168 lines
15 KiB
XML
168 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1174.107314" ID-Pensoft-Pub="1313-2970-1174-35" ID-Pensoft-UUID="635A059AEF505E17B338F6484B5F4DBF" ID-ZooBank="91E908240C45471A86BBA24ADC35E743" ModsDocID="1313-2970-1174-35" checkinTime="1691559557484" checkinUser="pensoft" docAuthor="Harris, Steven C. & Armitage, Brian J." docDate="2023" docId="9D01A6249EBE50549CBB10C9DB9B6221" docLanguage="en" docName="ZooKeys 1174: 35-74" docOrigin="ZooKeys 1174" docPubDate="2023-08-08" docSource="http://dx.doi.org/10.3897/zookeys.1174.107314" docTitle="Metrichia escobilla Harris & Armitage 2023, sp. nov." docType="treatment" docUuid="368B4424-5833-4E39-937F-AAF99D1E183D" docUuidSource="ZooBank" docVersion="1" id="635A059AEF505E17B338F6484B5F4DBF" lastPageNumber="35" masterDocId="635A059AEF505E17B338F6484B5F4DBF" masterDocTitle="The Trichoptera of Panama XXII. Sixteen new microcaddisfly species (Trichoptera, Hydroptilidae)" masterLastPageNumber="74" masterPageNumber="35" pageNumber="35" updateTime="1691559557484" updateUser="pensoft">
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<mods:mods id="014B0D229F0F69BECEFB6563B1CE457B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E2B40E5CF9E48BAEF8AB5218911445A2">
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<mods:title id="51D768E66A5F7CE2CE49305DFE69D201">The Trichoptera of Panama XXII. Sixteen new microcaddisfly species (Trichoptera, Hydroptilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A4D056CF16967C5BDB4E2A08C95928E9" type="personal">
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<mods:role id="456C54171211F7AC06DE2BBF00BEEE5E">
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<mods:roleTerm id="023F26B81B903422916063BBDABC298E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A6390056DF86521E80C0FC98B75F2CFE">Harris, Steven C.</mods:namePart>
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<mods:nameIdentifier id="6BE32AE4445CB2C2170E1791BA83A09F" type="ORCID">https://orcid.org/0000-0002-6432-7462</mods:nameIdentifier>
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<mods:affiliation id="B871D7B80EE3F834FA5053C39706D94A">Department of Biology and Environmental Sciences, Western Pennsylvania University, Clarion, PA 16214, USA & Museo de Peces de Agua Dulce e Invertebrados, Universidad Autonoma de Chiriqui, David, Panama</mods:affiliation>
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</mods:name>
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<mods:name id="0A4E56DBDE7ED115E7CF825BAA8EE544" type="personal">
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<mods:role id="2DD6328519EF7FF5282681DE764139C9">
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<mods:roleTerm id="9A4E691DE6D597DC7CDC9CD8D63DB9C8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="50764627AF3CB73BEB6232A516C6E248">Armitage, Brian J.</mods:namePart>
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<mods:nameIdentifier id="B31E28FA09D422CD9C9FE3292B5F4521" type="ORCID">https://orcid.org/0000-0003-3182-1533</mods:nameIdentifier>
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<mods:affiliation id="99A3DE2E12AE024A037BF0A9D94D1F06">Museo de Peces de Agua Dulce e Invertebrados, Universidad Autonoma de Chiriqui, David, Panama</mods:affiliation>
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<mods:nameIdentifier id="4B550F0E6392A8EBA0130D189C3FC200" type="email">tobikera89@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="2A0FB088E451C4240073527A6909C1E4">text</mods:typeOfResource>
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<mods:titleInfo id="D8325DD08A83C9A77A89F2236BABDA98">
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<mods:title id="6D9E4C4840E14A0454C2457AC49D7698">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="C9CD6EC8BC74F068D45D9018830322E3">
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<mods:date id="3FD460956CD2642E1C906925CC8F8EFF">2023</mods:date>
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<mods:detail id="CDB7DFF3E9F682560C8502842D2927D0" type="pubDate">
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<mods:number id="B27FE0DD2F269D67099E31BA44C8BF01">2023-08-08</mods:number>
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</mods:detail>
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<mods:detail id="EACB0C587DA8FF21639DD1F6504778F0" type="volume">
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<mods:number id="B37816133AD7A06AA0D1833340506CC2">1174</mods:number>
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</mods:detail>
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<mods:extent id="D9160865366E4C69DBC0A47A7EF066A1" unit="page">
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<mods:start id="EA45E4DEC7326BCE5EE649C759888846">35</mods:start>
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<mods:end id="5A36BDF6154185A3D680D81042071853">74</mods:end>
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<mods:location id="3FEBBC96BAB01B33C388E472BFC3AD18">
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<mods:url id="A5E95A829C4FE4D5564CED88F54503D3">http://dx.doi.org/10.3897/zookeys.1174.107314</mods:url>
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</mods:location>
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<mods:classification id="4D8078E17DA29DBC7A4D886615EF417A">journal article</mods:classification>
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<mods:identifier id="5FD8E06B5C787D08A84BA4C703480E9F" type="DOI">http://dx.doi.org/10.3897/zookeys.1174.107314</mods:identifier>
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<mods:identifier id="C66926F611B232C206AD9E5E8F51FC12" type="Pensoft-Pub">1313-2970-1174-35</mods:identifier>
