple setae; carpus has four groups of spines among most groups of short setae; propodus with five groups of spines and few short setae (
<figureCitation id="B1ED2A33FFB6FFE7F9E5F922FDFED76A" box="[458,527,1777,1801]" captionStart="FIGURE 4" captionStartId="5.[151,250,1949,1972]" captionTargetBox="[152,1425,198,1948]" captionTargetId="figure@5.[151,1436,109,2008]" captionTargetPageId="5" captionText="FIGURE 4. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 4), B (P 5), C (P 6) and D (P 7) on scale i; E (US) on scale ii; F (dactyl of P 4), G (dactyl of P 5), H (dactyl of P 6) and I (dactyl of P 7) on scale iii." httpUri="https://zenodo.org/record/202035/files/figure.png" pageId="3" pageNumber="42">Fig. 4</figureCitation>
, A).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAF8C5FE03D7FA" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAF8C5FEBAD74D" bold="true" box="[197,331,1814,1838]" pageId="3" pageNumber="42">Pereopod 5</emphasis>
: Basis subrectangular, postero-distal corner rounded, posterior margin with 11 short setae, anterior margin with 45 spines, anterior margin of merus with three groups of short setae (without spine) and posterior margin with one spine; carpus with two groups of anterior spines and 03 short setae; propodus bears five groups of spines anteriorly (
<figureCitation id="B1ED2A33FFB6FFE7F950F852FE32D7FA" box="[383,451,1921,1945]" captionStart="FIGURE 4" captionStartId="5.[151,250,1949,1972]" captionTargetBox="[152,1425,198,1948]" captionTargetId="figure@5.[151,1436,109,2008]" captionTargetPageId="5" captionText="FIGURE 4. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 4), B (P 5), C (P 6) and D (P 7) on scale i; E (US) on scale ii; F (dactyl of P 4), G (dactyl of P 5), H (dactyl of P 6) and I (dactyl of P 7) on scale iii." httpUri="https://zenodo.org/record/202035/files/figure.png" pageId="3" pageNumber="42">Fig. 4</figureCitation>
, B).
</paragraph>
<paragraph id="296936B6FFB6FFE7F8EAF875FC51D782" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="42">
<emphasis id="1BA2EAA4FFB6FFE7F8EAF875FEBCD7DD" bold="true" box="[197,333,1957,1982]" pageId="3" pageNumber="42">Pereopod 6</emphasis>
: Longer than pereopod 5; basis slender and posterior margin with 10 setae and anterior margin with 23 spines; other segments are similar to pereopod 5 (
<figureCitation id="B1ED2A33FFB6FFE7FB03F81AFC80D782" box="[812,881,1993,2017]" captionStart="FIGURE 4" captionStartId="5.[151,250,1949,1972]" captionTargetBox="[152,1425,198,1948]" captionTargetId="figure@5.[151,1436,109,2008]" captionTargetPageId="5" captionText="FIGURE 4. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 4), B (P 5), C (P 6) and D (P 7) on scale i; E (US) on scale ii; F (dactyl of P 4), G (dactyl of P 5), H (dactyl of P 6) and I (dactyl of P 7) on scale iii." httpUri="https://zenodo.org/record/202035/files/figure.png" pageId="3" pageNumber="42">Fig. 4</figureCitation>
, C).
</paragraph>
<caption id="7DA9663EFFB1FFE0F8B8F87AFB14D7BD" httpUri="https://zenodo.org/record/202034/files/figure.png" pageId="4" pageNumber="43" targetBox="[200,1374,208,1845]" targetPageId="4">
<paragraph id="296936B6FFB1FFE0F8B8F87AFB14D7BD" blockId="4.[151,1436,1961,2015]" pageId="4" pageNumber="43">
<emphasis id="1BA2EAA4FFB1FFE0F8B8F87AFEE3D7DC" bold="true" box="[151,274,1961,1984]" pageId="4" pageNumber="43">FIGURE 3.</emphasis>
<taxonomicName id="EED64D35FFB1FFE0F93DF878FDEFD7A3" box="[274,542,1962,1984]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="hegmatanensis" status="sp. nov.">
<emphasis id="1BA2EAA4FFB1FFE0F93DF878FDEFD7A3" box="[274,542,1962,1984]" italics="true" pageId="4" pageNumber="43">Gammarus hegmatanensis</emphasis>
</taxonomicName>
<emphasis id="1BA2EAA4FFB1FFE0FA0AF879FDACD7DC" bold="true" box="[549,605,1962,1983]" pageId="4" pageNumber="43">
<taxonomicNameLabel id="009157DFFFB1FFE0FA0AF879FDACD7DC" box="[549,605,1962,1983]" pageId="4" pageNumber="43" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (GN1) and C (GN2) on scale i; B (PGN1) and D (PGN1) on scale ii; E (P3) on scale iii; F (dactyl of P3) on scale iv.
</paragraph>
</caption>
<caption id="7DA9663EFFB0FFE1F8B8F84EFEFAD793" httpUri="https://zenodo.org/record/202035/files/figure.png" pageId="5" pageNumber="44" targetBox="[152,1425,198,1948]" targetPageId="5">
<paragraph id="296936B6FFB0FFE1F8B8F84EFEFAD793" blockId="5.[151,1436,1949,2033]" pageId="5" pageNumber="44">
<emphasis id="1BA2EAA4FFB0FFE1F8B8F84EFEE2D7D0" bold="true" box="[151,275,1949,1972]" pageId="5" pageNumber="44">FIGURE 4.</emphasis>
<taxonomicName id="EED64D35FFB0FFE1F93CF84DFDD1D7D0" box="[275,544,1949,1971]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="hegmatanensis" status="sp. nov.">
<emphasis id="1BA2EAA4FFB0FFE1F93CF84DFDD1D7D0" box="[275,544,1949,1971]" italics="true" pageId="5" pageNumber="44">Gammarus hegmatanensis</emphasis>
</taxonomicName>
<emphasis id="1BA2EAA4FFB0FFE1FA07F84DFD90D7D0" bold="true" box="[552,609,1950,1971]" pageId="5" pageNumber="44">
<taxonomicNameLabel id="009157DFFFB0FFE1FA07F84DFD90D7D0" box="[552,609,1950,1971]" pageId="5" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P4), B (P5), C (P6) and D (P7) on scale i; E (US) on scale ii; F (dactyl of P4), G (dactyl of P5), H (dactyl of P6) and I (dactyl of P7) on scale iii.
</paragraph>
</caption>
<caption id="7DA9663EFFB3FFE2F8B8F8D8FB53D723" httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="6" pageNumber="45" targetBox="[152,1432,225,1802]" targetPageId="6">
<paragraph id="296936B6FFB3FFE2F8B8F8D8FB53D723" blockId="6.[151,1436,1803,1856]" pageId="6" pageNumber="45">
<emphasis id="1BA2EAA4FFB3FFE2F8B8F8D8FEE2D742" bold="true" box="[151,275,1803,1826]" pageId="6" pageNumber="45">FIGURE 5.</emphasis>
<taxonomicName id="EED64D35FFB3FFE2F93CF8DFFDD0D742" box="[275,545,1803,1825]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="hegmatanensis" status="sp. nov.">
<emphasis id="1BA2EAA4FFB3FFE2F93CF8DFFDD0D742" box="[275,545,1803,1825]" italics="true" pageId="6" pageNumber="45">Gammarus hegmatanensis</emphasis>
</taxonomicName>
<emphasis id="1BA2EAA4FFB3FFE2FA07F8DFFD93D742" bold="true" box="[552,610,1804,1825]" pageId="6" pageNumber="45">
<taxonomicNameLabel id="009157DFFFB3FFE2FA07F8DFFD93D742" box="[552,610,1804,1825]" pageId="6" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U1) and B (U2) on scale i; C (U3) on scale ii; D (T) on scale iii; E (EP1), F (EP2) and G (EP3) on scale iv.
</paragraph>
</caption>
<paragraph id="296936B6FFB3FFE2F8EAF8BEFDBFD793" blockId="6.[151,1436,1900,2032]" pageId="6" pageNumber="45">
<emphasis id="1BA2EAA4FFB3FFE2F8EAF8BEFEBAD7E6" bold="true" box="[197,331,1901,1925]" pageId="6" pageNumber="45">Pereopod 7</emphasis>
: Basis wider proximally, no protruding lobe at postero-distal corner of basis, antero-proximal part of basis with three long setae and anterior margin with three spines, posterior margin with 14 short setae; merus anteriorly with 23 groups of spines and short setae; anterior margin of carpus and propodus with some groups of spines without simple setae (
<figureCitation id="B1ED2A33FFB3FFE2F9F6F80BFDEFD793" box="[473,542,2008,2032]" captionStart="FIGURE 4" captionStartId="5.[151,250,1949,1972]" captionTargetBox="[152,1425,198,1948]" captionTargetId="figure@5.[151,1436,109,2008]" captionTargetPageId="5" captionText="FIGURE 4. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 4), B (P 5), C (P 6) and D (P 7) on scale i; E (US) on scale ii; F (dactyl of P 4), G (dactyl of P 5), H (dactyl of P 6) and I (dactyl of P 7) on scale iii." httpUri="https://zenodo.org/record/202035/files/figure.png" pageId="6" pageNumber="45">Fig. 4</figureCitation>
, D).
</paragraph>
<paragraph id="296936B6FFB2FFE3F8EAFF44FEE0D0B7" blockId="7.[151,1436,151,356]" pageId="7" pageNumber="46">
<emphasis id="1BA2EAA4FFB2FFE3F8EAFF44FE7CD0D3" bold="true" box="[197,397,151,176]" pageId="7" pageNumber="46">Epimeral plate 1</emphasis>
: Antero-distal margin rounded with some long setae, posterior margin with three short setae (
<figureCitation id="B1ED2A33FFB2FFE3F8B0FF6FFF12D0B7" box="[159,227,188,212]" captionStart="FIGURE 5" captionStartId="6.[151,249,1803,1826]" captionTargetBox="[152,1432,225,1802]" captionTargetId="figure@6.[151,1436,61,1828]" captionTargetPageId="6" captionText="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="7" pageNumber="46">Fig. 5</figureCitation>
, E).
</paragraph>
<paragraph id="296936B6FFB2FFE3F8EAFF0CFF22D17F" blockId="7.[151,1436,151,356]" pageId="7" pageNumber="46">
<emphasis id="1BA2EAA4FFB2FFE3F8EAFF0CFE7DD09B" bold="true" box="[197,396,223,248]" pageId="7" pageNumber="46">Epimeral plate 2</emphasis>
: Antero-distal margin lobate, posterior margin moderately pointed with four short setae (
<figureCitation id="B1ED2A33FFB2FFE3FD40FF0CFF56D17F" captionStart="FIGURE 5" captionStartId="6.[151,249,1803,1826]" captionTargetBox="[152,1432,225,1802]" captionTargetId="figure@6.[151,1436,61,1828]" captionTargetPageId="6" captionText="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="7" pageNumber="46">Fig. 5</figureCitation>
, F).
</paragraph>
<paragraph id="296936B6FFB2FFE3F8EAFEF4FC00D107" blockId="7.[151,1436,151,356]" pageId="7" pageNumber="46">
<emphasis id="1BA2EAA4FFB2FFE3F8EAFEF4FE7CD123" bold="true" box="[197,397,295,320]" pageId="7" pageNumber="46">Epimeral plate 3</emphasis>
: With very sharply pointed posterior corner and markedly lobate antero-distal margin, some short setae on posterior margin and three spines on lower margin (
<figureCitation id="B1ED2A33FFB2FFE3FB54FE9FFC31D107" box="[891,960,332,356]" captionStart="FIGURE 5" captionStartId="6.[151,249,1803,1826]" captionTargetBox="[152,1432,225,1802]" captionTargetId="figure@6.[151,1436,61,1828]" captionTargetPageId="6" captionText="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="7" pageNumber="46">Fig. 5</figureCitation>
, G).
</paragraph>
<caption id="7DA9663EFFB2FFE3F8B8F880FB23D7EB" httpUri="https://zenodo.org/record/202037/files/figure.png" pageId="7" pageNumber="46" targetBox="[309,1242,459,1739]" targetPageId="7">
<paragraph id="296936B6FFB2FFE3F8B8F880FB23D7EB" blockId="7.[151,1436,1875,1929]" pageId="7" pageNumber="46">
<emphasis id="1BA2EAA4FFB2FFE3F8B8F880FEE4D70A" bold="true" box="[151,277,1875,1898]" pageId="7" pageNumber="46">FIGURE 6.</emphasis>
<taxonomicName id="EED64D35FFB2FFE3F932F886FDDCD709" box="[285,557,1876,1898]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="hegmatanensis" status="sp. nov.">
<emphasis id="1BA2EAA4FFB2FFE3F932F886FDDCD709" box="[285,557,1876,1898]" italics="true" pageId="7" pageNumber="46">Gammarus hegmatanensis</emphasis>
</taxonomicName>
<emphasis id="1BA2EAA4FFB2FFE3FA19F887FD80D70A" bold="true" box="[566,625,1876,1897]" pageId="7" pageNumber="46">
<taxonomicNameLabel id="009157DFFFB2FFE3FA19F887FD80D70A" box="[566,625,1876,1897]" pageId="7" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, Allotype, Ƥ, 15.1 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A1) and B (A2) on scale i; C (P3) and D (P4) on scale ii; E (dactyl of P3) and F (dactyl of P4) on scale iii.
</paragraph>
</caption>
<caption id="7DA9663EFFBDFFECF8B8F8F2FA62D735" httpUri="https://zenodo.org/record/202038/files/figure.png" pageId="8" pageNumber="47" targetBox="[152,1428,204,1824]" targetPageId="8">
<paragraph id="296936B6FFBDFFECF8B8F8F2FA62D735" blockId="8.[151,1436,1825,1878]" pageId="8" pageNumber="47">
<emphasis id="1BA2EAA4FFBDFFECF8B8F8F2FEE4D754" bold="true" box="[151,277,1825,1848]" pageId="8" pageNumber="47">FIGURE 7.</emphasis>
<taxonomicName id="EED64D35FFBDFFECF932F8F1FDDCD754" box="[285,557,1825,1847]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="hegmatanensis" status="sp. nov.">
<emphasis id="1BA2EAA4FFBDFFECF932F8F1FDDCD754" box="[285,557,1825,1847]" italics="true" pageId="8" pageNumber="47">Gammarus hegmatanensis</emphasis>
</taxonomicName>
<emphasis id="1BA2EAA4FFBDFFECFA19F8F1FD80D754" bold="true" box="[566,625,1826,1847]" pageId="8" pageNumber="47">
<taxonomicNameLabel id="009157DFFFBDFFECFA19F8F1FD80D754" box="[566,625,1826,1847]" pageId="8" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, Allotype, Ƥ, 15.1 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P5), B (P6), C (P7) and D (O) on scale i; E (U3) on scale ii; F (dactyl of P5), G (dactyl of P6) and H (dactyl of 7) on scale iii.
</paragraph>
</caption>
<paragraph id="296936B6FFBDFFECF8EAF851FA76D781" blockId="8.[151,1436,1922,2018]" pageId="8" pageNumber="47">
<emphasis id="1BA2EAA4FFBDFFECF8EAF851FE8CD7F8" bold="true" box="[197,381,1922,1947]" pageId="8" pageNumber="47">Urosomites 13</emphasis>
: With moderate dorsal elevation, urosomite 1 armed with four dorso-medial spines and three setae and two dorso-lateral spines and three setae; urosomite 2 with 23 dorso-medial spines and four setae and 1 2 dorso-lateral spines and two setae; urosomite 3 armed with three dorso-medial spines and 78 setae (
<figureCitation id="B1ED2A33FFBDFFECFD3AF819FAA8D781" box="[1301,1369,1994,2018]" captionStart="FIGURE 4" captionStartId="5.[151,250,1949,1972]" captionTargetBox="[152,1425,198,1948]" captionTargetId="figure@5.[151,1436,109,2008]" captionTargetPageId="5" captionText="FIGURE 4. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 4), B (P 5), C (P 6) and D (P 7) on scale i; E (US) on scale ii; F (dactyl of P 4), G (dactyl of P 5), H (dactyl of P 6) and I (dactyl of P 7) on scale iii." httpUri="https://zenodo.org/record/202035/files/figure.png" pageId="8" pageNumber="47">Fig. 4</figureCitation>
, E).
</paragraph>
<paragraph id="296936B6FFBCFFEDF8EAFF4BFCD0D0B0" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<emphasis id="1BA2EAA4FFBCFFEDF8EAFF4BFEC6D0D3" bold="true" box="[197,311,151,176]" pageId="9" pageNumber="48">Uropod 1</emphasis>
: Longer than uropod 2, mid-dorsal margin of peduncle with two spines, endopodite and exopodite with two mid-dorsal and some terminal spines (
<figureCitation id="B1ED2A33FFBCFFEDFA83FF68FD01D0B0" box="[684,752,187,211]" captionStart="FIGURE 5" captionStartId="6.[151,249,1803,1826]" captionTargetBox="[152,1432,225,1802]" captionTargetId="figure@6.[151,1436,61,1828]" captionTargetPageId="6" captionText="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="9" pageNumber="48">Fig. 5</figureCitation>
, A).
</paragraph>
<paragraph id="296936B6FFBCFFEDF8EAFF33FEAED178" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<emphasis id="1BA2EAA4FFBCFFEDF8EAFF33FEC8D09B" bold="true" box="[197,313,223,248]" pageId="9" pageNumber="48">Uropod 2</emphasis>
: Exopodite slightly shorter than endopodite, both rami with two mid-dorsal and some terminal spines (
<figureCitation id="B1ED2A33FFBCFFEDF8C3FED0FEC1D178" box="[236,304,259,283]" captionStart="FIGURE 5" captionStartId="6.[151,249,1803,1826]" captionTargetBox="[152,1432,225,1802]" captionTargetId="figure@6.[151,1436,61,1828]" captionTargetPageId="6" captionText="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="9" pageNumber="48">Fig. 5</figureCitation>
, B).
</paragraph>
<paragraph id="296936B6FFBCFFEDF8EAFEF4FE01D1E5" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<emphasis id="1BA2EAA4FFBCFFEDF8EAFEF4FEC6D15C" bold="true" box="[197,311,294,319]" pageId="9" pageNumber="48">Uropod 3</emphasis>
: Endopodite length about 80% of exopodite, distal article minute; outer margin of exopodite with simple setae, without any plumose setae; inner margin of endopodite and exopodite and outer margin of endopodite with plumose setae (
<figureCitation id="B1ED2A33FFBCFFEDF952FEBDFE30D1E5" box="[381,449,366,390]" captionStart="FIGURE 5" captionStartId="6.[151,249,1803,1826]" captionTargetBox="[152,1432,225,1802]" captionTargetId="figure@6.[151,1436,61,1828]" captionTargetPageId="6" captionText="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="9" pageNumber="48">Fig. 5</figureCitation>
, C).
</paragraph>
<paragraph id="296936B6FFBCFFEDF8EAFE40FEBDD1AD" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<emphasis id="1BA2EAA4FFBCFFEDF8EAFE40FEFED1C8" bold="true" box="[197,271,403,427]" pageId="9" pageNumber="48">Telson</emphasis>
: Lobes more than twice as long as width with 24 subdistal long setae, one distal spine and some long setae (
<figureCitation id="B1ED2A33FFBCFFEDF8F2FE65FEEBD1AD" box="[221,282,438,462]" captionStart="FIGURE 5" captionStartId="6.[151,249,1803,1826]" captionTargetBox="[152,1432,225,1802]" captionTargetId="figure@6.[151,1436,61,1828]" captionTargetPageId="6" captionText="FIGURE 5. Gammarus hegmatanensis sp. n., holotype, 3, 18 mm., from Gardaneh Asadabad spring, W of Hamadan. A (U 1) and B (U 2) on scale i; C (U 3) on scale ii; D (T) on scale iii; E (EP 1), F (EP 2) and G (EP 3) on scale iv." httpUri="https://zenodo.org/record/202036/files/figure.png" pageId="9" pageNumber="48">Fig 5</figureCitation>
, D).
</paragraph>
<paragraph id="296936B6FFBCFFEDF8EAFE09FD05D2E3" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<emphasis id="1BA2EAA4FFBCFFEDF8EAFE09FEEDD191" bold="true" box="[197,284,474,498]" pageId="9" pageNumber="48">Female</emphasis>
: Smaller than male (
<quantity id="EE2E9B53FFBCFFEDFA2AFE0AFD9ED192" box="[517,623,473,497]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="9" pageNumber="48" unit="mm" value="15.1">15.1 mm</quantity>
.), antenna 1 and 2 shorter, propodus of gnathopod 1 wider than propodus of gnathopod 2, propodus of gnathopod 1 and 2 smaller than in male, pereopod 3,4 similar to male, pereopod 5 similar to male but with some long setae on antero-inferior margin of basis. In female, compared to male, merus and carpus of pereopod 57 with more and longer setae, uropod 3 and telson are smaller than male and less setose. Oostegite 1 (of gnathopod 2) is illustrated (
<figureCitation id="B1ED2A33FFBCFFEDFA55FDBBFD16D2E3" box="[634,743,616,640]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="7.[151,249,1875,1898]" captionStartId-1="8.[151,249,1825,1848]" captionTargetBox-0="[309,1242,459,1739]" captionTargetBox-1="[152,1428,204,1824]" captionTargetId-0="figure@7.[151,1436,391,1873]" captionTargetId-1="figure@8.[151,1436,121,1900]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 6. Gammarus hegmatanensis sp. n., Allotype, Ƥ, 15.1 mm., from Gardaneh Asadabad spring, W of Hamadan. A (A 1) and B (A 2) on scale i; C (P 3) and D (P 4) on scale ii; E (dactyl of P 3) and F (dactyl of P 4) on scale iii." captionText-1="FIGURE 7. Gammarus hegmatanensis sp. n., Allotype, Ƥ, 15.1 mm., from Gardaneh Asadabad spring, W of Hamadan. A (P 5), B (P 6), C (P 7) and D (O) on scale i; E (U 3) on scale ii; F (dactyl of P 5), G (dactyl of P 6) and H (dactyl of 7) on scale iii." httpUri-0="https://zenodo.org/record/202037/files/figure.png" httpUri-1="https://zenodo.org/record/202038/files/figure.png" pageId="9" pageNumber="48">Figs. 67</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61CC653DFFBCFFEDF8EAFD5FFF3BD4FA" pageId="9" pageNumber="48" type="discussion">
<paragraph id="296936B6FFBCFFEDF8EAFD5FFDD9D2AB" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<emphasis id="1BA2EAA4FFBCFFEDF8EAFD5FFEBBD2C6" bold="true" box="[197,330,652,677]" pageId="9" pageNumber="48">Discussion.</emphasis>
The similarities and differences of
<taxonomicName id="EED64D35FFBCFFEDFACBFD5EFBFED2C6" box="[740,1039,653,677]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="hegmatanensis">
<emphasis id="1BA2EAA4FFBCFFEDFACBFD5EFBFED2C6" box="[740,1039,653,677]" italics="true" pageId="9" pageNumber="48">Gammarus hegmatanensis</emphasis>
</taxonomicName>
with some of the morphologically closest species are discussed below.
</paragraph>
<paragraph id="296936B6FFBCFFEDF8E8FD07FF3BD4FA" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
The most remarkable feature of
<taxonomicName id="EED64D35FFBCFFEDFA02FD06FCA6D28F" box="[557,855,724,748]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="hegmatanensis">
<emphasis id="1BA2EAA4FFBCFFEDFA02FD06FCA6D28F" box="[557,855,724,748]" italics="true" pageId="9" pageNumber="48">Gammarus hegmatanensis</emphasis>
</taxonomicName>
is the shape of epimeral plate 3, with a very sharply pointed and an extended posterior corner and distinct lobate anterior corner. The very sharply pointed posterior corner resembles
<taxonomicName id="EED64D35FFBCFFEDF914FCCFFD89D350" authority="Chevreux, 1895" authorityName="Chevreux" authorityYear="1895" box="[315,632,795,820]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="1BA2EAA4FFBCFFEDF914FCCFFE49D357" box="[315,440,796,820]" italics="true" pageId="9" pageNumber="48">G. syriacus</emphasis>
Chevreux, 1895
</taxonomicName>
, which has been reported by
<bibRefCitation id="4D474B47FFBCFFEDFBE8FCC8FB62D350" author="Stock" box="[967,1171,795,820]" pageId="9" pageNumber="48" refString="Stock, J. H., Mirzajani, A. R., Vonk, R., Naderi, S. &amp; Kiabi, B. H. (1998) Limnic and brackish water Amphipoda (Crustacea) from Iran. Beaufortia, 48 (9), 173 - 234." type="journal article" year="1998">
Stock
<emphasis id="1BA2EAA4FFBCFFEDFC20FCCEFBB0D357" box="[1039,1089,796,820]" italics="true" pageId="9" pageNumber="48">et al</emphasis>
., 1998
</bibRefCitation>
, from Kohgiluye-Buier Ahmad Province (South-east of the location, but not bounded). However,
<taxonomicName id="EED64D35FFBCFFEDFBFBFC93FB0CD334" box="[980,1277,831,855]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="hegmatanensis">
<emphasis id="1BA2EAA4FFBCFFEDFBFBFC93FB0CD334" box="[980,1277,831,855]" italics="true" pageId="9" pageNumber="48">Gammarus hegmatanensis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="EED64D35FFBCFFEDF91AFCB7FE43D318" box="[309,434,868,891]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="1BA2EAA4FFBCFFEDF91AFCB7FE43D318" box="[309,434,868,891]" italics="true" pageId="9" pageNumber="48">G. syriacus</emphasis>
</taxonomicName>
by lateral head lobes that are more rounded, anterior lower part of head more extended, the second and the third peduncular articles of antenna 1 not equal, flagellum with 26 articles; the fourth and fifth peduncular articles of antenna 2 not equal, flagellum has only eight articles, highly setose with long setae; posterior margin of pereopod 4 has much shorter setae and the number of setae and groups is lower; anterior margin of pereopods 57 is spinous (only with few short setae) versus long and numerus setae in
<emphasis id="1BA2EAA4FFBCFFEDFC01FC20FB42D469" box="[1070,1203,1011,1034]" italics="true" pageId="9" pageNumber="48">
<taxonomicName id="EED64D35FFBCFFEDFC01FC20FB5DD469" box="[1070,1196,1011,1034]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="syriacus">G. syriacus</taxonomicName>
;
</emphasis>
epimeral plate 1 and 2 without extended posterior corner and epimeral plate 2 and 3 with very lobate anterior corner. Moreover, pereopod
<quantity id="EE2E9B53FFBCFFEDF8E5FBE9FF09D431" box="[202,248,1082,1106]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="48" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="EED64D35FFBCFFEDF92EFBE8FE71D431" box="[257,384,1083,1106]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="syriacus">
<emphasis id="1BA2EAA4FFBCFFEDF92EFBE8FE71D431" box="[257,384,1083,1106]" italics="true" pageId="9" pageNumber="48">G. syriacus</emphasis>
</taxonomicName>
has a group of spines and setae at antero-inferior corner of basis; pereopod 57 basis with groups of marginal setae anteriorly, and pereopod 7 has a group of several setae at sub-marginal of antero-proximal part.
</paragraph>
</subSubSection>
<subSubSection id="61CC653DFFBCFFEEF8E8FB75FDA6D107" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="reference_group">
<paragraph id="296936B6FFBCFFEDF8E8FB75FD4BD5D4" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<taxonomicName id="EED64D35FFBCFFEDF8E8FB75FE01D4DE" ID-CoL="3F8CV" box="[199,496,1189,1213]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="hegmatanensis">
<emphasis id="1BA2EAA4FFBCFFEDF8E8FB75FE01D4DE" box="[199,496,1189,1213]" italics="true" pageId="9" pageNumber="48">Gammarus hegmatanensis</emphasis>
</taxonomicName>
is similar in some characters such as antenna 1 and 2 and gnathopod 1 to
<taxonomicName id="EED64D35FFBCFFEDFD00FB75FE7FD482" ID-CoL="6K98T" authority="Stock et al., 1998" authorityName="Stock et al." authorityYear="1998" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="parthicus">
<emphasis id="1BA2EAA4FFBCFFEDFD00FB75FF4CD482" italics="true" pageId="9" pageNumber="48">G. parthicus</emphasis>
<bibRefCitation id="4D474B47FFBCFFEDF8EAFB1AFE7FD482" author="Stock" box="[197,398,1225,1249]" pageId="9" pageNumber="48" refString="Stock, J. H., Mirzajani, A. R., Vonk, R., Naderi, S. &amp; Kiabi, B. H. (1998) Limnic and brackish water Amphipoda (Crustacea) from Iran. Beaufortia, 48 (9), 173 - 234." type="journal article" year="1998">
Stock
<emphasis id="1BA2EAA4FFBCFFEDF923FB19FEB1D482" box="[268,320,1225,1249]" italics="true" pageId="9" pageNumber="48">et al</emphasis>
., 1998
</bibRefCitation>
</taxonomicName>
reported and described from Markazi Province (the Eastern neighbouring province). However,
<taxonomicName id="EED64D35FFBCFFEDF8FBFB3EFEAFD566" ID-CoL="6K98T" box="[212,350,1261,1285]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="parthicus">
<emphasis id="1BA2EAA4FFBCFFEDF8FBFB3EFEAFD566" box="[212,350,1261,1285]" italics="true" pageId="9" pageNumber="48">G. parthicus</emphasis>
</taxonomicName>
differs from new species markedly with truncate lateral head lobes (versus rounded lateral head lobes in
<taxonomicName id="EED64D35FFBCFFEDF8DBFAC2FDEFD54B" ID-CoL="3F8CV" box="[244,542,1296,1320]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="hegmatanensis">
<emphasis id="1BA2EAA4FFBCFFEDF8DBFAC2FDEFD54B" box="[244,542,1296,1320]" italics="true" pageId="9" pageNumber="48">Gammarus hegmatanensis</emphasis>
</taxonomicName>
); different shapes of gnathopod 2 and epimeral plate 2; anterior corner of EP3 is not lobate and posteriorly more weakly pointed than the one in new species; basis of pereopod 57 anteriorly with some groups of marginal setae; existence of one group of setae and spines at postero-distal corner of basis of pereopod 6 and 7; uropod 3 outer margin of exopodite has plumose setae and telson lobes are very elongate and the length of lobes is longer 2.5 times than its width.
</paragraph>
<paragraph id="296936B6FFBCFFEDF8E8FA17FBC5D69A" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<taxonomicName id="EED64D35FFBCFFEDF8E8FA17FD13D5B8" ID-CoL="3F8N7" authority="Zamanpoore" authorityName="Zamanpoore" authorityYear="2009" box="[199,738,1475,1499]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="1BA2EAA4FFBCFFEDF8E8FA17FE3DD5B8" box="[199,460,1476,1499]" italics="true" pageId="9" pageNumber="48">Gammarus zagrosensis</emphasis>
<bibRefCitation id="4D474B47FFBCFFEDF9FCFA17FD13D5B8" author="Zamanpoore" box="[467,738,1475,1499]" pageId="9" pageNumber="48" refString="Zamanpoore, M., Poeckl, M., Grabowski, M. &amp; Schiemer, F. (2009) Two new sympatric species of freshwater Gammarus (Crustacea: Amphipoda) from Southern Zagros Region, Iran. Zootaxa, 2136, 21 - 39." type="journal article" year="2009">
Zamanpoore
<emphasis id="1BA2EAA4FFBCFFEDFA48FA17FD68D5B8" box="[615,665,1475,1499]" italics="true" pageId="9" pageNumber="48">et al</emphasis>
, 2009
</bibRefCitation>
</taxonomicName>
, described firstly in the Southern Zagros Mountains, has similar head lobe and antenna 1 as the new species; in contrast it has longer antenna 2 with 1117 flagellar articles armed with calceoli, short setae not forming a flag-like brush. Carpus and propodus of pereopod 3 bear some groups of long setae (23 times longer than the diameter of the segments). Merus and carpus of pereopod 4 have some groups of long setae. Anterior basis of pereopod 5 and 6 has one to two groups of 3 marginal setae and one seta at sub-margin of antero-proximal part of basis. Pereopod 6 bears 13 setae at the inner surface of the posterodistal corner of basis. Pereopod 7 is set with few long setae at postero-inferior corner basis. EP3 moderately pointed in posterior corner with 47 long setae on distal sub-marginal area of the second and the third epimeral plate. Dorsal surface of metasomites posteriorly is covered with very short setules.
</paragraph>
<paragraph id="296936B6FFBCFFEDF8E8F8D5FAB5D797" blockId="9.[151,1437,151,2036]" pageId="9" pageNumber="48">
<taxonomicName id="EED64D35FFBCFFEDF8E8F8D5FD73D77E" ID-CoL="3F8KG" authority="Zamanpoore" authorityName="Zamanpoore" authorityYear="2009" box="[199,642,1797,1821]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="sepidannus">
<emphasis id="1BA2EAA4FFBCFFEDF8E8F8D5FE97D77E" box="[199,358,1797,1821]" italics="true" pageId="9" pageNumber="48">G. sepidannus</emphasis>
<bibRefCitation id="4D474B47FFBCFFEDF940F8D5FD73D77E" author="Zamanpoore" box="[367,642,1797,1821]" pageId="9" pageNumber="48" refString="Zamanpoore, M., Poeckl, M., Grabowski, M. &amp; Schiemer, F. (2009) Two new sympatric species of freshwater Gammarus (Crustacea: Amphipoda) from Southern Zagros Region, Iran. Zootaxa, 2136, 21 - 39." type="journal article" year="2009">
Zamanpoore
<emphasis id="1BA2EAA4FFBCFFEDFA29F8D5FDB0D77E" box="[518,577,1797,1821]" italics="true" pageId="9" pageNumber="48">et al,</emphasis>
2009
</bibRefCitation>
</taxonomicName>
, has been described as a sympatric species with
<taxonomicName id="EED64D35FFBCFFEDFC96F8D5FAADD77E" ID-CoL="3F8N7" box="[1209,1372,1798,1821]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="9" pageNumber="48" phylum="Arthropoda" rank="species" species="zagrosensis">
<emphasis id="1BA2EAA4FFBCFFEDFC96F8D5FAADD77E" box="[1209,1372,1798,1821]" italics="true" pageId="9" pageNumber="48">G. zagrosensis</emphasis>
</taxonomicName>
from southern Zagros Mountains. It differs from the new species in the following ways: Antenna 2 flagellum with 1114 articles, gland cone is slightly longer than the third peduncle segment and peduncle segments 4 and 5 enjoy the same length. All segments of pereopod 3 and 4 posteriorly with dense groups of setae about 12 times the diameter of the segments. Pereopod 57 anterior margin of basis with a row of 48 setae, 34 spines and 13 long setae at antero-proximal inner surface. Endopodite of Uropod 3 half the length of the exopodite and plumose setae only on outer margin of exopodite and endopodite. Postero-inferior corner of EP3 is weakly to moderately pointed.
</paragraph>
<paragraph id="296936B6FFBFFFEEF8E8FF44FB4AD094" blockId="10.[151,1436,151,356]" pageId="10" pageNumber="49">
Finally,
<taxonomicName id="EED64D35FFBFFFEEF90AFF4BFE5FD0D3" ID-CoL="6K99D" box="[293,430,152,176]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="komareki">
<emphasis id="1BA2EAA4FFBFFFEEF90AFF4BFE5FD0D3" box="[293,430,152,176]" italics="true" pageId="10" pageNumber="49">G. komareki</emphasis>
</taxonomicName>
shares some features with
<emphasis id="1BA2EAA4FFBFFFEEFAC3FF4BFC4BD0D3" box="[748,954,152,176]" italics="true" pageId="10" pageNumber="49">
<taxonomicName id="EED64D35FFBFFFEEFAC3FF4BFC47D0D3" ID-CoL="3F8CV" box="[748,950,152,176]" class="Malacostraca" family="Gammaridae" genus="Gammarus" kingdom="Animalia" order="Amphipoda" pageId="10" pageNumber="49" phylum="Arthropoda" rank="species" species="hegmatanensis">G. hegmatanensis</taxonomicName>
,
</emphasis>
like small eyes, long setae on antenna 2, more or less the same ratio of U3 endo- and exopodite, but it has some differences with this new species including short gland cone, shape of basis in pereopod 7, shape of EP 2 &amp; 3 and elevation of urosomites.
</paragraph>
<paragraph id="296936B6FFBFFFEEF8E8FED0FDA6D107" blockId="10.[151,1436,151,356]" pageId="10" pageNumber="49">
<emphasis id="1BA2EAA4FFBFFFEEF8E8FED0FEBCD17F" bold="true" box="[199,333,259,284]" pageId="10" pageNumber="49">Variability.</emphasis>
The examined
<typeStatus id="F66D8814FFBFFFEEF9D5FED6FD94D17E" box="[506,613,261,285]" pageId="10" pageNumber="49" type="paratype">paratypes</typeStatus>
show variations in some characters such as number of terminal spines in uropod 1 &amp; 2, number of setae and spines on distal and subdistal position of telson, and number of setae and spines in palp of right maxilla and left maxilla.
</paragraph>
</subSubSection>
</treatment>
</document>