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<document id="1DF5435610223D5A29B25F4CA55E5FB3" ID-CLB-Dataset="5008" ID-DOI="10.1649/0010-065X-74.4.699" ID-GBIF-Dataset="6f145036-397c-4332-8c04-de9e31643aa7" ID-ISSN="1938-4394" ID-Zenodo-Dep="4791137" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1621947350721" checkinUser="carolina" docAuthor="Steury, Brent W. &amp; Steiner, Warren E." docDate="2020" docId="902587FBFFF6921DFD646EEF8FAD166B" docLanguage="en" docName="TheColeopBull.74.4.699-709.pdf" docOrigin="The Coleopterists Bulletin 74 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-74.4.699" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Pseudotolida syphaxi Steury and Steiner 2020, new species" docType="treatment" docUuid="1177B262-2A90-4C5C-927F-BCF8179B8585" docUuidSource="ZooBank" docVersion="10" lastPageNumber="708" masterDocId="6C1CFF83FFF19214FFB56C2A8C501504" masterDocTitle="Descriptions of Four New Species of Tumbling Flower Beetles (Coleoptera: Mordellidae) from Eastern North America" masterLastPageNumber="709" masterPageNumber="699" pageNumber="706" updateTime="1698989847044" updateUser="ExternalLinkService">
<mods:mods id="848F082B38D1DF25CFF5921FFA9F1052" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="80CE555208B753928F31A3C056269E0A">
<mods:title id="6A00F9A650D02493C60FA385B54444E0">Descriptions of Four New Species of Tumbling Flower Beetles (Coleoptera: Mordellidae) from Eastern North America</mods:title>
</mods:titleInfo>
<mods:name id="32ADB8D34116FB6078F087FCF7ADD224" type="personal">
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</mods:role>
<mods:namePart id="B55BE07370F488DD6624879C9F9F6982">Steury, Brent W.</mods:namePart>
</mods:name>
<mods:name id="A6936DB2CE3C23A5399B7DAAFA5974D4" type="personal">
<mods:role id="697B1F844EA6E5CA25845B1659BA92A0">
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</mods:role>
<mods:namePart id="07C4B2B6F99E50D87E32746CD025F725">Steiner, Warren E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="387B5B388A8417C93422C62EA1BC39DB">text</mods:typeOfResource>
<mods:relatedItem id="A7E0D66FF1039982597AE1FAA00BBD74" type="host">
<mods:titleInfo id="4C9CCBA7ADB6341DEA2FBB3889C31AA1">
<mods:title id="21C78C2843C99935FE5D64EC27C15773">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="537690BFFE3CA886DB5CDDF73489AE51">
<mods:date id="A1DC40EEA9E054D4638A496CCE5EEFA2">2020</mods:date>
<mods:detail id="8FFF7514C9149B08805D9647F0B2B56A" type="pubDate">
<mods:number id="CA4C4AFF8FBDC4002DC7E4308ED7943C">2020-12-22</mods:number>
</mods:detail>
<mods:detail id="B0A392BBA6EE217E00EF45316F3FB147" type="volume">
<mods:number id="168E4F91513437215FD2D672D3BAA95A">74</mods:number>
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<mods:detail id="1BC487800998C86C22528C71506C8126" type="issue">
<mods:number id="762AE1B520A9953600B67947F07B6B1E">4</mods:number>
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<mods:start id="53A5EB6A578DEA653D8E115DC55488D8">699</mods:start>
<mods:end id="86E7FDD7C34D92A3E2A16BCA869F9E0C">709</mods:end>
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<mods:url id="A202B06A09A25F1B0D3673105355E963">http://dx.doi.org/10.1649/0010-065x-74.4.699</mods:url>
</mods:location>
<mods:classification id="A06A85521120F94290BDC72E814DAF97">journal article</mods:classification>
<mods:identifier id="77C5910206F622DD1091D853055AC803" type="CLB-Dataset">5008</mods:identifier>
<mods:identifier id="591B6F78253C26E8D4012360C3AB9DF8" type="DOI">10.1649/0010-065X-74.4.699</mods:identifier>
<mods:identifier id="54C5B38F2D87F9F39EF8E0E3C81032F4" type="GBIF-Dataset">6f145036-397c-4332-8c04-de9e31643aa7</mods:identifier>
<mods:identifier id="F5CA6E2895EF961980CD63F73669A87F" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="936D166BAB71CC6E920FD0A1D8051115" type="Zenodo-Dep">4791137</mods:identifier>
<mods:identifier id="79490E93CC2708283C245B5FF3EE3790" type="ZooBank">3FF23BA0-B042-4F1B-AE59-6002DCBB12C0</mods:identifier>
</mods:mods>
<treatment id="902587FBFFF6921DFD646EEF8FAD166B" ID-DOI="http://doi.org/10.5281/zenodo.5191591" ID-GBIF-Taxon="184096793" ID-Zenodo-Dep="5191591" LSID="urn:lsid:zoobank.org:act:1177B262-2A90-4C5C-927F-BCF8179B8585" httpUri="http://treatment.plazi.org/id/902587FBFFF6921DFD646EEF8FAD166B" lastPageId="9" lastPageNumber="708" pageId="7" pageNumber="706">
<subSubSection id="50966566FFF69213FD646EEF8F8B17F1" pageId="7" pageNumber="706" type="nomenclature">
<paragraph id="183336EDFFF69213FD646EEF8F8B17F1" blockId="7.[706,1150,709,838]" pageId="7" pageNumber="706">
<heading id="437B8181FFF69213FD646EEF8F8B17F1" centered="true" fontSize="9" level="2" pageId="7" pageNumber="706" reason="2">
<emphasis id="2AF8EAFFFFF69213FD646EEF8F8B17F1" bold="true" pageId="7" pageNumber="706">
<taxonomicName id="DF8C4D6EFFF69213FD646EEF883C17DF" authority="Steury and Steiner" authorityName="Steury and Steiner" authorityYear="2020" box="[721,1132,709,731]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="706" phylum="Arthropoda" rank="species" species="syphaxi" status="sp. nov.">
<emphasis id="2AF8EAFFFFF69213FD646EEF8FF417DE" bold="true" box="[721,932,709,730]" italics="true" pageId="7" pageNumber="706">Pseudotolida syphaxi</emphasis>
Steury and Steiner
</taxonomicName>
,
<taxonomicNameLabel id="31CB5784FFF69213FCD36ECA8F8B17F1" box="[870,987,736,757]" pageId="7" pageNumber="706" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF69213FD596ED1882E162F" pageId="7" pageNumber="706" type="description">
<paragraph id="183336EDFFF69213FD596ED1882E162F" blockId="7.[706,1150,709,838]" pageId="7" pageNumber="706">
zoobank.org/
<uri id="6C1D3AEFFFF69213FCD86ED1882E162F" pageId="7" pageNumber="706">
urn:lsid:zoobank.org:act:
<uuid id="6C2A0C38FFF69213FD776F3C882E162F" box="[706,1150,790,811]" pageId="7" pageNumber="706">1177B262-2A90-4C5C-927F-BCF8179B8585</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF69213FCD56F1A8FB11641" box="[864,993,816,838]" pageId="7" pageNumber="706" type="description">
<paragraph id="183336EDFFF69213FCD56F1A8FB11641" blockId="7.[706,1150,709,838]" box="[864,993,816,838]" pageId="7" pageNumber="706">
(
<figureCitation id="80B72A68FFF69213FCD26F1A8F8A1641" box="[871,986,816,838]" captionStart="Fig" captionStartId="8.[197,228,1543,1562]" captionTargetBox="[179,1188,196,1525]" captionTargetId="figure-58@8.[175,1190,194,1527]" captionTargetPageId="8" captionText="Fig. 9. Pseudotolida syphaxi, new species, holotype, male. A) Dorsal, B) Lateral, C) Antenna, D) Terminal maxillary palpomeres. Virginia, Fairfax County, Turkey Run Park, gulch, Malaise trap, 19–30 June 2009, D. Smith and B. Steury. Length 3.5 mm to tip of elytra, 4.2 mm to tip of pygidium." figureDoi="http://doi.org/10.5281/zenodo.4791335" httpUri="https://zenodo.org/record/4791335/files/figure.png" pageId="7" pageNumber="706">Figs. 9A–D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF69213FD706F718FCE11CD" pageId="7" pageNumber="706" type="materials_examined">
<paragraph id="183336EDFFF69213FD706F718FCE11CD" blockId="7.[685,1173,859,1730]" pageId="7" pageNumber="706">
<emphasis id="2AF8EAFFFFF69213FD706F718F301674" bold="true" box="[709,864,859,880]" pageId="7" pageNumber="706">Type Material.</emphasis>
<materialsCitation id="A8E43CB0FFF69213FCD96F718FD016C4" ID-GBIF-Occurrence="3333864314" collectingDate="2009-06-19" collectingDateMax="2009-06-30" collectingDateMin="2009-06-19" collectionCode="USNM, GWMP" collectorName="D. Smith &amp; B. Steury." country="Turkey" county="Fairfax Co." location="Male" pageId="7" pageNumber="706" specimenCount="1" typeStatus="holotype">
<emphasis id="2AF8EAFFFFF69213FCD96F718F9E1674" bold="true" box="[876,974,859,880]" pageId="7" pageNumber="706">
<typeStatus id="C737884FFFF69213FCD96F718F9A1674" box="[876,970,859,880]" pageId="7" pageNumber="706" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="1D536036FFF69213FC6C6F71885A1674" LSID="urn:lsid:plazi:treatment:902587FBFFF6921DFD646EEF8FAD166B:1D536036FFF69213FC6C6F71885A1674" box="[985,1034,859,880]" country="Turkey" county="Fairfax Co." name="Male" pageId="7" pageNumber="706">Male</location>
(
<collectionCode id="7E9DAE28FFF69213FBA96F7188351674" box="[1052,1125,859,880]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="706" type="Museum">USNM</collectionCode>
), labeled “Virginia,
<collectingCounty id="F1524E61FFF69213FCFB6F5F8FEF168E" box="[846,959,885,906]" pageId="7" pageNumber="706">Fairfax Co.</collectingCounty>
,
<collectionCode id="7E9DAE28FFF69213FC7F6F5F8844168E" box="[970,1044,885,906]" pageId="7" pageNumber="706">GWMP</collectionCode>
,
<collectingCountry id="609B767DFFF69213FB956F5F8834168E" box="[1056,1124,885,906]" name="Turkey" pageId="7" pageNumber="706">Turkey</collectingCountry>
Run Park, gulch,
<collectingMethod id="C1CD4EFAFFF69213FC846FBA8FFE16A1" box="[817,942,912,933]" pageId="7" pageNumber="706">Malaise trap</collectingMethod>
,
<date id="6C32102DFFF69213FC086FBA883D16A1" box="[957,1133,911,933]" pageId="7" pageNumber="706" value="2009-06-19" valueMax="2009-06-30" valueMin="2009-06-19">
<collectingDate id="7C76E9C5FFF69213FC086FBA883D16A1" box="[957,1133,911,933]" pageId="7" pageNumber="706" value="2009-06-19" valueMax="2009-06-30" valueMin="2009-06-19">19–30 June 2009</collectingDate>
</date>
,
<collectorName id="B579533BFFF69213FBC96FBA8EB516BB" pageId="7" pageNumber="706">D. Smith</collectorName>
&amp;
<collectorName id="B579533BFFF69213FCA76F808F2816BB" box="[786,888,938,959]" pageId="7" pageNumber="706">B. Steury.</collectorName>
”
</materialsCitation>
<materialsCitation id="A8E43CB0FFF69213FC3B6F808FCA11CD" ID-GBIF-Occurrence="3333864302" collectingDate="2009-07-31" collectingDateMax="2018-08-17" collectingDateMin="2009-07-31" collectionCode="USNM, GWMP" collectorName="B. Steury" country="Turkey" county="Fairfax Co." location="Little Hunting Creek" pageId="7" pageNumber="706" specimenCount="4" typeStatus="paratype">
<emphasis id="2AF8EAFFFFF69213FC3B6F808FA316BB" bold="true" box="[910,1011,938,959]" pageId="7" pageNumber="706">
<typeStatus id="C737884FFFF69213FC3B6F808FA316BB" box="[910,1011,938,959]" pageId="7" pageNumber="706" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<emphasis id="2AF8EAFFFFF69213FBB26F80884316BB" box="[1031,1043,938,959]" italics="true" pageId="7" pageNumber="706">n</emphasis>
= 12). One
<typeStatus id="C737884FFFF69213FD186FEF8F5116DE" box="[685,769,965,986]" pageId="7" pageNumber="706" type="paratype">paratype</typeStatus>
, male (
<collectionCode id="7E9DAE28FFF69213FCE76FEF8FCB16DE" box="[850,923,965,986]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="706" type="Museum">USNM</collectionCode>
), “Virginia,
<collectingCounty id="F1524E61FFF69213FBAE6FEF88C016DE" box="[1051,1168,965,986]" pageId="7" pageNumber="706">Fairfax Co.</collectingCounty>