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<mods:identifier id="CB8DEC710E12BF817BABEB678E3DE233" type="ZooBank">91E908240C45471A86BBA24ADC35E743</mods:identifier>
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<mods:identifier id="334DDF824B37C5D597E85AD4F75B7927" type="Pensoft-UUID">635A059AEF505E17B338F6484B5F4DBF</mods:identifier>
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</mods:mods>
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<treatment id="9D01A6249EBE50549CBB10C9DB9B6221" LSID="urn:lsid:zoobank.org:act:368B4424-5833-4E39-937F-AAF99D1E183D" httpUri="http://treatment.plazi.org/id/9D01A6249EBE50549CBB10C9DB9B6221" lastPageNumber="35" pageId="0" pageNumber="35">
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<subSubSection id="7BEDB745B9B7F78555395B1545B23CBC" pageId="0" pageNumber="35" type="nomenclature">
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<paragraph id="EF61D5D6FAA2EB2B76A3D280D6837DB6" pageId="0" pageNumber="35">
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<taxonomicName id="CFE30C43EE325AC58015067C23314DD6" LSID="https://zoobank.org/368B4424-5833-4E39-937F-AAF99D1E183D" authority="Harris & Armitage, 2023" authorityName="Harris & Armitage" authorityYear="2023" class="Insecta" family="Hydroptilidae" genus="Metrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrichia escobilla" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="escobilla" status="sp. nov.">Metrichia escobilla</taxonomicName>
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<taxonomicNameLabel id="A787DDCD6008782A810CF76AA836B84B" pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="E24B22E2BA145121639652C3EBF40EDC" pageId="0" pageNumber="35" type="description">
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<paragraph id="62517F087FA867E643C45567CFCA7637" pageId="0" pageNumber="35">
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<figureCitation id="99945076F45ECE1A6D859585381AF022" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Metrichia escobilla sp. nov., male holotype, genitalia A left lateral B dorsal C ventral D phallus apex, left lateral E phallus, dorsal." figureDoi="10.3897/zookeys.1174.107314.figure12" httpUri="https://binary.pensoft.net/fig/890137" pageId="0" pageNumber="35">Fig. 12</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="58729242E0CC5676500B67F8E1D146EE" pageId="0" pageNumber="35" type="type locality">
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<paragraph id="5A5DA18650A1F70FF99C548B0A50D5C1" pageId="0" pageNumber="35">Type locality.</paragraph>
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<paragraph id="55FF1111BD9B2A3B9026C44F6235F657" pageId="0" pageNumber="35">
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<emphasis id="86A0FF0B8752AD629326175AE68DB798" bold="true" pageId="0" pageNumber="35">Panama: Veraguas Province</emphasis>
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: Cuenca 097; Santa Fe District; Santa Fe NP;
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<normalizedToken id="302E32D3396A5271ABD1113D900A3401" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="3BD7D84E413ACC2E23019676A2CDF8B9" originalValue="Calovébora">Calovebora</normalizedToken>
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; PSPSCB-PNSF-C097-2017-005;
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<geoCoordinate id="41ABF0C939632E28C7888D89E6FBEC36" degrees="8.54318" direction="north" orientation="latitude" precision="1" value="8.54318">8.54318°N</geoCoordinate>
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,
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<geoCoordinate id="500C3B04B31A90AB52FD2BB74880742A" degrees="81.16398" direction="west" orientation="longitude" precision="1" value="-81.16398">81.16398°W</geoCoordinate>
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; 536 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="D728C3A4C1537AD395599EBCA126A85B" pageId="0" pageNumber="35" type="type specimen">
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<paragraph id="916194DFDEBFC6F316AAEC5A568C8005" pageId="0" pageNumber="35">Type specimen.</paragraph>
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<paragraph id="6F0D030BF8A4B76835FD7B77F3BA29BA" pageId="0" pageNumber="35">
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<emphasis id="53BD7208AC3363C5B2424803F065314A" bold="true" pageId="0" pageNumber="35">
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<emphasis id="81688B0219B12D454D41D6FB11AB486B" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
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: male, Panama: Veraguas Province
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</emphasis>
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: Cuenca 097; Santa Fe District; Santa Fe NP;
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<normalizedToken id="6450BDEA62D1303E4085B15796ADE7DD" originalValue="Río">Rio</normalizedToken>
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<normalizedToken id="5227A0A5113F3606B11FEA368A5ADF0F" originalValue="Calovébora">Calovebora</normalizedToken>
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; PSPSCB-PNSF-C097-2017-005;
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<geoCoordinate id="E2F26DEA393B1637C5432FDC6513D528" degrees="8.54318" direction="north" orientation="latitude" precision="1" value="8.54318">8.54318°N</geoCoordinate>
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,
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<geoCoordinate id="4DB0B232B3C9671EF49284DAB42EC649" degrees="81.16398" direction="west" orientation="longitude" precision="1" value="-81.16398">81.16398°W</geoCoordinate>
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; 536 m a.s.l.; UV light trap; T.
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<normalizedToken id="EC1056520407865888104623B8D2CFAB" originalValue="Ríos">Rios</normalizedToken>
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, E.
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<normalizedToken id="FA9797F96EB87CAD8C98B5C41608321F" originalValue="Álvarez">Alvarez</normalizedToken>
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, C. Nieto, leg.; 21.iv.2017; MIUP-010-T-2023 (in alcohol).
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</paragraph>
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</subSubSection>
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<subSubSection id="138439A45835AA8B0D5003B195730271" pageId="0" pageNumber="35" type="diagnosis">
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<paragraph id="6E96A89573D5ECB4CBCB045374392602" pageId="0" pageNumber="35">Diagnosis.</paragraph>
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<paragraph id="E07ACA18C4B30A3FFE9205F3A3C610A6" pageId="0" pageNumber="35">
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This species shares a number of character states with
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<taxonomicName id="A34578AE6F847D2CD5A786A5618E6C91" lsidName="M. angulosa" pageId="0" pageNumber="35" rank="species" species="angulosa">
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<emphasis id="9DB556CC059B68EE10BA3B9183A6A63B" italics="true" pageId="0" pageNumber="35">M. angulosa</emphasis>
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</taxonomicName>
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Bueno-Soria & Holzenthal, which occurs in Costa Rica and Panama. Both species have a similarly shaped inferior appendage, with dentate posterior margin, and both have apical spines on the phallus. However, while
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<taxonomicName id="E750AB9219F10A62C49C38BFAFC287AB" lsidName="M. angulosa" pageId="0" pageNumber="35" rank="species" species="angulosa">
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<emphasis id="EE24A031DA13F973AD49E22C35318642" italics="true" pageId="0" pageNumber="35">M. angulosa</emphasis>
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</taxonomicName>
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has prominent reniform pouches on the dorsum of abdominal segment V, the abdomen of
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<taxonomicName id="4A8E092841F1FFA9A9A21E38F07B3EDE" lsidName="M. escobilla" pageId="0" pageNumber="35" rank="species" species="escobilla">
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<emphasis id="F211E3FFD49F26045BE8ACA9FC920B9F" italics="true" pageId="0" pageNumber="35">M. escobilla</emphasis>
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</taxonomicName>
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sp. nov. lacks abdominal modifications. As well, the new species has both phallic spines pointed, rather than one truncate as in
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<taxonomicName id="3EFDE298034E8B973BCEA61978BBF0AD" lsidName="M. angulosa" pageId="0" pageNumber="35" rank="species" species="angulosa">
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<emphasis id="89135A381EE5DEB066F3D4C974947280" italics="true" pageId="0" pageNumber="35">M. angulosa</emphasis>
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</taxonomicName>
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, the dentation on the posterior margin of the inferior appendage is more pronounced, and the brush-like apex of the dorsolateral hook is unique to
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<taxonomicName id="7FD98B4072594D8C14A5EE3E42EF67CB" lsidName="M. escobilla" pageId="0" pageNumber="35" rank="species" species="escobilla">
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<emphasis id="63BB49C3EED3935051493B43033FB67C" italics="true" pageId="0" pageNumber="35">M. escobilla</emphasis>
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</taxonomicName>
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sp. nov..
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</paragraph>
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<caption id="B0C7D996882A0665D7E2305924C9A4F2" doi="10.3897/zookeys.1174.107314.figure12" httpUri="https://binary.pensoft.net/fig/890137" pageId="0" pageNumber="35" start="Figure 12" startId="F12">
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<paragraph id="CF82170B4D9387F136D77E8DD6707C20" pageId="0" pageNumber="35">
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<emphasis id="1B90463F42AB3B7914069F58B0BE512A" bold="true" pageId="0" pageNumber="35">Figure 12.</emphasis>
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<taxonomicName id="BFB14BA2E38A4A4F3A1F9A9B357C2B84" authorityName="Harris & Armitage" authorityYear="2023" class="Insecta" family="Hydroptilidae" genus="Metrichia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrichia escobilla" order="Trichoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="escobilla">
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<emphasis id="F72A39895E0E40EBD81E192839489FE7" italics="true" pageId="0" pageNumber="35">Metrichia escobilla</emphasis>
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</taxonomicName>
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sp. nov., male holotype, genitalia
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<emphasis id="0BBDFEF41AE51BA6A5E7CBDEE119DE70" bold="true" pageId="0" pageNumber="35">A</emphasis>
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left lateral
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<emphasis id="934A16E64950C83D7EC6919EA7BEBBDF" bold="true" pageId="0" pageNumber="35">B</emphasis>
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dorsal
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<emphasis id="416E84D190B872D7094BC16044C1F440" bold="true" pageId="0" pageNumber="35">C</emphasis>
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ventral
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<emphasis id="406AFCEDFDBBCB8ED756E742B001CA22" bold="true" pageId="0" pageNumber="35">D</emphasis>
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phallus apex, left lateral
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<emphasis id="D576A19105BA928614C90D5540CBA0FD" bold="true" pageId="0" pageNumber="35">E</emphasis>
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phallus, dorsal.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="17F1C79D5DA78722311C9DCD703BB014" pageId="0" pageNumber="35" type="description">
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<paragraph id="20F30400418913DBCE1B21E6C14F9C55" pageId="0" pageNumber="35">Description.</paragraph>
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<paragraph id="4A799B25D0F0BF779379628C3C789AFE" pageId="0" pageNumber="35">
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<emphasis id="1081E583B1D01088F6454E060DCF671F" bold="true" pageId="0" pageNumber="35">Male.</emphasis>
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Total length 1.6 mm, 18 antennal segments, wings and body brown in alcohol, abdominal terga without modifications.
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<emphasis id="49DABE9F207B7C611AA36C1FDE38C446" bold="true" pageId="0" pageNumber="35">
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<emphasis id="B9769E32F0BF2F00B5319896711D80F0" italics="true" pageId="0" pageNumber="35">Genitalia</emphasis>
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.
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</emphasis>
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Abdominal segment VII annular without short ventromesal process. Segment VIII triangular, tapering ventrally; in ventral view deeply incised mesally; in dorsal view quadrate. Segment IX triangular, truncate posteriorly, tapering anteriorly into segment VII. Preanal appendage (cercus) oval in lateral and dorsal views. Dorsolateral hook in lateral view wide basally, converging near midlength, apex serrate ventrally; in dorsal view wide basally, tapering to serrate margins at apices. Segment X lobate; in dorsal view triangular, membranous apically. Inferior appendage in lateral view narrow at base then widening dorsally, rounded apically; in dorsal and ventral view quadrate, diverging basally, posterior margins dentate. Phallus very long, extending into segment V; in dorsal view wide basally, narrowing posteriorly, pair of spines apically, subapical spine longer than apical, lateral process originating at midlength; in lateral view, subapical spine slightly longer than apical, both nearly straight, phallus apex widening into plate-like structure.
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</paragraph>
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</subSubSection>
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<subSubSection id="6C7F8D40C952F59738FC31D3E997EB86" pageId="0" pageNumber="35" type="distribution">
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<paragraph id="F48FC360101359A4AE06E827DB0F74FD" pageId="0" pageNumber="35">Distribution.</paragraph>
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<paragraph id="50B05A7192D706D043A0E014F2D1CC6D" pageId="0" pageNumber="35">Panama.</paragraph>
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</subSubSection>
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<subSubSection id="43193186F5B2EA011131E86599061733" pageId="0" pageNumber="35" type="etymology">
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<paragraph id="E4CCC9C8550A14577F271B1061110961" pageId="0" pageNumber="35">Etymology.</paragraph>
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<paragraph id="8381A28D56D8510481215E2A824C8AA4" pageId="0" pageNumber="35">
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The species name
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<emphasis id="397F5BEA1BE2892B219EE071DB9843E4" italics="true" pageId="0" pageNumber="35">escobillla</emphasis>
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(brush) derives from Spanish, referring to the brush-like apex of the dorsolateral hook in lateral view. The name is a noun in the nominative singular standing in apposition.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |