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<document id="082590585B234A35DFD9DC046D4FFE61" ID-CLB-Dataset="8118" ID-DOI="10.5252/z2010n1a2" ID-GBIF-Dataset="4cecd899-5900-4411-8446-0d9552b1b098" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612808309234" checkinUser="felipe" docAuthor="Randriamanantsoa, Richard, Aberlenc, Henri-Pierre, Ralisoa, Olga Bakoly, Ratnadass, Alain &amp; Vercambre, Bernard" docDate="2010" docId="901F8794FF8D747AFEFFFF3AFBFAFAA0" docLanguage="fr" docName="Zoosystema.32.1.19-72.pdf" docOrigin="Zoosystema 32 (1)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n1a2" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Scarabaeoidea Latreille 1802" docType="key" docVersion="8" lastPageNumber="46" masterDocId="6C26FFECFF947461FFB5FFEDFF94FFE8" masterDocTitle="Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar" masterLastPageNumber="72" masterPageNumber="19" pageNumber="44" updateTime="1698928337295" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2DCFC86B0728551E64614CADC9000187">Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar</mods:title>
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<mods:namePart id="64B1E0EDB755A6692496FC087C8A0225">Randriamanantsoa, Richard</mods:namePart>
<mods:affiliation id="E457292E62B331B1B491E9B12C0C5FD0">Fofifa - URP Scrid, 110 Antsirabe (Madagascar) r _ randriamanantsoa @ yahoo. fr</mods:affiliation>
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<mods:namePart id="FD11C416B4D3A363582F34B61AD5A91B">Aberlenc, Henri-Pierre</mods:namePart>
<mods:affiliation id="3E1554A1BBD9B6E1FDBB37ED46635C2F">Cirad Bios, UMR CBGP (INRA / IRD / Cirad / Montpellier SupAgro), Campus international de Baillarguet, Ta A- 55 / L, F- 34398 Montpellier cedex 5 (France) henri-pierre. aberlenc @ cirad. fr</mods:affiliation>
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<mods:namePart id="EF5A479BF7BD9A3E22073AF78497F808">Ralisoa, Olga Bakoly</mods:namePart>
<mods:affiliation id="116FF17612A3D47030742B2BFB01D460">Département d’Entomologie, Faculté des Sciences, Université d’Antananarivo, Antananarivo 101 (Madagascar)</mods:affiliation>
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<mods:namePart id="3D9EFFBCD8BF66E0FAE00EAEFC4725EA">Ratnadass, Alain</mods:namePart>
<mods:affiliation id="B5284D9BFBD7142A7C921FAA1736A5DA">Cirad, UPR HortSys - ICRISAT BP 12404, Niamey (Niger) a. ratnadass @ cgiar. org</mods:affiliation>
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<mods:namePart id="D695045D821141FD1CF0A25D710CBFD4">Vercambre, Bernard</mods:namePart>
<mods:affiliation id="97C4F765AC148B2A63B2E6B8A2806104">Cirad, UPR Systèmes de Cultures annuels, avenue Agropolis, F- 34398 Montpellier cedex 5 (France) bernard. vercambre @ cirad. fr</mods:affiliation>
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<mods:date id="91A7CDD5CE2171BDC0064D1818FB294D">2010</mods:date>
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<treatment id="901F8794FF8D747AFEFFFF3AFBFAFAA0" ID-DOI="http://doi.org/10.5281/zenodo.4560182" ID-GBIF-Taxon="177425630" ID-Zenodo-Dep="4560182" LSID="urn:lsid:plazi:treatment:901F8794FF8D747AFEFFFF3AFBFAFAA0" httpUri="http://treatment.plazi.org/id/901F8794FF8D747AFEFFFF3AFBFAFAA0" lastPageId="27" lastPageNumber="46" pageId="25" pageNumber="44">
<subSubSection id="50AC6509FF8D7478FEFFFF3AFC7BFEF8" pageId="25" pageNumber="44" type="nomenclature">
<paragraph id="18093682FF8D7478FEFFFF3AFC7BFEF8" blockId="25.[113,1209,215,341]" pageId="25" pageNumber="44">
CLÉ D’ IDENTIFICATION DES «VERS BLANCS » (
<taxonomicName id="DFB64D01FF8D7478FC99FF3AFC4CFF18" ID-CoL="SC" ID-ENA="75546" authorityName="Latreille" authorityYear="1802" box="[812,984,215,242]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="20" phylum="Arthropoda" rank="superFamily" superFamily="Scarabaeoidea">SCARABAEOIDEA</taxonomicName>
) PRÉSENTS EN RIZICULTURE PLUVIALE D’ ALTITUDE À MADAGASCAR
</paragraph>
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<subSubSection id="50AC6509FF8D7478FFC4FECFFDE4FEBD" pageId="25" pageNumber="44" type="description">
<paragraph id="18093682FF8D7478FFC4FECFFDE4FEBD" blockId="25.[113,1209,215,341]" pageId="25" pageNumber="44">Le nombre de dents stridulatoires maxillaires, comme le nombre d’épines des palidia, sont mentionnés à titre indicatif, car ils sont sujets à une certaine variabilité.</paragraph>
</subSubSection>
<subSubSection id="50AC6509FF8D747AFF08FE82FBFAFAA0" lastPageId="27" lastPageNumber="46" pageId="25" pageNumber="44" type="key">
<key id="6C2761B3FF8D747AFF08FE82FBFAFAA0" lastPageId="27" lastPageNumber="46" pageId="25" pageNumber="44">
<keyStep id="A3422E27FF8D7478FF08FE82FBFAFE12" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08FE82FBF9FE29" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FE82FBF9FE29" pageId="25" pageNumber="44">
1. Maxilles à galea et lacinia distinctement séparées (
<figureCitation id="808D2A07FF8D7478FD50FE82FCD3FE6F" box="[741,839,367,391]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="25" pageNumber="44">Fig. 17H</figureCitation>
). Fente anale en « Y ». Palidia formant un cercle ouvert vers l’arrière d’épines convergeant vers la fente anale (
<figureCitation id="808D2A07FF8D7478FBA0FE61FBF2FE4C" box="[1045,1126,396,420]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="25" pageNumber="44">Fig. 5G</figureCitation>
) ...................................................................................
<taxonomicName id="DFB64D01FF8D7478FC96FE44FC00FE29" authorityName="Erichson" authorityYear="1847" box="[803,916,425,449]" class="Insecta" family="Orphnidae" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="family">Orphnidae</taxonomicName>
:
<taxonomicName id="DFB64D01FF8D7478FC10FE44FBF9FE29" baseAuthorityName="Guerin" baseAuthorityYear="1844" box="[933,1133,425,449]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="2AC2EA90FF8D7478FC10FE44FBF9FE29" box="[933,1133,425,449]" italics="true" pageId="25" pageNumber="44">Triodontus nitidulus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8D7478FF08FE2BFBFAFE12" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FE2BFBFAFE12" pageId="25" pageNumber="44">— Maxilles à galea et lacinia au moins partiellement soudées ou étroitement accolées l’une à l’autre. Fente anale transverse ou en « Y » .................................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8D7478FF08FDFBFBFAFD49" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08FDFBFBFAFD6C" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FDFBFBFAFD6C" pageId="25" pageNumber="44">
2. Fente anale transverse, droite ou faiblement arquée. Mandibules à aire stridulatoire ventrale formant une plage ovale, avec un nombre variable de stries transversales (
<figureCitation id="808D2A07FF8D7478FC0AFDDFFB87FDA2" box="[959,1043,562,586]" captionStart="FIG" captionStartId="40.[113,124,1612,1629]" captionTargetBox="[116,1208,217,1563]" captionTargetId="figure-21@40.[113,1202,214,1628]" captionTargetPageId="40" captionText="FIG. 13. — Larves de Scarabaeoidea de Madagascar: A-E, labre; A, Anochilia bifida (Olivier, 1789); B, Heteronychus plebeius (Klug, 1833); C, Paramorphochelus cornutus (Nonfried, 1892); D, Enaria melanictera (Klug, 1833); E, Triodontus nitidulus (Guérin, 1844); F-J, Celidota parvula (Janson, 1881), mandibule; F, gauche, face dorsale; G, gauche, face ventrale; H, droite, face dorsale; I, droite, face ventrale; J, aire stridulatoire mandibulaire. Échelles: A, B, D, F-I, 1 mm; C, E, J, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520793" httpUri="https://zenodo.org/record/4520793/files/figure.png" pageId="25" pageNumber="44">Fig. 13J</figureCitation>
), réduite à 5 stries difficilement visibles chez
<taxonomicName id="DFB64D01FF8D7478FDEAFDBDFCD5FD8F" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[607,833,591,615]" class="Insecta" family="Dynastidae" genus="Heteroconus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="2AC2EA90FF8D7478FDEAFDBDFCD5FD8F" box="[607,833,591,615]" italics="true" pageId="25" pageNumber="44">Heteroconus paradoxus</emphasis>
</taxonomicName>
. Galea et lacinia soudées sur toute leur longueur, la suture plus ou moins visible ou effacée .................................... 3
</keyLead>
</paragraph>
<paragraph id="18093682FF8D7478FF08FD64FBFAFD49" blockId="25.[189,1136,367,1699]" box="[189,1134,648,673]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FD64FBFAFD49" box="[189,1134,648,673]" pageId="25" pageNumber="44">— Fente anale en « Y ». Mandibules sans aire stridulatoire ventrale .................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8D7478FF08FD51FBFAFC52" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08FD51FBFAFC8C" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FD51FBFAFC8C" pageId="25" pageNumber="44">
3. Palidia en général présents, parfois absents. Lacinia avec un uncus formé de 2 dents sclérifiées. Dents stridulatoires maxillaires constituées de fortes pointes sclérifiées subconiques et pointues. Labre bilobé ou trilobé, en général presque symétrique, avec une tache pigmentée de part et d’autre du milieu, sur le bord antérieur (clithra) (
<figureCitation id="808D2A07FF8D7478FC8FFCFFFC03FCC2" box="[826,919,786,810]" captionStart="FIG" captionStartId="40.[113,124,1612,1629]" captionTargetBox="[116,1208,217,1563]" captionTargetId="figure-21@40.[113,1202,214,1628]" captionTargetPageId="40" captionText="FIG. 13. — Larves de Scarabaeoidea de Madagascar: A-E, labre; A, Anochilia bifida (Olivier, 1789); B, Heteronychus plebeius (Klug, 1833); C, Paramorphochelus cornutus (Nonfried, 1892); D, Enaria melanictera (Klug, 1833); E, Triodontus nitidulus (Guérin, 1844); F-J, Celidota parvula (Janson, 1881), mandibule; F, gauche, face dorsale; G, gauche, face ventrale; H, droite, face dorsale; I, droite, face ventrale; J, aire stridulatoire mandibulaire. Échelles: A, B, D, F-I, 1 mm; C, E, J, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520793" httpUri="https://zenodo.org/record/4520793/files/figure.png" pageId="25" pageNumber="44">Fig. 13A</figureCitation>
). Haptomère avec en général une rangée transverse arquée de courtes et fortes soies épineuses, jamais conformé en pointe saillante (
<figureCitation id="808D2A07FF8D7478FE06FCA1FDB3FC8B" box="[435,551,843,868]" captionStart="FIG" captionStartId="33.[113,124,1635,1652]" captionTargetBox="[117,1206,214,1443]" captionTargetId="figure-14@33.[117,1206,214,1443]" captionTargetPageId="33" captionText="FIG. 6. — Larves de Scarabaeoidea de Madagascar: A-F, tête; A, Anochilia bifida (Olivier, 1789); B, Hexodon unicolor (Olivier, 1789); C, sp.D2; D, Paramorphochelus cornutus (Nonfried,1892); E, Empecta scutata (Fairmaire,1901); F, sp.S2; G, H, épipharynx; G, Bricoptis variolosa (Gory &amp; Percheron, 1833); H, Celidota parvula (Janson, 1881). Échelles: A-C, 2 mm; D-F, 1 mm; G, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520767" httpUri="https://zenodo.org/record/4520767/files/figure.png" pageId="25" pageNumber="44">Figs 6G, H</figureCitation>
;
<figureCitation id="808D2A07FF8D7478FD8CFCA1FDE3FC8C" box="[569,631,844,868]" captionStart="FIG" captionStartId="34.[113,124,1635,1652]" captionTargetBox="[155,1168,220,1591]" captionTargetId="figure-21@34.[155,1168,220,1591]" captionTargetPageId="34" captionText="FIG. 7. — Larves de Scarabaeoidea de Madagascar,épipharynx: A, Anochilia bifida (Olivier,1789); B, Euryomia argentea (Olivier,1789); C, Heteronychus arator rugifrons (Fairmaire, 1871); D, Heteronychus bituberculatus (Kolbe, 1900); E, F, sp. D1; G, Hexodon unicolor (Olivier, 1789); H, sp. D2. Échelles: A, B, D, E, G, 0,5 mm; C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520771" httpUri="https://zenodo.org/record/4520771/files/figure.png" pageId="25" pageNumber="44">7A, B</figureCitation>
). Larves se déplaçant sur le dos ....
<taxonomicName id="DFB64D01FF8D7478FC57FCA1FBC3FC8C" authorityName="Leach" authorityYear="1815" box="[994,1111,844,868]" class="Insecta" family="Cetoniidae" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="family">Cetoniidae</taxonomicName>
. 4
</keyLead>
</paragraph>
<paragraph id="18093682FF8D7478FF08FC84FBFAFC52" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FC84FBFAFC52" pageId="25" pageNumber="44">
— Palidia toujours absents. Lacinia avec uncus formé de 2 ou 3, parfois 4 dents sclérifiées. Labre asymétrique (
<figureCitation id="808D2A07FF8D7478FE00FC68FD85FC75" box="[437,529,901,925]" captionStart="FIG" captionStartId="40.[113,124,1612,1629]" captionTargetBox="[116,1208,217,1563]" captionTargetId="figure-21@40.[113,1202,214,1628]" captionTargetPageId="40" captionText="FIG. 13. — Larves de Scarabaeoidea de Madagascar: A-E, labre; A, Anochilia bifida (Olivier, 1789); B, Heteronychus plebeius (Klug, 1833); C, Paramorphochelus cornutus (Nonfried, 1892); D, Enaria melanictera (Klug, 1833); E, Triodontus nitidulus (Guérin, 1844); F-J, Celidota parvula (Janson, 1881), mandibule; F, gauche, face dorsale; G, gauche, face ventrale; H, droite, face dorsale; I, droite, face ventrale; J, aire stridulatoire mandibulaire. Échelles: A, B, D, F-I, 1 mm; C, E, J, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520793" httpUri="https://zenodo.org/record/4520793/files/figure.png" pageId="25" pageNumber="44">Fig. 13B</figureCitation>
). Haptomère en pointe saillante (
<figureCitation id="808D2A07FF8D7478FCDDFC68FC32FC75" box="[872,934,901,925]" captionStart="FIG" captionStartId="34.[113,124,1635,1652]" captionTargetBox="[155,1168,220,1591]" captionTargetId="figure-21@34.[155,1168,220,1591]" captionTargetPageId="34" captionText="FIG. 7. — Larves de Scarabaeoidea de Madagascar,épipharynx: A, Anochilia bifida (Olivier,1789); B, Euryomia argentea (Olivier,1789); C, Heteronychus arator rugifrons (Fairmaire, 1871); D, Heteronychus bituberculatus (Kolbe, 1900); E, F, sp. D1; G, Hexodon unicolor (Olivier, 1789); H, sp. D2. Échelles: A, B, D, E, G, 0,5 mm; C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520771" httpUri="https://zenodo.org/record/4520771/files/figure.png" pageId="25" pageNumber="44">Figs 7</figureCitation>
C-H; 8A, B; 11A). Larves se déplaçant sur le ventre ...........................................................
<taxonomicName id="DFB64D01FF8D7478FC67FC4FFBDDFC52" authorityName="MacLeay" authorityYear="1819" box="[978,1097,930,954]" class="Insecta" family="Dynastidae" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="family">Dynastidae</taxonomicName>
. 18
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8D7478FF08FC38FBFAFBE2" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08FC38FBFAFC05" blockId="25.[189,1136,367,1699]" box="[189,1134,981,1005]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FC38FBFAFC05" box="[189,1134,981,1005]" pageId="25" pageNumber="44">
4. Raster sans palidia (
<figureCitation id="808D2A07FF8D7478FE00FC38FD90FC05" box="[437,516,981,1005]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3C</figureCitation>
) ............................................................................... sp. C1
</keyLead>
</paragraph>
<paragraph id="18093682FF8D7478FF08FC1FFBFAFBE2" blockId="25.[189,1136,367,1699]" box="[189,1134,1010,1034]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FC1FFBFAFBE2" box="[189,1134,1010,1034]" pageId="25" pageNumber="44">
— Raster avec palidia (
<figureCitation id="808D2A07FF8D7478FE03FC1FFDF5FBE2" box="[438,609,1010,1034]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3A, B, D, E</figureCitation>
) ......................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8D7478FF08FBC8FBF8FB58" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08FBC8FBF8FBB2" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FBC8FBF8FBB2" pageId="25" pageNumber="44">
5. Palidia (
<figureCitation id="808D2A07FF8D7478FE8FFBC8FE10FBD5" box="[314,388,1061,1085]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3A</figureCitation>
) formés par 2 rangées longitudinales de 12 à 19 épines légèrement concaves. 5 + 1 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8D7478FDDDFBAFFD56FBB2" box="[616,706,1090,1114]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="25" pageNumber="44">Fig. 16E</figureCitation>
) .................................
<taxonomicName id="DFB64D01FF8D7478FC0CFBAEFBF8FBB2" baseAuthorityName="Gory &amp; Percheron" baseAuthorityYear="1833" box="[953,1132,1090,1114]" class="Insecta" family="Cetoniidae" genus="Bricoptis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="species" species="variolosa">
<emphasis id="2AC2EA90FF8D7478FC0CFBAEFBF8FBB2" box="[953,1132,1090,1114]" italics="true" pageId="25" pageNumber="44">Bricoptis variolosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8D7478FF08FBB2FBF8FB58" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FBB2FBF8FB58" pageId="25" pageNumber="44">
— Palidia conformés autrement, soit avec 2 rangées d’épines divergentes vers l’arrière (
<figureCitation id="808D2A07FF8D7478FBA2FBB2FBF6FB9F" box="[1047,1122,1119,1143]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3B</figureCitation>
), soit avec 2 rangées d’épines subparallèles (
<figureCitation id="808D2A07FF8D7478FD32FB96FD4CFB7C" box="[647,728,1147,1172]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3D</figureCitation>
), soit avec 2 rangées d’épines légèrement convexes (
<figureCitation id="808D2A07FF8D7478FEE4FB75FE09FB58" box="[337,413,1176,1200]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3E</figureCitation>
) ...................................................................................................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8D7478FF08FB26FBFBFA67" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08FB26FBFAFAD2" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FB26FBFAFAD2" pageId="25" pageNumber="44">
6. Palidia (
<figureCitation id="808D2A07FF8D7478FEF4FB21FE05FB0C" box="[321,401,1228,1252]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3B</figureCitation>
) formés par 2 rangées longitudinales de 16 à 21 épines divergeant à l’extrémité postérieure, avec une rupture de la courbe au tiers basal environ (formant ainsi le dessin d’un « fer de lance »). 5 + 1 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8D7478FBBEFAE8FBF3FAF5" box="[1035,1127,1285,1309]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="25" pageNumber="44">Fig. 16F</figureCitation>
) ............................................................................................................
<taxonomicName id="DFB64D01FF8D7478FC7DFACFFBFAFAD2" baseAuthorityName="Janson" baseAuthorityYear="1881" box="[968,1134,1314,1338]" class="Insecta" family="Cetoniidae" genus="Celidota" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="2AC2EA90FF8D7478FC7DFACFFBFAFAD2" box="[968,1134,1314,1338]" italics="true" pageId="25" pageNumber="44">Celidota parvula</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8D7478FF08FAD2FBFBFA67" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FAD2FBFBFA67" pageId="25" pageNumber="44">
— Palidia formés d’alignements de 11 à 16 épines disposées soit en 2 rangées subparallèles (
<figureCitation id="808D2A07FF8D7478FF47FAB6FED3FA9B" box="[242,327,1371,1395]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3D</figureCitation>
), soit en 2 rangées légèrement convexes dessinant une « voûte gothique » vers l’avant (
<figureCitation id="808D2A07FF8D7478FE8BFA95FE1EFA78" box="[318,394,1400,1424]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3E</figureCitation>
) .......................................................................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8D7478FF08FA41FBFAF9DE" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08FA41FBFAFA15" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08FA41FBFAFA15" pageId="25" pageNumber="44">
7. Palidia (
<figureCitation id="808D2A07FF8D7478FE89FA46FE13FA2B" box="[316,391,1451,1475]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3E</figureCitation>
) formés par deux rangées légèrement convexes de 13 à 16 épines, formant une pointe antérieure en « ogive gothique ». 4 + 1 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8D7478FBBDFA25FBF0FA08" box="[1032,1124,1480,1504]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="25" pageNumber="44">Fig. 16H</figureCitation>
) ........................................................................................................
<taxonomicName id="DFB64D01FF8D7478FC01FA08FBFAFA15" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[948,1134,1509,1533]" class="Insecta" family="Cetoniidae" genus="Euryomia" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="species" species="argentea">
<emphasis id="2AC2EA90FF8D7478FC01FA08FBFAFA15" box="[948,1134,1509,1533]" italics="true" pageId="25" pageNumber="44">Euryomia argentea</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8D7478FF08F9EFFBFAF9DE" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08F9EFFBFAF9DE" pageId="25" pageNumber="44">
— Palidia (
<figureCitation id="808D2A07FF8D7478FE8AF9ECFE05F9F2" box="[319,401,1537,1562]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="25" pageNumber="44">Fig. 3D</figureCitation>
) formés par 2 rangées subparallèles de 11 à 14 épines. 6 à 8 + 1 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8D7478FE44F9F3FDDBF9DE" box="[497,591,1566,1590]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="25" pageNumber="44">Fig. 16G</figureCitation>
) ......................................................
<taxonomicName id="DFB64D01FF8D7478FC7AF9F3FBFAF9DE" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[975,1134,1566,1590]" class="Insecta" family="Cetoniidae" genus="Anochilia" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="2AC2EA90FF8D7478FC7AF9F3FBFAF9DE" box="[975,1134,1566,1590]" italics="true" pageId="25" pageNumber="44">Anochilia bifida</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8D747BFF08F9BFFBFAFF18" lastPageId="26" lastPageNumber="45" pageId="25" pageNumber="44">
<paragraph id="18093682FF8D7478FF08F9BFFBFAF94B" blockId="25.[189,1136,367,1699]" pageId="25" pageNumber="44">
<keyLead id="A34795B7FF8D7478FF08F9BFFBFAF94B" pageId="25" pageNumber="44">
8. Palidia absents, raster avec un groupe de fortes soies aplaties et recourbées à l’apex du côté ventral de la fente anale, avec de nombreuses soies plus fines autour de la fente anale et du côté dorsal de celle-ci (
<figureCitation id="808D2A07FF8D7478FE40F966FDD4F94B" box="[501,576,1675,1699]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="25" pageNumber="44">Fig. 4F</figureCitation>
) ...................... Hopliidae:
<taxonomicName id="DFB64D01FF8D7478FCD7F966FBFAF94B" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[866,1134,1675,1699]" class="Insecta" family="Melolonthidae" genus="Paramorphochelus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="44" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="2AC2EA90FF8D7478FCD7F966FBFAF94B" box="[866,1134,1675,1699]" italics="true" pageId="25" pageNumber="44">Paramorphochelus cornutus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FF35FBFAFF18" blockId="26.[189,1136,216,1693]" box="[189,1134,216,240]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FF35FBFAFF18" box="[189,1134,216,240]" pageId="26" pageNumber="45">— Palidia présents ........................................................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FEE0FBFAFE39" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FEE0FBFAFE70" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FEE0FBFAFE70" pageId="26" pageNumber="45">
9. Palidia formés par 2 rangées ou groupes symétriques d’épines séparés par le septula glabre (
<figureCitation id="808D2A07FF8E747BFF45FEC4FEFDFEA9" box="[240,361,297,322]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="26" pageNumber="45">Figs 4G, H</figureCitation>
;
<figureCitation id="808D2A07FF8E747BFECEFEC4FE0FFEA9" box="[379,411,297,321]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">5A</figureCitation>
, C-F). Si le palidium est formé par une rangée d’épines disposées en arc de cercle, alors celles-ci sont regroupées en « touffes » de 2, 3 ou 4 soies rapprochées (
<figureCitation id="808D2A07FF8E747BFF47FE8EFED5FE93" box="[242,321,355,379]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5B</figureCitation>
). Galea et lacinia fusionnées à la base mais séparées et accolées à l’apex ............ ........................................................................................................
<taxonomicName id="DFB64D01FF8E747BFC1AFE6DFBDEFE70" authorityName="MacLeay" authorityYear="1819" box="[943,1098,384,408]" class="Insecta" family="Melolonthidae" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="family">Melolonthidae</taxonomicName>
. 10
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FE70FBFAFE39" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FE70FBFAFE39" pageId="26" pageNumber="45">
— Palidium formé par une rangée d’épines régulièrement disposées en arc de cercle ouvert vers l’arrière (
<figureCitation id="808D2A07FF8E747BFEC1FE54FE48FE39" box="[372,476,441,465]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5H, I</figureCitation>
) ........................................................................... Sericidae. 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FE03FBFAFD27" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FE03FBE4FDB4" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FE03FBE4FDB4" pageId="26" pageNumber="45">
10. Soit palidium formé par une rangée de fortes soies disposées en arc de cercle et regroupées en « touffes » de 2, 3 ou 4 soies rapprochées (
<figureCitation id="808D2A07FF8E747BFD1AFDE6FD6AFDCB" box="[687,766,523,547]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5B</figureCitation>
), soit palidia formés par une double rangée symétrique de soies dessinant un V renversé (
<figureCitation id="808D2A07FF8E747BFCADFDCAFCFDFDD7" box="[792,873,551,575]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5A</figureCitation>
) ou un fuseau (
<figureCitation id="808D2A07FF8E747BFBA6FDCAFBF2FDD7" box="[1043,1126,551,575]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5C</figureCitation>
) ................................................................................................................................. 15
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FD8CFBFAFD27" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FD8CFBFAFD27" pageId="26" pageNumber="45">
— Palidia formés non par une double rangée symétrique de soies mais soit par un double groupe symétrique de soies (
<figureCitation id="808D2A07FF8E747BFDB3FD93FDECFD7D" box="[518,632,638,662]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="26" pageNumber="45">Figs 4G, H</figureCitation>
;
<figureCitation id="808D2A07FF8E747BFD32FD90FD55FD7E" box="[647,705,637,662]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">5E, F</figureCitation>
), soit par une double rangée symétrique de courtes épines en avant et de 2 groupes symétriques de fortes et longues soies en arrière, l’ensemble formant un «T » inversé (
<figureCitation id="808D2A07FF8E747BFDD5FD5AFD25FD27" box="[608,689,695,719]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5D</figureCitation>
) ........................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FD01FBFAFCB2" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FD01FBFAFCD5" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FD01FBFAFCD5" pageId="26" pageNumber="45">
11. Palidia (
<figureCitation id="808D2A07FF8E747BFE8AFD01FE05FCEC" box="[319,401,748,772]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5D</figureCitation>
) formés par une double rangée symétrique de courtes épines en avant et par 2 groupes symétriques de fortes et longues soies en arrière, l’ensemble formant un « T» inversé. 13 à 14 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8E747BFCBDFCC8FCF5FCD6" box="[776,865,805,830]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="26" pageNumber="45">Fig. 17F</figureCitation>
) .............
<taxonomicName id="DFB64D01FF8E747BFC65FCCBFBFAFCD5" baseAuthorityName="Fairmaire" baseAuthorityYear="1901" box="[976,1134,806,829]" class="Insecta" family="Melolonthidae" genus="Empecta" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="scutata">
<emphasis id="2AC2EA90FF8E747BFC65FCCBFBFAFCD5" box="[976,1134,806,829]" italics="true" pageId="26" pageNumber="45">Empecta scutata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FCAEFBFAFCB2" blockId="26.[189,1136,216,1693]" box="[189,1134,834,859]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FCAEFBFAFCB2" box="[189,1134,834,859]" pageId="26" pageNumber="45">
— Palidia formés par 2 groupes symétriques de fortes soies (
<figureCitation id="808D2A07FF8E747BFC84FCAFFC32FCB2" box="[817,934,834,858]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="26" pageNumber="45">Figs 4G, H</figureCitation>
;
<figureCitation id="808D2A07FF8E747BFC02FCAFFC64FCB2" box="[951,1008,834,858]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">5E, F</figureCitation>
) ........... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FC9AFBFAFC0D" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FC9AFBFBFC44" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FC9AFBFBFC44" pageId="26" pageNumber="45">
12. Palidia (
<figureCitation id="808D2A07FF8E747BFEF7FC9AFE0CFC67" box="[322,408,887,911]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="26" pageNumber="45">Fig. 4G</figureCitation>
) formés par 2 groupes symétriques de fortes soies en forme de « J » .......................................................................................................................... sp. M1
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FC5CFBFAFC0D" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FC5CFBFAFC0D" pageId="26" pageNumber="45">
— Palidia (
<figureCitation id="808D2A07FF8E747BFE8EFC5DFE1AFC21" box="[315,398,944,969]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="26" pageNumber="45">Figs 4H</figureCitation>
;
<figureCitation id="808D2A07FF8E747BFE28FC5DFE41FC21" box="[413,469,944,969]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">5E, F</figureCitation>
) formés par 2 groupes subtriangulaires symétriques de fortes soies ................................................................................................................................. 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FBEFFBFAFB98" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FBEFFBE4FBDF" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FBEFFBE4FBDF" pageId="26" pageNumber="45">
13. Palidia (
<figureCitation id="808D2A07FF8E747BFEF7FBEFFE06FBF2" box="[322,402,1026,1050]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5F</figureCitation>
) formés par 2 groupes de fortes soies disposées sur 2 ou 3 rangs au maximum .......................................................................................................... sp. M3
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FBD1FBFAFB98" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FBD1FBFAFB98" pageId="26" pageNumber="45">
— Palidia (
<figureCitation id="808D2A07FF8E747BFE89FBD1FE04FBBC" box="[316,400,1084,1108]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="26" pageNumber="45">Figs 4H</figureCitation>
;
<figureCitation id="808D2A07FF8E747BFE15FBD6FE2AFBBB" box="[416,446,1083,1107]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">5E</figureCitation>
) formés par 2 groupes de fortes soies plus nombreuses disposées sur 4 rangs ou plus ......................................................................................................... 14
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FB60FBFAFAF0" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FB60FBFAFB2A" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FB60FBFAFB2A" pageId="26" pageNumber="45">
14. Palidia (
<figureCitation id="808D2A07FF8E747BFE88FB60FE1AFB4D" box="[317,398,1165,1189]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="26" pageNumber="45">Fig. 4H</figureCitation>
) en triangle plus large que haut. Teges plus étroits, aux soies plus courtes disposées sur 3 rangs. 17 à 21 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8E747BFCEEFB47FC23FB2A" box="[859,951,1194,1218]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="26" pageNumber="45">Fig. 17C</figureCitation>
) ....
<taxonomicName id="DFB64D01FF8E747BFC5BFB46FBFAFB2A" baseAuthorityName="Brenske" baseAuthorityYear="1891" box="[1006,1134,1194,1218]" class="Insecta" family="Melolonthidae" genus="Encya" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="sikorai">
<emphasis id="2AC2EA90FF8E747BFC5BFB46FBFAFB2A" box="[1006,1134,1194,1218]" italics="true" pageId="26" pageNumber="45">Encya sikorai</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FB2AFBFAFAF0" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FB2AFBFAFAF0" pageId="26" pageNumber="45">
— Palidia (
<figureCitation id="808D2A07FF8E747BFEF4FB2AFE04FB37" box="[321,400,1223,1247]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5E</figureCitation>
) en triangle plus haut que large, en « ogive gothique » dont la pointe serait tronquée. Teges plus large, jusqu’à 5 rangs de soies plus allongées. 15 à 16 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8E747BFE44FAEDFDDAFAF0" box="[497,590,1280,1304]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="26" pageNumber="45">Fig. 17G</figureCitation>
) ..........................................
<taxonomicName id="DFB64D01FF8E747BFCCBFAEDFBFAFAF0" baseAuthorityName="Kunckel" baseAuthorityYear="1887" box="[894,1134,1280,1304]" class="Insecta" family="Melolonthidae" genus="Hoplochelus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="betanimena">
<emphasis id="2AC2EA90FF8E747BFCCBFAEDFBFAFAF0" box="[894,1134,1280,1304]" italics="true" pageId="26" pageNumber="45">Hoplochelus betanimena</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FAD8FBFAFA6E" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FAD8FBE4FA82" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FAD8FBE4FA82" pageId="26" pageNumber="45">
15. Palidia (
<figureCitation id="808D2A07FF8E747BFEF7FAD8FE00FAA5" box="[322,404,1333,1357]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5A</figureCitation>
) en forme de « V » renversé. 11 à 13 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8E747BFF41FABFFEC6FA82" box="[244,338,1362,1386]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="26" pageNumber="45">Fig. 17D</figureCitation>
) .......................................................................................
<taxonomicName id="DFB64D01FF8E747BFC06FABEFBE4FA82" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[947,1136,1362,1386]" class="Insecta" family="Melolonthidae" genus="Enaria" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="melanictera">
<emphasis id="2AC2EA90FF8E747BFC06FABEFBE4FA82" box="[947,1136,1362,1386]" italics="true" pageId="26" pageNumber="45">Enaria melanictera</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FA82FBFAFA6E" blockId="26.[189,1136,216,1693]" box="[189,1134,1390,1415]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FA82FBFAFA6E" box="[189,1134,1390,1415]" pageId="26" pageNumber="45">
— Palidia (
<figureCitation id="808D2A07FF8E747BFE88FA83FE3EFA6F" box="[317,426,1390,1415]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5B, C</figureCitation>
) arrondis ..................................................................................... 16
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08FA4EFBFAF9FA" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08FA4EFBE4FA30" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FA4EFBE4FA30" pageId="26" pageNumber="45">
16. Palidium (
<figureCitation id="808D2A07FF8E747BFEECFA4EFE3DFA53" box="[345,425,1443,1467]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5B</figureCitation>
) formé par une rangée de fortes soies disposées en arc de cercle et regroupées en « touffes » de 2, 3 ou 4 soies rapprochées ...................................... sp. M2
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08FA30FBFAF9FA" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08FA30FBFAF9FA" pageId="26" pageNumber="45">
— Palidia (
<figureCitation id="808D2A07FF8E747BFE89FA30FE1DFA1D" box="[316,393,1501,1525]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5C</figureCitation>
) formés par une double rangée symétrique de 13-15 fortes soies dessinant un fuseau. 16 à 17 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8E747BFD46FA17FCD9F9FA" box="[755,845,1530,1554]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="26" pageNumber="45">Fig. 17E</figureCitation>
) ...........
<taxonomicName id="DFB64D01FF8E747BFC1AFA17FBFAF9FA" baseAuthorityName="Dewailly" baseAuthorityYear="1950" box="[943,1134,1530,1554]" class="Insecta" family="Melolonthidae" genus="Apicencya" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="45" phylum="Arthropoda" rank="species" species="waterloti">
<emphasis id="2AC2EA90FF8E747BFC1AFA17FBFAF9FA" box="[943,1134,1530,1554]" italics="true" pageId="26" pageNumber="45">Apicencya waterloti</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8E747BFF08F9C2FBFAF975" pageId="26" pageNumber="45">
<paragraph id="18093682FF8E747BFF08F9C2FBFAF98B" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08F9C2FBFAF98B" pageId="26" pageNumber="45">
17. Palidia (
<figureCitation id="808D2A07FF8E747BFEF1F9C3FE08F9AE" box="[324,412,1582,1606]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5H</figureCitation>
) formés par 15 à 18 longues épines aplaties très serrées, pointues ........................................................................................................................... sp. S1
</keyLead>
</paragraph>
<paragraph id="18093682FF8E747BFF08F985FBFAF975" blockId="26.[189,1136,216,1693]" pageId="26" pageNumber="45">
<keyLead id="A34795B7FF8E747BFF08F985FBFAF975" pageId="26" pageNumber="45">
— Palidia (
<figureCitation id="808D2A07FF8E747BFE88F985FE17F968" box="[317,387,1640,1664]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="26" pageNumber="45">Fig. 5I</figureCitation>
) formés par 17 à 19 courtes épines aplaties, plus espacées, à apex arrondi ........................................................................................................................... sp.
<geoCoordinate id="7D825045FF8E747BFBE6F968FBFAF975" box="[1107,1134,1669,1693]" degrees="2" direction="south" minutes="18" orientation="latitude" pageId="26" pageNumber="45" precision="925" value="-2.3">S2</geoCoordinate>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8F747AFF08FF35FBFAFE8B" pageId="27" pageNumber="46">
<paragraph id="18093682FF8F747AFF08FF35FBFAFEAE" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FF35FBFAFEAE" pageId="27" pageNumber="46">
18. Raster (
<figureCitation id="808D2A07FF8F747AFE83FF35FE15FF18" box="[310,385,216,240]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="27" pageNumber="46">Fig. 4E</figureCitation>
) avec symétriquement de part et d’autre du milieu, séparées par une étroite zone glabre, une très forte et épaisse épine variable orientée vers l’intérieur (simple, bifide ou trifide) et dissymétrique (trifide ou bifide du côté gauche et bifide ou simple du côté droit). 9 à 10 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8F747AFD08FEC3FC8CFEAE" box="[701,792,302,326]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="27" pageNumber="46">Fig. 17A</figureCitation>
) .................
<taxonomicName id="DFB64D01FF8F747AFC2AFEC3FBFAFEAE" baseAuthorityName="Fairmaire" baseAuthorityYear="1871" box="[927,1134,302,326]" class="Insecta" family="Dynastidae" genus="Paranodon" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="coquerelii">
<emphasis id="2AC2EA90FF8F747AFC2AFEC3FBFAFEAE" box="[927,1134,302,326]" italics="true" pageId="27" pageNumber="46">Paranodon coquerelii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8F747AFF08FEA6FBFAFE8B" blockId="27.[189,1136,216,1352]" box="[189,1134,331,355]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FEA6FBFAFE8B" box="[189,1134,331,355]" pageId="27" pageNumber="46">— Raster sans fortes soies bifides ou trifides .................................................................. 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8F747AFF08FE93FBFAFE04" pageId="27" pageNumber="46">
<paragraph id="18093682FF8F747AFF08FE93FBFAFE38" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FE93FBFAFE38" pageId="27" pageNumber="46">
19. Raster (
<figureCitation id="808D2A07FF8F747AFE8DFE93FE15FE7E" box="[312,385,382,406]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="27" pageNumber="46">Fig. 3F</figureCitation>
) avec une large zone médiane glabre centrale bordée d’une rangée de 5 ou 6 courtes épines du côté dorsal de la fente anale. 7 à 10 + 1 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8F747AFF47FE55FED1FE27" box="[242,325,439,464]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="27" pageNumber="46">Fig. 16J</figureCitation>
) ..........................................................................
<taxonomicName id="DFB64D01FF8F747AFCF9FE55FBFAFE38" baseAuthorityName="Fairmaire" baseAuthorityYear="1871" box="[844,1134,440,464]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="arator" subSpecies="rugifrons">
<emphasis id="2AC2EA90FF8F747AFCF9FE55FBFAFE38" box="[844,1134,440,464]" italics="true" pageId="27" pageNumber="46">Heteronychus arator rugifrons</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8F747AFF08FE38FBFAFE04" blockId="27.[189,1136,216,1352]" box="[189,1134,468,492]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FE38FBFAFE04" box="[189,1134,468,492]" pageId="27" pageNumber="46">— Raster sans large zone médiane glabre centrale du côté dorsal de la fente anale ......... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8F747AFF08FDE5FBE4FD9E" pageId="27" pageNumber="46">
<paragraph id="18093682FF8F747AFF08FDE5FBF9FDD5" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FDE5FBF9FDD5" pageId="27" pageNumber="46">
20. Raster (
<figureCitation id="808D2A07FF8F747AFE83FDE5FE17FDC8" box="[310,387,520,544]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="27" pageNumber="46">Fig. 4C</figureCitation>
): teges à soies épineuses courtes non recourbées à l’apex. Larve L3 toujours de très grande taille ............................................................................................ sp. D2
</keyLead>
</paragraph>
<paragraph id="18093682FF8F747AFF08FDAFFBE4FD9E" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FDAFFBE4FD9E" pageId="27" pageNumber="46">— Raster: teges à soies épineuses aplaties recourbées à l’apex. Larve L3 de taille petite à grande ...................................................................................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8F747AFF08FD7CFBFAFCF4" pageId="27" pageNumber="46">
<paragraph id="18093682FF8F747AFF08FD7CFBF8FD2E" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FD7CFBF8FD2E" pageId="27" pageNumber="46">
21. Raster (
<figureCitation id="808D2A07FF8F747AFE83FD7CFE16FD41" box="[310,386,657,681]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="27" pageNumber="46">Fig. 4A</figureCitation>
): teges avec des épines aplaties et recourbées assez étroites, très nombreuses et avec de longues soies sur la périphérie. Larve L3 toujours de grande taille ...... sp. D1
</keyLead>
</paragraph>
<paragraph id="18093682FF8F747AFF08FD26FBFAFCF4" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FD26FBFAFCF4" pageId="27" pageNumber="46">
— Raster (
<figureCitation id="808D2A07FF8F747AFE8DFD26FE3AFD0A" box="[312,430,715,739]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="27" pageNumber="46">Figs 3G, H</figureCitation>
;
<figureCitation id="808D2A07FF8F747AFE75FD26FD94FD0B" box="[448,512,715,739]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="27" pageNumber="46">4B, D</figureCitation>
): teges jamais entourés par de longues soies, avec des épines aplaties et recourbées assez larges, moins nombreuses. Larve L3 de taille petite à grande ................................................................................................................................. 22
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8F747AFF08FCD5FBFAFC37" pageId="27" pageNumber="46">
<paragraph id="18093682FF8F747AFF08FCD5FBFAFC4E" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FCD5FBFAFC4E" pageId="27" pageNumber="46">
22. Raster (
<figureCitation id="808D2A07FF8F747AFE8FFCD5FE1DFCB8" box="[314,393,824,848]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="27" pageNumber="46">Fig. 4B</figureCitation>
): teges avec de nombreuses soies aplaties et recourbées. Du côté dorsal de la fente anale, nombreuses soies courtes mêlées de quelques longues soies. Une étroite bande glabre le long de la fente anale du côté dorsal. 9 à 10 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8F747AFF47FC63FEC7FC4E" box="[242,339,910,934]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="27" pageNumber="46">Fig. 16N</figureCitation>
). Larves L3 de grande taille ...............................................
<taxonomicName id="DFB64D01FF8F747AFC0AFC63FBFAFC4E" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[959,1134,910,934]" class="Insecta" family="Dynastidae" genus="Hexodon" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="2AC2EA90FF8F747AFC0AFC63FBFAFC4E" box="[959,1134,910,934]" italics="true" pageId="27" pageNumber="46">Hexodon unicolor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8F747AFF08FC46FBFAFC37" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FC46FBFAFC37" pageId="27" pageNumber="46">
— Raster conformé autrement (
<figureCitation id="808D2A07FF8F747AFDA4FC47FD12FC2A" box="[529,646,938,963]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="27" pageNumber="46">Figs 3G, H</figureCitation>
;
<figureCitation id="808D2A07FF8F747AFD22FC46FD28FC2B" box="[663,700,939,963]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="27" pageNumber="46">4D</figureCitation>
), teges à épines moins nombreuses. Larves L3 de petite taille ..................................................................................................... 23
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8F747AFF08FC16FBFAFB6E" pageId="27" pageNumber="46">
<paragraph id="18093682FF8F747AFF08FC16FBFAFB81" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FC16FBFAFB81" pageId="27" pageNumber="46">
23. Raster (
<figureCitation id="808D2A07FF8F747AFE8DFC16FE1DFBFB" box="[312,393,1019,1043]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="27" pageNumber="46">Fig. 4D</figureCitation>
): teges avec de très fortes soies dispersées, épaisses, recourbées et aplaties. Du côté dorsal de la fente anale, nombreuses soies courtes avec quelques très longues soies fines. Mandibules à aire stridulatoire réduite à 5 stries diffi cilement visibles. 12 à 15 dents stridulatoires maxillaires (
<figureCitation id="808D2A07FF8F747AFE44FBBCFDC4FB81" box="[497,592,1105,1129]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="27" pageNumber="46">Fig. 16O</figureCitation>
) ............................................
<taxonomicName id="DFB64D01FF8F747AFC3BFBBCFBFAFB81" baseAuthorityName="Endrodi" baseAuthorityYear="1968" box="[910,1134,1105,1129]" class="Insecta" family="Dynastidae" genus="Heteroconus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="2AC2EA90FF8F747AFC3BFBBCFBFAFB81" box="[910,1134,1105,1129]" italics="true" pageId="27" pageNumber="46">Heteroconus paradoxus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8F747AFF08FB83FBFAFB6E" blockId="27.[189,1136,216,1352]" box="[189,1134,1133,1158]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FB83FBFAFB6E" box="[189,1134,1133,1158]" pageId="27" pageNumber="46">
— Raster conformé autrement (
<figureCitation id="808D2A07FF8F747AFDA7FB80FD11FB6E" box="[530,645,1133,1158]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="27" pageNumber="46">Fig. 3G, H</figureCitation>
) .................................................................. 24
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A3422E27FF8F747AFF08FB4CFBFAFAA0" pageId="27" pageNumber="46">
<paragraph id="18093682FF8F747AFF08FB4CFBF9FB1A" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FB4CFBF9FB1A" pageId="27" pageNumber="46">
24. Raster (
<figureCitation id="808D2A07FF8F747AFE8CFB4CFE1EFB51" box="[313,394,1185,1209]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="27" pageNumber="46">Fig. 3G</figureCitation>
): teges avec de fortes et longues soies aplaties, à l’apex incurvé, avec de chaque côté un groupe de soies plus longues. 5 dents stridulatoires maxillaires .............. ...........................................................................................
<taxonomicName id="DFB64D01FF8F747AFCECFB37FBF9FB1A" baseAuthorityName="Kolbe" baseAuthorityYear="1900" box="[857,1133,1242,1266]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="bituberculatus">
<emphasis id="2AC2EA90FF8F747AFCECFB37FBF9FB1A" box="[857,1133,1242,1266]" italics="true" pageId="27" pageNumber="46">Heteronychus bituberculatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="18093682FF8F747AFF08FB15FBFAFAA0" blockId="27.[189,1136,216,1352]" pageId="27" pageNumber="46">
<keyLead id="A34795B7FF8F747AFF08FB15FBFAFAA0" pageId="27" pageNumber="46">
— Raster (
<figureCitation id="808D2A07FF8F747AFE8CFB1AFE1FFAE7" box="[313,395,1271,1295]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="27" pageNumber="46">Fig. 3H</figureCitation>
): teges avec des soies bien plus courtes que chez
<taxonomicName id="DFB64D01FF8F747AFC3DFB15FBA1FAE7" baseAuthorityName="Kolbe" baseAuthorityYear="1900" box="[904,1077,1271,1295]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="bituberculatus">
<emphasis id="2AC2EA90FF8F747AFC3DFB15FBA1FAE7" box="[904,1077,1271,1295]" italics="true" pageId="27" pageNumber="46">H. bituberculatus</emphasis>
</taxonomicName>
, sans groupe de longues soies de chaque côté. 7 + 1 dents stridulatoires maxillaires ................ .....................................................................................................
<taxonomicName id="DFB64D01FF8F747AFC2DFADDFBFAFAA0" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[920,1134,1328,1352]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="46" phylum="Arthropoda" rank="species" species="plebeius">
<emphasis id="2AC2EA90FF8F747AFC2DFADDFBFAFAA0" box="[920,1134,1328,1352]" italics="true" pageId="27" pageNumber="46">Heteronychus plebeius</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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