,
<collectionCode id="7E9DAE28FFF69213FD186FF58EA716F0" box="[685,759,991,1012]" pageId="7" pageNumber="706">GWMP</collectionCode>
,
<location id="1D536036FFF69213FCBD6FF58F8D16F0" LSID="urn:lsid:plazi:treatment:902587FBFFF6921DFD646EEF8FAD166B:1D536036FFF69213FCBD6FF58F8D16F0" box="[776,989,991,1012]" country="Turkey" county="Fairfax Co." name="Little Hunting Creek" pageId="7" pageNumber="706">Little Hunting Creek</location>
,
<collectingMethod id="C1CD4EFAFFF69213FC586FF5883B16F0" box="[1005,1131,991,1012]" pageId="7" pageNumber="706">Malaise trap</collectingMethod>
,
<date id="6C32102DFFF69213FBC96FF58FD0110B" pageId="7" pageNumber="706" value="2018-07-30" valueMax="2018-08-17" valueMin="2018-07-30">
<collectingDate id="7C76E9C5FFF69213FBC96FF58FD0110B" pageId="7" pageNumber="706" value="2018-07-30" valueMax="2018-08-17" valueMin="2018-07-30">30 July–17 August 2018</collectingDate>
</date>
,
<collectorName id="B579533BFFF69213FC386FD08FBE110B" box="[909,1006,1018,1039]" pageId="7" pageNumber="706">B. Steury.</collectorName>
” Additional
<typeStatus id="C737884FFFF69213FBDB6FD08EBC112E" pageId="7" pageNumber="706" type="paratype">paratypes</typeStatus>
, males (
<collectionCode id="7E9DAE28FFF69213FC88683F8FDC112E" box="[829,908,1045,1066]" pageId="7" pageNumber="706">GWMP</collectionCode>
), same data as USNM
<typeStatus id="C737884FFFF69213FBDB683F8EB01140" pageId="7" pageNumber="706" type="paratype">paratype</typeStatus>
(
<emphasis id="2AF8EAFFFFF69213FD5868058EA91140" box="[749,761,1071,1092]" italics="true" pageId="7" pageNumber="706">n</emphasis>
= 2); same data but “
<date id="6C32102DFFF69213FC08680588D91140" box="[957,1161,1071,1093]" pageId="7" pageNumber="706" value="2018-06-28" valueMax="2018-07-16" valueMin="2018-06-28">
<collectingDate id="7C76E9C5FFF69213FC08680588D91140" box="[957,1161,1071,1093]" pageId="7" pageNumber="706" value="2018-06-28" valueMax="2018-07-16" valueMin="2018-06-28">28 June–16 July 2018</collectingDate>
</date>
” (
<emphasis id="2AF8EAFFFFF69213FD0068608E91115B" box="[693,705,1098,1119]" italics="true" pageId="7" pageNumber="706">n</emphasis>
= 3); “
<date id="6C32102DFFF69213FCBB68608FE3115B" box="[782,947,1097,1119]" pageId="7" pageNumber="706" value="2018-07-16" valueMax="2018-07-30" valueMin="2018-07-16">
<collectingDate id="7C76E9C5FFF69213FCBB68608FE3115B" box="[782,947,1097,1119]" pageId="7" pageNumber="706" value="2018-07-16" valueMax="2018-07-30" valueMin="2018-07-16">16–30 July 2018</collectingDate>
</date>
” (
<emphasis id="2AF8EAFFFFF69213FC7B68608F8A115B" box="[974,986,1098,1119]" italics="true" pageId="7" pageNumber="706">n</emphasis>
= 2); “
<date id="6C32102DFFF69213FB9268608F77117D" pageId="7" pageNumber="706" value="2018-07-30" valueMax="2018-08-17" valueMin="2018-07-30">
<collectingDate id="7C76E9C5FFF69213FB9268608F77117D" pageId="7" pageNumber="706" value="2018-07-30" valueMax="2018-08-17" valueMin="2018-07-30">30 July–17 August 2018</collectingDate>
</date>
” (
<emphasis id="2AF8EAFFFFF69213FC8B684F8F1A117E" box="[830,842,1125,1146]" italics="true" pageId="7" pageNumber="706">n</emphasis>
= 1); “
<date id="6C32102DFFF69213FC39684E8825117D" box="[908,1141,1124,1146]" pageId="7" pageNumber="706" value="2017-07-28" valueMax="2017-08-11" valueMin="2017-07-28">
<collectingDate id="7C76E9C5FFF69213FC39684E8825117D" box="[908,1141,1124,1146]" pageId="7" pageNumber="706" value="2017-07-28" valueMax="2017-08-11" valueMin="2017-07-28">28 July–11 August 2017</collectingDate>
</date>
,
<collectorName id="B579533BFFF69213FBCB684E8EBC1190" pageId="7" pageNumber="706">B. Steury</collectorName>
,
<collectorName id="B579533BFFF69213FD4368558F1D1190" box="[758,845,1151,1172]" pageId="7" pageNumber="706">C. Davis</collectorName>
,
<collectorName id="B579533BFFF69213FCE268558FE61190" box="[855,950,1151,1172]" pageId="7" pageNumber="706">C. Acosta</collectorName>
” (
<emphasis id="2AF8EAFFFFF69213FC7A68558F8B1190" box="[975,987,1151,1172]" italics="true" pageId="7" pageNumber="706">n</emphasis>
= 2); “
<collectingCountry id="609B767DFFF69213FB94685588351190" box="[1057,1125,1151,1172]" name="Turkey" pageId="7" pageNumber="706">Turkey</collectingCountry>
Run Park, river trail,
<collectingMethod id="C1CD4EFAFFF69213FCE768B38F9D11AA" box="[850,973,1177,1198]" pageId="7" pageNumber="706">Malaise trap</collectingMethod>
,
<date id="6C32102DFFF69213FC6C68B38E8F11CD" pageId="7" pageNumber="706" value="2009-07-31" valueMax="2009-08-17" valueMin="2009-07-31">
<collectingDate id="7C76E9C5FFF69213FC6C68B38E8F11CD" pageId="7" pageNumber="706" value="2009-07-31" valueMax="2009-08-17" valueMin="2009-07-31">31 July–17 August 2009</collectingDate>
</date>
,
<collectorName id="B579533BFFF69213FD5E689E8F1211CD" box="[747,834,1204,1225]" pageId="7" pageNumber="706">D. Smith</collectorName>
” (
<emphasis id="2AF8EAFFFFF69213FCE9689E8F3811CD" box="[860,872,1204,1225]" italics="true" pageId="7" pageNumber="706">n</emphasis>
= 1)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF6921DFD7068E58E12163E" lastPageId="9" lastPageNumber="708" pageId="7" pageNumber="706" type="description">
<paragraph id="183336EDFFF6921DFD7068E58E6014FF" blockId="7.[685,1173,859,1730]" lastBlockId="9.[151,638,194,1730]" lastPageId="9" lastPageNumber="708" pageId="7" pageNumber="706">
<emphasis id="2AF8EAFFFFF69213FD7068E58FAE11E0" bold="true" box="[709,1022,1231,1252]" pageId="7" pageNumber="706">
Description.
<typeStatus id="C737884FFFF69213FCE668E58FE111E0" box="[851,945,1231,1252]" pageId="7" pageNumber="706" type="holotype">Holotype</typeStatus>
, male.
</emphasis>
Body length
<quantity id="DF749B08FFF69213FD1868C38EA711FA" box="[685,759,1257,1278]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="706" unit="mm" value="3.3">3.3 mm</quantity>
to tip of elytra,
<quantity id="DF749B08FFF69213FC2468C38F8A11FA" box="[913,986,1257,1278]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="706" unit="mm" value="4.2">4.2 mm</quantity>
to tip of pygidium.
<emphasis id="2AF8EAFFFFF69213FD18692E8F18101D" bold="true" box="[685,840,1284,1305]" pageId="7" pageNumber="706">Dorsal habitus</emphasis>
robust; head black; labrum yellowish; pronotum black with rufous edging on base of midbasal lobe, basal corners and lateral edges; elytra pale rufous, with paler humeral angle.
<emphasis id="2AF8EAFFFFF69213FBD069798F721087" bold="true" pageId="7" pageNumber="706">Ventral habitus</emphasis>
black with rufous edging basally on all ventrites. Front and middle legs yellowish, hind legs pale brown.
<emphasis id="2AF8EAFFFFF69213FC9669898FD410BC" bold="true" box="[803,900,1443,1464]" pageId="7" pageNumber="706">Pygidium</emphasis>
black, somewhat stout, 2.2× length of hypopygidium.
<emphasis id="2AF8EAFFFFF69213FC166994884610D7" bold="true" box="[931,1046,1470,1491]" pageId="7" pageNumber="706">Pubescence</emphasis>
dorsally and ventrally recumbent, pale, slightly shorter on head than on elytra and pronotum.
<emphasis id="2AF8EAFFFFF69213FC5169D98844130C" bold="true" box="[996,1044,1523,1544]" pageId="7" pageNumber="706">Eyes</emphasis>
small, oval, entire, with many short, erect hairs, temple absent.
<emphasis id="2AF8EAFFFFF69213FD186A028F5E1339" bold="true" box="[685,782,1576,1597]" pageId="7" pageNumber="706">Antennae</emphasis>
(
<figureCitation id="80B72A68FFF69213FC976A028F221339" box="[802,882,1576,1597]" captionStart="Fig" captionStartId="8.[197,228,1543,1562]" captionTargetBox="[179,1188,196,1525]" captionTargetId="figure-58@8.[175,1190,194,1527]" captionTargetPageId="8" captionText="Fig. 9. Pseudotolida syphaxi, new species, holotype, male. A) Dorsal, B) Lateral, C) Antenna, D) Terminal maxillary palpomeres. Virginia, Fairfax County, Turkey Run Park, gulch, Malaise trap, 19–30 June 2009, D. Smith and B. Steury. Length 3.5 mm to tip of elytra, 4.2 mm to tip of pygidium." figureDoi="http://doi.org/10.5281/zenodo.4791335" httpUri="https://zenodo.org/record/4791335/files/figure.png" pageId="7" pageNumber="706">Fig. 9C</figureCitation>
) with antennomeres all yellowish, setose with recumbent, anteriorly directed setae and a few erect setae near the apex of each antennomere, reaching to base of pronotum, antennomeres 3 and 4 the shortest and subequal, 1 and 2 the widest, 5 one-third longer than 4 and subequal to 2, 5–11 slightly lengthening toward apex, 11 the longest.
<emphasis id="2AF8EAFFFFF9921CFE376A528CB913A3" bold="true" pageId="8" pageNumber="707">Terminal maxillary palpomeres</emphasis>
pale brown, globular, malleiform with slightly concave apex (
<figureCitation id="80B72A68FFF9921CFE066A878E5E13C6" box="[435,526,1709,1730]" captionStart="Fig" captionStartId="8.[197,228,1543,1562]" captionTargetBox="[179,1188,196,1525]" captionTargetId="figure-58@8.[175,1190,194,1527]" captionTargetPageId="8" captionText="Fig. 9. Pseudotolida syphaxi, new species, holotype, male. A) Dorsal, B) Lateral, C) Antenna, D) Terminal maxillary palpomeres. Virginia, Fairfax County, Turkey Run Park, gulch, Malaise trap, 19–30 June 2009, D. Smith and B. Steury. Length 3.5 mm to tip of elytra, 4.2 mm to tip of pygidium." figureDoi="http://doi.org/10.5281/zenodo.4791335" httpUri="https://zenodo.org/record/4791335/files/figure.png" pageId="8" pageNumber="707">Fig. 9D</figureCitation>
).
<emphasis id="2AF8EAFFFFF9921CFD996A878EC313C6" bold="true" box="[556,659,1709,1730]" pageId="8" pageNumber="707">Pronotum</emphasis>
broadest just before the base, the midbasal lobe convex, lateral margins slightly concave near base.
<emphasis id="2AF8EAFFFFF9921CFD766AB88F3013A3" bold="true" box="[707,864,1682,1703]" pageId="8" pageNumber="707">Scutellar shield</emphasis>
black with rufous anterior central area, small and triangular, the apex rounded.
<emphasis id="2AF8EAFFFFF9921CFBDD6A8788F813C6" bold="true" box="[1128,1192,1709,1730]" pageId="8" pageNumber="707">Elytra</emphasis>
together 1.9 times as long as broad, broadest at basal third, gradually narrowing apically, narrower at base than pronotum.
<emphasis id="2AF8EAFFFFF8921DFE9E6CDD8D321408" bold="true" box="[299,354,247,268]" pageId="9" pageNumber="708">Legs:</emphasis>
Metatibia with two spines, the inner one longest, half the length of metatarsomere 1, the outer spine half the length of the inner one; one subapical and three lateral subequal ridges, the second slightly more oblique than, and thus converging at the hind margin with, the first; metatarsomere 1 with three slightly oblique ridges and three black spinules anterior to the proximal ridge, metatarsomere 2 with one ridge; pro- and mesotarsomeres 4 short and deeply emarginate.
</paragraph>
<caption id="4CF36665FFF69213FF1A6A558D6A13C6" ID-DOI="http://doi.org/10.5281/zenodo.4791321" ID-Zenodo-Dep="4791321" httpUri="https://zenodo.org/record/4791321/files/figure.png" pageId="7" pageNumber="706" startId="7.[175,206,1663,1682]" targetBox="[155,629,1226,1644]" targetPageId="7">
<paragraph id="183336EDFFF69213FF1A6A558D6A13C6" blockId="7.[151,639,1662,1730]" pageId="7" pageNumber="706">
<emphasis id="2AF8EAFFFFF69213FF1A6A558CBC1396" bold="true" box="[175,236,1663,1682]" pageId="7" pageNumber="706">Fig. 8.</emphasis>
Parameres.
<taxonomicName id="DF8C4D6EFFF69213FEDC6A548E5E1395" authorityName="Liljeblad" authorityYear="1917" box="[361,526,1662,1681]" class="Insecta" family="Mordellidae" genus="Mordellistena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="706" phylum="Arthropoda" rank="species" species="vera">
<emphasis id="2AF8EAFFFFF69213FEDC6A548E5E1395" box="[361,526,1662,1681]" italics="true" pageId="7" pageNumber="706">Mordellistena vera</emphasis>
</taxonomicName>
: A) Dorsal, B) Lateral.
<taxonomicName id="DF8C4D6EFFF69213FEB26ABC8D8413AD" authority="Steury &amp; Steiner, 2020" authorityName="Steury &amp; Steiner" authorityYear="2020" box="[263,468,1686,1705]" class="Insecta" family="Mordellidae" genus="Mordellistena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="706" phylum="Arthropoda" rank="species" species="virginica" status="sp. nov.">
<emphasis id="2AF8EAFFFFF69213FEB26ABC8D8413AD" box="[263,468,1686,1705]" italics="true" pageId="7" pageNumber="706">Mordellistena virginica</emphasis>
</taxonomicName>
,
<emphasis id="2AF8EAFFFFF69213FE526ABD8E0613AE" bold="true" box="[487,598,1687,1706]" pageId="7" pageNumber="706">
<taxonomicNameLabel id="31CB5784FFF69213FE526ABD8E0613AE" box="[487,598,1687,1706]" pageId="7" pageNumber="706" rank="species">new species</taxonomicNameLabel>
</emphasis>
: C) Dorsal, D) Lateral.
</paragraph>
</caption>
<caption id="4CF36665FFF9921CFF706A2D8F7D134E" ID-DOI="http://doi.org/10.5281/zenodo.4791335" ID-Zenodo-Dep="4791335" httpUri="https://zenodo.org/record/4791335/files/figure.png" pageId="8" pageNumber="707" startId="8.[197,228,1543,1562]" targetBox="[179,1188,196,1525]" targetPageId="8">
<paragraph id="183336EDFFF9921CFF706A2D8F7D134E" blockId="8.[173,1193,1542,1610]" pageId="8" pageNumber="707">
<emphasis id="2AF8EAFFFFF9921CFF706A2D8D51131E" bold="true" box="[197,257,1543,1562]" pageId="8" pageNumber="707">Fig. 9.</emphasis>
<taxonomicName id="DF8C4D6EFFF9921CFEA26A2C8D81131D" authority="Steury &amp; Steiner, 2020" authorityName="Steury &amp; Steiner" authorityYear="2020" box="[279,465,1542,1561]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="707" phylum="Arthropoda" rank="species" species="syphaxi" status="sp. nov.">
<emphasis id="2AF8EAFFFFF9921CFEA26A2C8D81131D" box="[279,465,1542,1561]" italics="true" pageId="8" pageNumber="707">Pseudotolida syphaxi</emphasis>
</taxonomicName>
,
<emphasis id="2AF8EAFFFFF9921CFE546A2D8E1F131E" bold="true" box="[481,591,1543,1562]" pageId="8" pageNumber="707">
<taxonomicNameLabel id="31CB5784FFF9921CFE546A2D8E1F131E" box="[481,591,1543,1562]" pageId="8" pageNumber="707" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype, male. A) Dorsal, B) Lateral, C) Antenna, D) Terminal maxillary palpomeres. Virginia, Fairfax County, Turkey Run Park, gulch, Malaise trap, 19–30 June 2009, D. Smith and B. Steury. Length 3.5 mm to tip of elytra, 4.2 mm to tip of pygidium.
</paragraph>
</caption>
<paragraph id="183336EDFFF8921DFF1A6E2B8E12163E" blockId="9.[151,638,194,1730]" pageId="9" pageNumber="708">
<emphasis id="2AF8EAFFFFF8921DFF1A6E2B8D231712" bold="true" box="[175,371,513,534]" pageId="9" pageNumber="708">Probable Females.</emphasis>
<collectingMunicipality id="F857AC97FFF8921DFE376E2B8D891712" box="[386,473,513,534]" pageId="9" pageNumber="708">Differing</collectingMunicipality>
from males in having scalene triangular terminal maxillary palpomeres, slightly shorter antennomeres 5–11, and a less prominent or absent pale humeral angle, the elytra thus more uniformly rufous colored.
<location id="1D536036FFF8921DFDF46E418C99179F" LSID="urn:lsid:plazi:treatment:902587FBFFF6921DFD646EEF8FAD166B:1D536036FFF8921DFDF46E418C99179F" county="Fairfax Co." municipality="Differing" name="Specimens" pageId="9" pageNumber="708" stateProvince="Virginia">Specimens</location>
(
<emphasis id="2AF8EAFFFFF8921DFF636EAC8CB2179F" box="[214,226,646,667]" italics="true" pageId="9" pageNumber="708">n</emphasis>
= 9, GWMP):
<location id="1D536036FFF8921DFED96EAC8DDA179F" LSID="urn:lsid:plazi:treatment:902587FBFFF6921DFD646EEF8FAD166B:1D536036FFF8921DFED96EAC8DDA179F" box="[364,394,646,667]" county="Fairfax Co." municipality="Differing" name="All" pageId="9" pageNumber="708" stateProvince="Virginia">All</location>
labeled “
<collectingRegion id="DA48F80FFFF8921DFE576EAC8E62179F" box="[482,562,646,667]" country="United States of America" name="Virginia" pageId="9" pageNumber="708">Virginia</collectingRegion>
,
<collectingCounty id="F1524E61FFF8921DFD8F6EAC8CEC17B1" pageId="9" pageNumber="708">Fairfax Co.</collectingCounty>
, GWMP” and “Little Hunting Creek,
<collectingMethod id="C1CD4EFAFFF8921DFD866E8A8CEE17D4" pageId="9" pageNumber="708">Malaise trap</collectingMethod>
,
<date id="6C32102DFFF8921DFF7B6E918D2C17D4" box="[206,380,698,720]" pageId="9" pageNumber="708" value="2017-07-17" valueMax="2017-07-28" valueMin="2017-07-17">
<collectingDate id="7C76E9C5FFF8921DFF7B6E918D2C17D4" box="[206,380,698,720]" pageId="9" pageNumber="708" value="2017-07-17" valueMax="2017-07-28" valueMin="2017-07-17">17–28 July 2017</collectingDate>
</date>
,
<collectorName id="B579533BFFF8921DFE396E918DBD17D4" box="[396,493,699,720]" pageId="9" pageNumber="708">B. Steury</collectorName>
,
<collectorName id="B579533BFFF8921DFE496E918E0817D4" box="[508,600,699,720]" pageId="9" pageNumber="708">C. Davis</collectorName>
,
<collectorName id="B579533BFFF8921DFDDD6E918C8917EE" pageId="9" pageNumber="708">C. Acosta</collectorName>
” (
<emphasis id="2AF8EAFFFFF8921DFF426EFF8D5317EE" box="[247,259,725,746]" italics="true" pageId="9" pageNumber="708">n</emphasis>
= 1); same data except “28 July–11 August” (
<emphasis id="2AF8EAFFFFF8921DFF466EDA8CAF1601" box="[243,255,752,773]" italics="true" pageId="9" pageNumber="708">n</emphasis>
= 2); same data except “
<date id="6C32102DFFF8921DFE506EDA8C99161B" pageId="9" pageNumber="708" value="2018-06-28" valueMax="2018-07-16" valueMin="2018-06-28">
<collectingDate id="7C76E9C5FFF8921DFE506EDA8C99161B" pageId="9" pageNumber="708" value="2018-06-28" valueMax="2018-07-16" valueMin="2018-06-28">28 June–16 July 2018</collectingDate>
</date>
,
<collectorName id="B579533BFFF8921DFF616F208D7E161B" box="[212,302,778,799]" pageId="9" pageNumber="708">B. Steury</collectorName>
” (
<emphasis id="2AF8EAFFFFF8921DFEF26F218D031624" box="[327,339,779,800]" italics="true" pageId="9" pageNumber="708">n</emphasis>
= 2), same data except “16–30 July” (
<emphasis id="2AF8EAFFFFF8921DFF6C6F0F8CB5163E" box="[217,229,805,826]" italics="true" pageId="9" pageNumber="708">n</emphasis>
= 1); “30 July–17 August” (
<emphasis id="2AF8EAFFFFF8921DFE4A6F0F8E5B163E" box="[511,523,805,826]" italics="true" pageId="9" pageNumber="708">n</emphasis>
= 3).
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF8921DFF1A6F6A8E12117D" pageId="9" pageNumber="708" type="diagnosis">
<paragraph id="183336EDFFF8921DFF1A6F6A8E12117D" blockId="9.[151,638,194,1730]" pageId="9" pageNumber="708">
<emphasis id="2AF8EAFFFFF8921DFF1A6F6A8D471651" bold="true" box="[175,279,832,853]" pageId="9" pageNumber="708">Diagnosis.</emphasis>
<taxonomicName id="DF8C4D6EFFF8921DFEAB6F158DBA1651" authorityName="Steury &amp; Steiner" authorityYear="2020" box="[286,490,831,853]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="species" species="syphaxi">
<emphasis id="2AF8EAFFFFF8921DFEAB6F158DBA1651" box="[286,490,831,853]" italics="true" pageId="9" pageNumber="708">Pseudotolida syphaxi</emphasis>
</taxonomicName>
has elytral and ventral coloration similar to
<taxonomicName id="DF8C4D6EFFF8921DFE0D6F738D6F168E" authority="(Liljeblad, 1945)" baseAuthorityName="Liljeblad" baseAuthorityYear="1945" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="species" species="knausi">
<emphasis id="2AF8EAFFFFF8921DFE0D6F738E2E166B" box="[440,638,857,879]" italics="true" pageId="9" pageNumber="708">Pseudotolida knausi</emphasis>
(
<bibRefCitation id="7C1D4B1CFFF8921DFF2B6F5F8D68168E" author="Liljeblad, E." box="[158,312,885,906]" pageId="9" pageNumber="708" pagination="1 - 229" refId="ref6627" refString="Liljeblad, E. 1945. Monograph of the family Mordellidae (Coleoptera) of North America, north of Mexico. Miscellaneous Publications, Museum of Zoology, University of Michigan 62: 1 - 229." type="journal article" year="1945">Liljeblad, 1945</bibRefCitation>
)
</taxonomicName>
(a species only reported from Cedar City,
<collectingRegion id="DA48F80FFFF8921DFEB86FA58D6D16A0" box="[269,317,911,932]" country="United States of America" name="Utah" pageId="9" pageNumber="708">Utah</collectingRegion>
, at an elevation of
<quantity id="DF749B08FFF8921DFE4F6FA58E1C16A0" box="[506,588,911,932]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="9" pageNumber="708" unit="m" value="1950.0">1,950 m</quantity>
), but differs from that species in the paler head and pronotum, having an additional metatibial ridge, one less ridge on metatarsomere 3, and the shape of the male terminal maxillary palpomeres. A key to
<taxonomicName id="DF8C4D6EFFF8921DFF2268398DE7112D" authority="Ermisch, 1950" authorityName="Ermisch" authorityYear="1950" box="[151,439,1043,1065]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAFFFFF8921DFF2268398D44112D" box="[151,276,1043,1065]" italics="true" pageId="9" pageNumber="708">Pseudotolida</emphasis>
Ermisch, 1950
</taxonomicName>
of the world was provided by
<bibRefCitation id="7C1D4B1CFFF8921DFEA668058D9B1140" author="Franciscolo, M." box="[275,459,1071,1092]" pageId="9" pageNumber="708" pagination="49 - 72" refId="ref6435" refString="Franciscolo, M. 1982. Mordellidae (Coleoptera: Heteromera) from Rennell and Bellona Islands (Solomons). The Natural History of Rennell Island, British Solomon Islands 8: 49 - 72." type="journal article" year="1982">Franciscolo (1982)</bibRefCitation>
. Females were associated by their presence in the same Malaise traps at Little Hunting Creek that captured males.
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF8921DFF1A68558E471087" pageId="9" pageNumber="708" type="description">
<paragraph id="183336EDFFF8921DFF1A68558E471087" blockId="9.[151,638,194,1730]" pageId="9" pageNumber="708">
<emphasis id="2AF8EAFFFFF8921DFF1A68558D451190" bold="true" box="[175,277,1151,1172]" pageId="9" pageNumber="708">Variation.</emphasis>
Pronotum sometimes all black or the rufous edging more extensive; elytra with the midlateral and apical sutural margins and apical half of the elytra darkened to various extents (making the pale humeral angle more obvious); scutellar shield sometimes all black; anterior metatibial ridge sometimes weaker than the two below, metatarsomere 1 with or without a fourth obvious but weaker anterior ridge, the angle of convergence of the first and second ridges variable, sometimes more parallel.
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF8921DFF1A69A28E79130C" pageId="9" pageNumber="708" type="etymology">
<paragraph id="183336EDFFF8921DFF1A69A28E79130C" blockId="9.[151,638,194,1730]" pageId="9" pageNumber="708">
<emphasis id="2AF8EAFFFFF8921DFF1A69A28D721099" bold="true" box="[175,290,1416,1437]" pageId="9" pageNumber="708">Etymology.</emphasis>
<taxonomicName id="DF8C4D6EFFF8921DFE9969A28DAC109A" authorityName="Steury &amp; Steiner" authorityYear="2020" box="[300,508,1416,1438]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="species" species="syphaxi">
<emphasis id="2AF8EAFFFFF8921DFE9969A28DAC109A" box="[300,508,1416,1438]" italics="true" pageId="9" pageNumber="708">Pseudotolida syphaxi</emphasis>
</taxonomicName>
is named in honor of Stephen Syphax, former Chief of Resources for National Capital Parks East, for his devotion to the protection of the natural and cultural resources of the National Capital Region.
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF8921DFF1A6A278FFF1734" pageId="9" pageNumber="708" type="biology_ecology">
<paragraph id="183336EDFFF8921DFF1A6A278FFF1734" blockId="9.[151,638,194,1730]" lastBlockId="9.[685,1172,193,879]" pageId="9" pageNumber="708">
<emphasis id="2AF8EAFFFFF8921DFF1A6A278D511326" bold="true" box="[175,257,1549,1570]" pageId="9" pageNumber="708">Biology.</emphasis>
<taxonomicName id="DF8C4D6EFFF8921DFEB36A278D9C1327" authorityName="Steury &amp; Steiner" authorityYear="2020" box="[262,460,1549,1571]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="species" species="syphaxi">
<emphasis id="2AF8EAFFFFF8921DFEB36A278D9C1327" box="[262,460,1549,1571]" italics="true" pageId="9" pageNumber="708">Pseudotolida syphaxi</emphasis>
</taxonomicName>
is currently known from
<specimenCount id="0E8AFD64FFF8921DFF7C6A028D431339" box="[201,275,1576,1597]" pageId="9" pageNumber="708" type="male">13 male</specimenCount>
and
<specimenCount id="0E8AFD64FFF8921DFE8B6A028DF91339" box="[318,425,1576,1597]" pageId="9" pageNumber="708" type="female">nine female</specimenCount>
specimens collected in the Piedmont and Coastal Plain physiographic provinces of
<collectingRegion id="DA48F80FFFF8921DFEB86A778D0B1376" box="[269,347,1629,1650]" country="United States of America" name="Virginia" pageId="9" pageNumber="708">Virginia</collectingRegion>
. In the Piedmont it was found in mature upland forest dominated by broadleaved hardwoods, especially tulip poplar (
<taxonomicName id="DF8C4D6EFFF8921DFDB66ABB8CBC13C6" class="Magnoliopsida" family="Magnoliaceae" genus="Liriodendron" kingdom="Plantae" order="Magnoliales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="tulipifera">
<emphasis id="2AF8EAFFFFF8921DFDB66ABB8CBC13C6" italics="true" pageId="9" pageNumber="708">Liriodendron tulipifera</emphasis>
</taxonomicName>
L.;
<taxonomicName id="DF8C4D6EFFF8921DFE906A878DFA13C6" authorityName="de Jussieu" authorityYear="1789" box="[293,426,1709,1730]" class="Magnoliopsida" family="Magnoliaceae" kingdom="Plantae" order="Magnoliales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="family">Magnoliaceae</taxonomicName>
), sugar maple (
<taxonomicName id="DF8C4D6EFFF8921DFDE76A868F2115D3" authority="Marshall" authorityName="Marshall" authorityYear="1785" box="[594,881,193,1730]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="saccharum">
<emphasis id="2AF8EAFFFFF8921DFDE76A868F4315D3" box="[594,787,193,1730]" italics="true" pageId="9" pageNumber="708">Acer saccharum</emphasis>
Marshall
</taxonomicName>
;
<taxonomicName id="DF8C4D6EFFF8921DFCCA6CE88FA615D3" box="[895,1014,194,215]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
), oaks (
<taxonomicName id="DF8C4D6EFFF8921DFBF36CEB8E9A15F5" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2AF8EAFFFFF8921DFBF36CEB88C415D3" box="[1094,1172,193,215]" italics="true" pageId="9" pageNumber="708">Quercus</emphasis>
sp.
</taxonomicName>
), and hickory (
<taxonomicName id="DF8C4D6EFFF8921DFCE26CF18FE515F5" box="[855,949,219,241]" class="Magnoliopsida" family="Juglandaceae" genus="Carya" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2AF8EAFFFFF8921DFCE26CF18FC115F5" box="[855,913,219,241]" italics="true" pageId="9" pageNumber="708">Carya</emphasis>
sp.
</taxonomicName>
). The site overlooks the Potomac Gorge and has a diverse spring ephemeral flora. On the Coastal Plain it was found in mature forest dominated by broadleaved hardwoods, especially oaks (
<taxonomicName id="DF8C4D6EFFF8921DFC966D6C8FC5145F" box="[803,917,326,348]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2AF8EAFFFFF8921DFC966D6C8F211458" box="[803,881,326,348]" italics="true" pageId="9" pageNumber="708">Quercus</emphasis>
sp.
</taxonomicName>
), hickory (
<taxonomicName id="DF8C4D6EFFF8921DFC4A6D6C880E145F" box="[1023,1118,326,348]" class="Magnoliopsida" family="Juglandaceae" genus="Carya" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2AF8EAFFFFF8921DFC4A6D6C88691458" box="[1023,1081,326,348]" italics="true" pageId="9" pageNumber="708">Carya</emphasis>
sp.
</taxonomicName>
), and American beech (
<taxonomicName id="DF8C4D6EFFF8921DFCD46D4B88401472" box="[865,1040,352,374]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="grandifolia">
<emphasis id="2AF8EAFFFFF8921DFCD46D4B88401472" box="[865,1040,352,374]" italics="true" pageId="9" pageNumber="708">Fagus grandifolia</emphasis>
</taxonomicName>
), with some
<collectingRegion id="DA48F80FFFF8921DFD186D568EA81495" box="[685,760,380,401]" country="United States of America" name="Virginia" pageId="9" pageNumber="708">Virginia</collectingRegion>
pine (
<taxonomicName id="DF8C4D6EFFF8921DFC816D5688541495" authority="Mill." authorityName="Mill." box="[820,1028,380,401]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="virginiana">
<emphasis id="2AF8EAFFFFF8921DFC816D568F9E1495" box="[820,974,380,401]" italics="true" pageId="9" pageNumber="708">Pinus virginiana</emphasis>
Mill.
</taxonomicName>
) and a strongly ericaceous understory dominated by blueberry and deerberry (
<taxonomicName id="DF8C4D6EFFF8921DFCA26D9B8FF914C2" box="[791,937,433,454]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2AF8EAFFFFF8921DFCA26D9B8F2714C2" box="[791,887,433,454]" italics="true" pageId="9" pageNumber="708">Vaccinium</emphasis>
spp.
</taxonomicName>
) and huckleberry (
<taxonomicName id="DF8C4D6EFFF8921DFBD06D9A8F7014E4" class="Magnoliopsida" family="Ericaceae" genus="Gaylussacia" kingdom="Plantae" order="Ericales" pageId="9" pageNumber="708" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="2AF8EAFFFFF8921DFBD06D9A8EA914E4" italics="true" pageId="9" pageNumber="708">Gaylussacia</emphasis>
sp.
</taxonomicName>
) located approximately
<quantity id="DF749B08FFF8921DFBA56DE1881514E4" box="[1040,1093,459,480]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="708" unit="m" value="20.0">20 m</quantity>
from a narrow band of shrubby marsh along the bank of a large creek. It was collected in Malaise traps set during 19 June–17 August.
</paragraph>
</subSubSection>
<subSubSection id="50966566FFF8921DFD706E1F8FAD166B" pageId="9" pageNumber="708" type="discussion">
<paragraph id="183336EDFFF8921DFD706E1F8FAD166B" blockId="9.[685,1172,193,879]" pageId="9" pageNumber="708">
<emphasis id="2AF8EAFFFFF8921DFD706E1F8F68174E" bold="true" box="[709,824,565,586]" pageId="9" pageNumber="708">Comments.</emphasis>
Two genera of North American
<taxonomicName id="DF8C4D6EFFF8921DFD186E7A8F701761" authorityName="Latreille" authorityYear="1802" box="[685,800,592,613]" class="Insecta" family="Mordellidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="family">Mordellidae</taxonomicName>
possess malleiform terminal maxillary palpomeres in males,
<taxonomicName id="DF8C4D6EFFF8921DFC316E4088C41784" authority="Emery, 1876" authorityName="Emery" authorityYear="1876" box="[900,1172,618,640]" class="Insecta" family="Mordellidae" genus="Mordellochroa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAFFFFF8921DFC316E40885F1784" box="[900,1039,618,640]" italics="true" pageId="9" pageNumber="708">Mordellochroa</emphasis>
Emery, 1876
</taxonomicName>
and
<taxonomicName id="DF8C4D6EFFF8921DFD666EAF8F1D179F" authorityName="Ermisch" authorityYear="1950" box="[723,845,645,667]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAFFFFF8921DFD666EAF8F1D179F" box="[723,845,645,667]" italics="true" pageId="9" pageNumber="708">Pseudotolida</emphasis>
</taxonomicName>
.
<taxonomicName id="DF8C4D6EFFF8921DFCED6EAF884D179F" authorityName="Steury &amp; Steiner" authorityYear="2020" box="[856,1053,645,667]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="species" species="syphaxi">
<emphasis id="2AF8EAFFFFF8921DFCED6EAF884D179F" box="[856,1053,645,667]" italics="true" pageId="9" pageNumber="708">Pseudotolida syphaxi</emphasis>
</taxonomicName>
has the body size, narrowly emarginate pro- and mesotarsomeres 4, and long, oblique, somewhat converging metatibial ridges as in
<taxonomicName id="DF8C4D6EFFF8921DFCFC6EFE8F9317EE" authorityName="Ermisch" authorityYear="1950" box="[841,963,724,746]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAFFFFF8921DFCFC6EFE8F9317EE" box="[841,963,724,746]" italics="true" pageId="9" pageNumber="708">Pseudotolida</emphasis>
</taxonomicName>
; however, its eyes are small, oval, and entire (unemarginated), which is more similar to the eyes of
<taxonomicName id="DF8C4D6EFFF8921DFC026F238812161B" authorityName="Emery" authorityYear="1876" box="[951,1090,777,799]" class="Insecta" family="Mordellidae" genus="Mordellochroa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAFFFFF8921DFC026F238812161B" box="[951,1090,777,799]" italics="true" pageId="9" pageNumber="708">Mordellochroa</emphasis>
</taxonomicName>
. Further study is needed to better understand the relationship of the one North American species of
<taxonomicName id="DF8C4D6EFFF8921DFBBC6F1488C41650" authorityName="Emery" authorityYear="1876" box="[1033,1172,830,852]" class="Insecta" family="Mordellidae" genus="Mordellochroa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAFFFFF8921DFBBC6F1488C41650" box="[1033,1172,830,852]" italics="true" pageId="9" pageNumber="708">Mordellochroa</emphasis>
</taxonomicName>
to the four species of
<taxonomicName id="DF8C4D6EFFF8921DFCC86F738FA7166B" authorityName="Ermisch" authorityYear="1950" box="[893,1015,857,879]" class="Insecta" family="Mordellidae" genus="Pseudotolida" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="708" phylum="Arthropoda" rank="genus">
<emphasis id="2AF8EAFFFFF8921DFCC86F738FA7166B" box="[893,1015,857,879]" italics="true" pageId="9" pageNumber="708">Pseudotolida</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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