173 lines
26 KiB
XML
173 lines
26 KiB
XML
<document id="B184286CA0AB5FCA56DD0BDB3F4A49EA" ID-CLB-Dataset="8118" ID-DOI="10.5252/z2010n1a2" ID-GBIF-Dataset="4cecd899-5900-4411-8446-0d9552b1b098" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612808309234" checkinUser="felipe" docAuthor="Randriamanantsoa, Richard, Aberlenc, Henri-Pierre, Ralisoa, Olga Bakoly, Ratnadass, Alain & Vercambre, Bernard" docDate="2010" docId="901F8794FF99746FFD6CFBC2FC48FCC5" docLanguage="fr" docName="Zoosystema.32.1.19-72.pdf" docOrigin="Zoosystema 32 (1)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n1a2" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Paranodon coquerelii" docType="treatment" docVersion="11" lastPageNumber="33" masterDocId="6C26FFECFF947461FFB5FFEDFF94FFE8" masterDocTitle="Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar" masterLastPageNumber="72" masterPageNumber="19" pageNumber="32" updateTime="1698928337295" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AA6953AC727030392277AD0A693455A3">Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar</mods:title>
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<mods:namePart id="DC690B6A2E6565E217FF7E6852AEF983">Randriamanantsoa, Richard</mods:namePart>
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<mods:affiliation id="2766FD059A13D658A2C82DFB0CB75C2B">Fofifa - URP Scrid, 110 Antsirabe (Madagascar) r _ randriamanantsoa @ yahoo. fr</mods:affiliation>
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<mods:namePart id="C5A52EAE8D2B5BEF51A1B7DBBB7D4E2D">Aberlenc, Henri-Pierre</mods:namePart>
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<mods:affiliation id="697E2A8CB4B07790F44004F8153A1B3F">Cirad Bios, UMR CBGP (INRA / IRD / Cirad / Montpellier SupAgro), Campus international de Baillarguet, Ta A- 55 / L, F- 34398 Montpellier cedex 5 (France) henri-pierre. aberlenc @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="3FA5C1CF90BAE72D18DAB369E250B109" type="email">aberlenc@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="0EEE8923F857AF429C93DB8D4D5F3A88">Ralisoa, Olga Bakoly</mods:namePart>
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<mods:affiliation id="D24F2098775E89D94111616612876051">Département d’Entomologie, Faculté des Sciences, Université d’Antananarivo, Antananarivo 101 (Madagascar)</mods:affiliation>
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<mods:namePart id="114A75F1DF0190CB2A306E0F39C4DAC8">Ratnadass, Alain</mods:namePart>
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<mods:affiliation id="5E0F0B157F95BDDDACA1B15E3CFBB89A">Cirad, UPR HortSys - ICRISAT BP 12404, Niamey (Niger) a. ratnadass @ cgiar. org</mods:affiliation>
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<mods:nameIdentifier id="32F4D09541E19F71DDF397301CA45F56" type="email">ratnadass@cgiar.org</mods:nameIdentifier>
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<mods:namePart id="C2376B6A1DFCB85CF7648F3639DB303C">Vercambre, Bernard</mods:namePart>
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<mods:affiliation id="3C6F6E770E180184EA32CC2A2D381A9F">Cirad, UPR Systèmes de Cultures annuels, avenue Agropolis, F- 34398 Montpellier cedex 5 (France) bernard. vercambre @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="9D78FEA65A7C1E353D3E7BC57FF2AA50" type="email">vercambre@cirad.fr</mods:nameIdentifier>
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<mods:title id="7697F49C7F260A1635A547265C7243BE">Zoosystema</mods:title>
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<mods:part id="3B03F03D6D6D68D2693EA158533E7365">
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<mods:date id="1063101F2DDC2D43AFA25A5CDABF35B7">2010</mods:date>
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<mods:number id="131650F8F17271D6453E526A728EFFFA">2010-03-31</mods:number>
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<mods:number id="92B4EDA09A76BCBB215EB1AE17EE204D">32</mods:number>
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<mods:url id="246C42608788DDBCE87E32024334AF44">http://www.bioone.org/doi/abs/10.5252/z2010n1a2</mods:url>
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<treatment id="901F8794FF99746FFD6CFBC2FC48FCC5" ID-DOI="http://doi.org/10.5281/zenodo.4557669" ID-GBIF-Taxon="177425655" ID-Zenodo-Dep="4557669" LSID="urn:lsid:plazi:treatment:901F8794FF99746FFD6CFBC2FC48FCC5" httpUri="http://treatment.plazi.org/id/901F8794FF99746FFD6CFBC2FC48FCC5" lastPageId="14" lastPageNumber="33" pageId="13" pageNumber="32">
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<subSubSection id="50AC6509FF99746CFD6CFBC2FCE1FB2F" pageId="13" pageNumber="32" type="nomenclature">
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<paragraph id="18093682FF99746CFD6CFBC2FB11FBA2" blockId="13.[729,1157,1071,1161]" box="[729,1157,1071,1098]" pageId="13" pageNumber="32">
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<taxonomicName id="DFB64D01FF99746CFD6CFBC2FB11FBA2" authority="(Fairmaire, 1871)" baseAuthorityName="Fairmaire" baseAuthorityYear="1871" box="[729,1157,1071,1098]" class="Insecta" family="Dynastidae" genus="Paranodon" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="32" phylum="Arthropoda" rank="species" species="coquerelii">
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<emphasis id="2AC2EA90FF99746CFD6CFBC2FC2BFBA2" bold="true" box="[729,959,1071,1098]" italics="true" pageId="13" pageNumber="32">Paranodon coquerelii</emphasis>
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(Fairmaire, 1871)
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</taxonomicName>
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</paragraph>
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<paragraph id="18093682FF99746CFD4AFBA2FBD1FB61" blockId="13.[729,1157,1071,1161]" pageId="13" pageNumber="32">
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(
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<figureCitation id="808D2A07FF99746CFCB3FBA2FCCCFB82" box="[774,856,1103,1130]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="13" pageNumber="32">Figs 4E</figureCitation>
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;
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<figureCitation id="808D2A07FF99746CFCDFFBA2FC1CFB82" box="[874,904,1103,1130]" captionStart="FIG" captionStartId="35.[113,124,1635,1652]" captionTargetBox="[141,1181,217,1589]" captionTargetId="figure-14@35.[141,1181,217,1589]" captionTargetPageId="35" captionText="FIG. 8. — Larves de Scarabaeoidea de Madagascar, épipharynx: A, Heteroconus paradoxus (Endrödi,1968); B, Paranodon coquerelii (Fairmaire, 1871); C, Paramorphochelus cornutus (Nonfried,1892); D, sp. M1; E, Encya sikorai (Brenske,1891); F, Enaria melanictera (Klug, 1833); G, sp. M2. Échelles: A, F, 0,5 mm; B, C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520775" httpUri="https://zenodo.org/record/4520775/files/figure.png" pageId="13" pageNumber="32">8B</figureCitation>
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;
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<figureCitation id="808D2A07FF99746CFC2FFBA2FC54FB82" box="[922,960,1103,1130]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory & Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="13" pageNumber="32">10J</figureCitation>
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;
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<figureCitation id="808D2A07FF99746CFC64FBA2FC63FB82" box="[977,1015,1103,1130]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="13" pageNumber="32">12J</figureCitation>
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;
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<figureCitation id="808D2A07FF99746CFBBDFBA2FBB5FB82" box="[1032,1057,1103,1130]" captionStart="FIG" captionStartId="42.[113,124,1589,1606]" captionTargetBox="[135,1177,220,1549]" captionTargetId="figure-21@42.[136,1178,214,1549]" captionTargetPageId="42" captionText="FIG. 15. — Larves de Scarabaeoidea de Madagascar, mandibule: A, E, I, M, Q, U, Y, gauche, face dorsale; B, F, J, N, R, V, Z1, gauche, face ventrale; C, G, K, O, S, W, Z2, droite,face dorsale; D, H, L, P, T, X, Z3, droite,face ventrale; A-D, Paranodon coquerelii (Fairmaire, 1871); E-H, Paramorphochelus cornutus (Nonfried, 1892); I-L, Encya sikorai (Brenske, 1891); M-P, Enaria melanictera (Klug, 1833); Q-T, Apicencya waterloti (Dewailly, 1950); U-X, Empecta scutata (Fairmaire, 1901); Y-Z3, Hoplochelus betanimena (Künckel, 1887). Échelles: A-D, M-P, 1 mm; E-H, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520799" httpUri="https://zenodo.org/record/4520799/files/figure.png" pageId="13" pageNumber="32">15</figureCitation>
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A-D; 17A; 19F; 22A-C; 24P, Q)
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</paragraph>
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<paragraph id="18093682FF99746CFD10FB43FCE1FB2F" blockId="13.[675,1211,1198,1383]" box="[677,885,1198,1225]" pageId="13" pageNumber="32">MATÉRIEL EXAMINÉ</paragraph>
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</subSubSection>
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<subSubSection id="50AC6509FF99746FFD10FB23FC48FCC5" lastPageId="14" lastPageNumber="33" pageId="13" pageNumber="32" type="description">
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<paragraph id="18093682FF99746CFD10FB23FB09FA8F" blockId="13.[675,1211,1198,1383]" pageId="13" pageNumber="32">
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Larves provenant d’Ampandrotrarana (
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<quantity id="DF4E9B67FF99746CFBD9FB23FB2DFB01" box="[1132,1209,1230,1257]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="32" unit="km" value="20.0">20 km</quantity>
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d’Antsirabe) dans une culture de Riz pluvial en haut de pente.
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<typeStatus id="C70D8820FF99746CFCF5FAE0FCE0FAC0" box="[832,884,1293,1320]" pageId="13" pageNumber="32">Type</typeStatus>
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de sol: ferrallitique. Spécimens conservés dans les collections du Fofifa (
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<specimenCount id="0EB0FD0BFF99746CFBF3FAC0FB19FAA0" box="[1094,1165,1325,1352]" pageId="13" pageNumber="32" type="larva">1 larve</specimenCount>
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), du Cirad-CBGP (
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<specimenCount id="0EB0FD0BFF99746CFCF1FAA1FC18FA8F" box="[836,908,1356,1383]" pageId="13" pageNumber="32" type="larva">1 larve</specimenCount>
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) et du MNHN (
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<specimenCount id="0EB0FD0BFF99746CFBF3FAA1FB05FA8F" box="[1094,1169,1356,1383]" pageId="13" pageNumber="32" type="larva">1 larve</specimenCount>
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).
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</paragraph>
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<paragraph id="18093682FF99746CFD10FA61FCA1FA4F" blockId="13.[677,864,1420,1478]" box="[677,821,1420,1447]" pageId="13" pageNumber="32">DESCRIPTION</paragraph>
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<paragraph id="18093682FF99746CFD10FA46FCF4FA2E" blockId="13.[677,864,1420,1478]" box="[677,864,1451,1478]" pageId="13" pageNumber="32">Couleur blanche.</paragraph>
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<paragraph id="18093682FF99746CFD10FA06FD5BF9ED" blockId="13.[677,1211,1515,1700]" box="[677,719,1515,1541]" pageId="13" pageNumber="32">
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<emphasis id="2AC2EA90FF99746CFD10FA06FD5BF9ED" box="[677,719,1515,1541]" italics="true" pageId="13" pageNumber="32">Tête</emphasis>
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</paragraph>
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<paragraph id="18093682FF99746FFD10F9E7FEEDFEFB" blockId="13.[677,1211,1515,1700]" lastBlockId="14.[113,646,216,275]" lastPageId="14" lastPageNumber="33" pageId="13" pageNumber="32">Couleur jaune. Épicrâne avec 3 soies dorso-épicraniales et 4 soies latéro-externes disposées symétriquement. Front avec 2 soies dans l’angle frontal, 1 soie frontale postérieure et 2 soies antérieures disposées symétriquement. Clypéus transverse avec 2 soies clypéales postérieures et 2 soies clypéales externes de chaque côté.</paragraph>
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<paragraph id="18093682FF9A746FFFC4FEDAFE5EFE18" blockId="14.[111,649,311,496]" pageId="14" pageNumber="33">
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<emphasis id="2AC2EA90FF9A746FFFC4FEDAFF4DFEB9" bold="true" box="[113,217,311,337]" pageId="14" pageNumber="33">Antennes</emphasis>
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(
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<figureCitation id="808D2A07FF9A746FFF52FEDAFED4FEBA" box="[231,320,311,338]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="14" pageNumber="33">Fig. 12J</figureCitation>
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). Les longueurs respectives des articles sont: I = 1,36; II = 1,54; III = 1,27; IV = 1. Une aire sensorielle apico-latérale sur l’avant-dernier article. Dernier article antennaire avec 5 plages sensorielles: 1 petite plage apicale ronde, 2 ventrales écartées et 2 dorsales adjacentes.
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</paragraph>
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<paragraph id="18093682FF9A746FFFC4FDF8FDA9FCC5" blockId="14.[111,649,533,813]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFFC4FDF8FF2CFDC7" bold="true" box="[113,184,533,559]" pageId="14" pageNumber="33">Labre.</emphasis>
|
||
Transverse, asymétrique, avec de longues soies.Zone médiane du bord antérieur non-saillante, différenciée du côté droit, avec 4 soies antérieures externes et2 soies antérieures internes rapprochées.De part et d’autre, 5 soies latéro-antérieures externes et 2 soies latéro-antérieures internes, puis 1 soie latérale.2 longues soies labro-internes en avant du milieu.Une rangée transverse de 7 soies postérieures en 2 groupes de 4 et 3 soies, la zone centrale étant glabre.
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFFC4FCBCFDB5FBE3" blockId="14.[112,649,849,1035]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFFC4FCBCFF66FC84" bold="true" box="[113,242,849,876]" pageId="14" pageNumber="33">Épipharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF9A746FFEB6FCBCFECCFC84" box="[259,344,849,876]" captionStart="FIG" captionStartId="35.[113,124,1635,1652]" captionTargetBox="[141,1181,217,1589]" captionTargetId="figure-14@35.[141,1181,217,1589]" captionTargetPageId="35" captionText="FIG. 8. — Larves de Scarabaeoidea de Madagascar, épipharynx: A, Heteroconus paradoxus (Endrödi,1968); B, Paranodon coquerelii (Fairmaire, 1871); C, Paramorphochelus cornutus (Nonfried,1892); D, sp. M1; E, Encya sikorai (Brenske,1891); F, Enaria melanictera (Klug, 1833); G, sp. M2. Échelles: A, F, 0,5 mm; B, C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520775" httpUri="https://zenodo.org/record/4520775/files/figure.png" pageId="14" pageNumber="33">Fig. 8B</figureCitation>
|
||
). Haptomère à bord relevé, sclérifié et émoussé. Clithrum sclérifié. Aire pédiale très petite, glabre. Chaetoparia dissymétriques, à fortes soies épaisses convergeant vers l’aire pédiale. Acanthoparia avec 9 courtes soies incurvées lamelliformes, de taille croissante vers l’avant.
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFFC4FBC2FDADFA6F" blockId="14.[111,649,1071,1415]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFFC4FBC2FF63FBA1" bold="true" box="[113,247,1071,1097]" pageId="14" pageNumber="33">Mandibules</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF9A746FFEBCFBC2FECCFBA2" box="[265,344,1071,1098]" captionStart="FIG" captionStartId="42.[113,124,1589,1606]" captionTargetBox="[135,1177,220,1549]" captionTargetId="figure-21@42.[136,1178,214,1549]" captionTargetPageId="42" captionText="FIG. 15. — Larves de Scarabaeoidea de Madagascar, mandibule: A, E, I, M, Q, U, Y, gauche, face dorsale; B, F, J, N, R, V, Z1, gauche, face ventrale; C, G, K, O, S, W, Z2, droite,face dorsale; D, H, L, P, T, X, Z3, droite,face ventrale; A-D, Paranodon coquerelii (Fairmaire, 1871); E-H, Paramorphochelus cornutus (Nonfried, 1892); I-L, Encya sikorai (Brenske, 1891); M-P, Enaria melanictera (Klug, 1833); Q-T, Apicencya waterloti (Dewailly, 1950); U-X, Empecta scutata (Fairmaire, 1901); Y-Z3, Hoplochelus betanimena (Künckel, 1887). Échelles: A-D, M-P, 1 mm; E-H, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520799" httpUri="https://zenodo.org/record/4520799/files/figure.png" pageId="14" pageNumber="33">Fig. 15</figureCitation>
|
||
A-D). Aire scissoriale de la mandibule gauche avec 4 dents, l’apicale forte, la deuxième arrondie et très émoussée, la troisième et la quatrième pointues et plus écartées que les autres. 5 soies latéro-externes et 2 soies dorso-externes. Aire scissoriale de la mandibule droite à 3 dents, l’apicale pointue, la deuxième arrondie et la troisième émoussée en angle obtus. 5 soies latéro-externes et 2 soies dorso-externes. Sur la face ventrale des 2 mandibules, présence d’une aire stridulatoire composée de 23 à 26 bourrelets parallèles.
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFFC4FA46FE67F9CD" blockId="14.[111,647,1451,1573]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFFC4FA46FF5EFA2D" bold="true" box="[113,202,1451,1477]" pageId="14" pageNumber="33">Maxilles</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF9A746FFF63FA46FEA7FA2E" box="[214,307,1451,1478]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="14" pageNumber="33">Fig. 17A</figureCitation>
|
||
). Galea et lacinia soudées.Lacinia à uncus avec 4 dents (3 grandes et 1 petite).Galea avec uncus sclérifié. Sur la face dorsale, aire stridulatoire composée de 9 à 10 dents crénelées.
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFFC4F9A7FCE6FEFB" blockId="14.[113,646,1610,1700]" lastBlockId="14.[674,1210,216,275]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFFC4F9A7FE9FF98C" bold="true" box="[113,267,1610,1636]" pageId="14" pageNumber="33">Hypopharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF9A746FFEACF9A7FEE7F98D" box="[281,371,1610,1637]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory & Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="14" pageNumber="33">Fig. 10J</figureCitation>
|
||
). Surface entre la base des palpes et les oncyli couverte de longues et fortes soies. Oncyli transverses, avec une dent sclérifiée postérieure très émoussée et une forte dent antérieure, sclérifiée et dressée.
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFD10FEDAFD79FEB9" blockId="14.[675,1210,311,465]" box="[677,749,311,337]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFD10FEDAFD79FEB9" box="[677,749,311,337]" italics="true" pageId="14" pageNumber="33">Thorax</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFD10FEBAFCE5FE39" blockId="14.[675,1210,311,465]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFD10FEBAFD73FE99" bold="true" box="[677,743,343,369]" pageId="14" pageNumber="33">Pattes</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF9A746FFD41FEBAFCC6FE9A" box="[756,850,343,370]" captionStart="FIG" captionStartId="46.[113,124,1635,1652]" captionTargetBox="[121,1206,215,1527]" captionTargetId="figure-21@46.[117,1206,214,1542]" captionTargetPageId="46" captionText="FIG. 19. — Larves de Scarabaeoidea de Madagascar, pattes: A, Heteronychus arator rugifrons (Fairmaire, 1871); B, Heteronychus bituberculatus (Kolbe, 1900); C, Heteronychus plebeius (Klug, 1833); D, Hexodon unicolor (Olivier, 1789); E, Heteroconus paradoxus (Endrödi, 1968); F, Paranodon coquerelii (Fairmaire, 1871). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520815" httpUri="https://zenodo.org/record/4520815/files/figure.png" pageId="14" pageNumber="33">Fig. 19F</figureCitation>
|
||
). Longues, de longueur croissante de P1 à P3, toutes de structure analogue: coxa, trochanter, fémur, tibiotarse, griffe longue, aiguë, incurvée, sclérifiée.
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFD10FE18FC93FDE7" blockId="14.[675,1212,501,813]" box="[677,775,501,527]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFD10FE18FC93FDE7" box="[677,775,501,527]" italics="true" pageId="14" pageNumber="33">Abdomen</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF9A746FFD10FDF8FC48FCC5" blockId="14.[675,1212,501,813]" pageId="14" pageNumber="33">
|
||
<emphasis id="2AC2EA90FF9A746FFD10FDF8FD7EFDC7" bold="true" box="[677,746,533,559]" pageId="14" pageNumber="33">Raster</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF9A746FFD43FDF8FCD3FDD8" box="[758,839,533,560]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="14" pageNumber="33">Fig. 4E</figureCitation>
|
||
). Fente anale transverse. Une bande glabre du côté dorsal de la fente anale, barbula composée de nombreuses longues soies fines. Une rangée de longues soies du côté ventral de la fente anale. Teges avec une trentaine de soies épaisses, aplaties et incurvées, avec de chaque côté une très forte et épaisse épine dissymétrique (trifide ou bifide du côté gauche et bifide ou simple du côté droit), séparées par une zone glabre.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |