239 lines
24 KiB
XML
239 lines
24 KiB
XML
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<mods:title id="93C2A38473E430C2641EB2A5DB45ADB5">Conchological and molecular analysis of the " non-scaly " Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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<mods:namePart id="624F9E5D2D395B3247AA4CEB8FC51B52">Zacaery Khalik, Mohd</mods:namePart>
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<mods:namePart id="C7638906C0BB88C588608BCC21A7E14C">P. Hendriks, Kasper</mods:namePart>
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<mods:namePart id="8A01FDC7DFC4BC905EE5FD8CD340EE1B">J. Vermeulen, Jaap</mods:namePart>
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<mods:namePart id="97D1C32B04EB1A3869879CB7DD09B297">Menno Schilthuizen,</mods:namePart>
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<treatment id="8EDD22CAAA52E5066D0DB84E772991BD" ID-GBIF-Taxon="156787941" LSID="urn:lsid:plazi:treatment:8EDD22CAAA52E5066D0DB84E772991BD" httpUri="http://treatment.plazi.org/id/8EDD22CAAA52E5066D0DB84E772991BD" lastPageId="16" lastPageNumber="51" pageId="13" pageNumber="48">
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<subSubSection id="C82F9CF53270C6B0A25EFCDC6EBD30CE" pageId="13" pageNumber="48" type="nomenclature">
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<paragraph id="0B775E0F1C0F7FC4245B32A4DA97DF68" pageId="13" pageNumber="48">
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<taxonomicName id="8665489447F8DB455DE78239B4D2B6C6" ID-CoL="3FRYS" authority="Godwin-Austen, 1889" authorityName="Godwin-Austen" authorityYear="1889" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa williamsi" order="Cycloneritida" pageId="13" pageNumber="48" phylum="Mollusca" rank="species" species="williamsi">
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<pageBreakToken id="4E4239F092C038418F0C1FD5DFAF3E94" pageId="13" pageNumber="48" start="start">Georissa</pageBreakToken>
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williamsi Godwin-Austen, 1889
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</taxonomicName>
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Figures 1C, 8
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<normalizedToken id="DABDE41A43AFAAA18969606E7629E2F7" originalValue="A–I">A-I</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection id="20D38B95EBC0D75FBAA4FC737F40A0D2" pageId="14" pageNumber="49" type="reference_group">
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<paragraph id="C6672E237D646424AD2FCDB75915FF49" pageId="14" pageNumber="49">
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<taxonomicName id="97F3537B2EB0233908F672954E045A7B" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa williamsi" order="Cycloneritida" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="williamsi">
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<pageBreakToken id="051F09EFA4E161F8E6B1A99F94B6D674" pageId="14" pageNumber="49" start="start">Georissa</pageBreakToken>
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williamsi
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</taxonomicName>
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Godwin-Austen, 1889: 353, Plate XXXIX fig. 10;
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<bibRefCitation id="7BAA8E36AA6EBC2CEA6727268A640EBF" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
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||
: 124 (non
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<taxonomicName id="189BF3DDF40E2A420A4EF3F714491F6A" family="Hydrocenidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="G. hungerfordi" order="Cycloneritida" pageId="14" pageNumber="49" rank="species" species="hungerfordi">G. hungerfordi</taxonomicName>
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Godwin-Austen, 1889;
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<taxonomicName id="2916A8D7C9C75DFE5B441ED0217AACFB" family="Hydrocenidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="G. javana" order="Cycloneritida" pageId="14" pageNumber="49" rank="species" species="javana">G. javana</taxonomicName>
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<normalizedToken id="2EB76B155AE6EF14961C3D80D4B2E151" originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1897;
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<taxonomicName id="ED7BEC96D86A08E0C2B0F3D5704101A8" family="Hydrocenidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="G. javana" order="Cycloneritida" pageId="14" pageNumber="49" rank="subspecies" species="javana" subspecies="intermedia">G. javana intermedia</taxonomicName>
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<normalizedToken id="34B781F3934B18AD0C66EAAAD4C10A78" originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1897).
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</paragraph>
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<paragraph id="85FF8A1784320A0B61CA3F65E7D7513D" pageId="14" pageNumber="49">
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<taxonomicName id="D05A0F360701288B7B165375E5FA3511" class="Gastropoda" family="Hydrocenidae" genus="Hydrocena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrocena williamsi" order="Cycloneritida" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="williamsi">Hydrocena williamsi</taxonomicName>
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(Godwin-Austen, 1889):
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<bibRefCitation id="CE61F9CA6ACE513EA09240CCC01192B2" author="Saul, M" journalOrPublisher="Sabah Society Journal" pageId="41" pageNumber="76" pagination="105 - 110" title="Shell collecting in the limestone caves of Borneo." volume="3" year="1967">Saul 1967</bibRefCitation>
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: 109.
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</paragraph>
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<paragraph id="AB314F4190FC6EFCB1BEF7A282771030" pageId="14" pageNumber="49">
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<taxonomicName id="B5B128AFB24331EC469799E524B741C5" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="14" pageNumber="49" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
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sp.1 (Godwin-Austen, 1889):
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<bibRefCitation id="DCB77F9DC8F5DABA06F33974033CE6B8" author="Clements, R" journalOrPublisher="Biological Conservation" pageId="39" pageNumber="74" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="https://doi.org/10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
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: Appendix D.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="663C004EB34BAA3CEBBDE5C9844DF84D" pageId="14" pageNumber="49" type="type locality">
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<paragraph id="BC945AD3E0E70B32DF68943F54EF6ACE" pageId="14" pageNumber="49">Type locality.</paragraph>
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<paragraph id="468B2341DCA63D18E5A1A9DAAB5E0907" pageId="14" pageNumber="49">Borneo.</paragraph>
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</subSubSection>
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<subSubSection id="C9B7B5B87FC6FA5C3B41F60870524323" pageId="14" pageNumber="49" type="type material">
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<paragraph id="C974333D365E4DDABED8A9D2631C1194" pageId="14" pageNumber="49">Type material.</paragraph>
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<paragraph id="E72DAB7C955DFF7EBEE9B469BBC4AFA9" pageId="14" pageNumber="49">
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Holotype (Holotype by original monotypy). Borneo: NHMUK 1889.12.7.71 (glued on paper) (Fig. 1C) (
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<bibRefCitation id="1CE37466AB07A9032700A61DD42D5EC4" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
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||
).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EAF7F0BCB302C3E044777AB7A13AA32F" pageId="14" pageNumber="49" type="materials_examined">
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<paragraph id="D240467A16DA41539C0FEC3504FFABA2" pageId="14" pageNumber="49">Other material.</paragraph>
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<paragraph id="B35AC622E4107D9089432EE842EBD45F" pageId="14" pageNumber="49">
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Batu Punggul, Sepulut valley, Interior province, Sabah (
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<geoCoordinate id="18C1A5E947FBF7741F1BC697615DA541" direction="north" orientation="latitude" precision="9" value="4.65">04°39.00'N</geoCoordinate>
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,
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<geoCoordinate id="7BD4220E733105550E2B7136FEFD8665" direction="east" orientation="longitude" precision="9" value="116.61667">116°37.00'E</geoCoordinate>
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): RMNH/MOL 187642, BOR/MOL 57, JJV 1907. Gua Pungiton, Sepulut valley, Interior province, Sabah (
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<geoCoordinate id="67DEE0A313C8B6BA9EF5C51ED560E5D0" direction="north" orientation="latitude" precision="9" value="4.7068334">04°42.41'N</geoCoordinate>
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,
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<geoCoordinate id="095F76579D8873B483BBAE7CDB9334D8" direction="east" orientation="longitude" precision="9" value="116.60067">116°36.04'E</geoCoordinate>
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): BOR/MOL 55, JJV 7543. Bukit Tinahas, Sepulut valley, East end of Batu Punggul limestone, Interior province, Sabah (
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<geoCoordinate id="FCCF3F713317A76BDF8D68F1FEA6956C" direction="north" orientation="latitude" precision="9" value="4.638">04°38.28'N</geoCoordinate>
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,
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<geoCoordinate id="9DB49CDE47C925090D5C097279A046F6" direction="east" orientation="longitude" precision="9" value="116.6175">116°37.05'E</geoCoordinate>
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): JJV 7623. Tinahas limestone hill, Interior Province, Sabah (
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<geoCoordinate id="83FED3298FEFE8D599D5828F486E4FE1" direction="north" orientation="latitude" precision="9" value="4.641">04°38.46'N</geoCoordinate>
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,
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<geoCoordinate id="489DCC4CE8AC8666A90284197EDDD4C3" direction="east" orientation="longitude" precision="9" value="116.618">116°37.08'E</geoCoordinate>
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): RMNH/MOL 333928, RMNH/MOL 334016, BOR/MOL 56, BOR/MOL 59. Batu Temurung, Sepulut valley, Interior province, Sabah (
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<geoCoordinate id="EA663D722052D4FF40AAD7358C94CE2A" direction="north" orientation="latitude" precision="9" value="4.7075">04°42.45'N</geoCoordinate>
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,
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<geoCoordinate id="6C37F18EC83CAB13307817C5B7170E11" direction="east" orientation="longitude" precision="9" value="116.57333">116°34.40'E</geoCoordinate>
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): BOR/MOL 58, BOR/MOL 60, JJV 8037. Simbaluyon limestone hill, Interior Province, Sabah, (
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<geoCoordinate id="856139E57931B9E590E5EC4552C7C5F3" direction="north" orientation="latitude" precision="9" value="4.7208333">04°43.25'N</geoCoordinate>
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,
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<geoCoordinate id="7D50C572334ADA5411EB1CFF83EAFE87" direction="east" orientation="longitude" precision="9" value="116.570335">116°34.22'E</geoCoordinate>
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): RMNH/MOL 333922, RMNH/MOL 333946 (Fig. 8), RMNH/MOL 334007. Batu Baturong ca. 50 km W.S.W. of Lahad Datu, Tawau province, Sabah (
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<geoCoordinate id="8B7E6031C25A5BC292712BA790B9B21D" direction="north" orientation="latitude" precision="9" value="4.6833334">04°41.00'N</geoCoordinate>
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,
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<geoCoordinate id="54AB5B4F4FDE2628B9854C630D6D7AA0" direction="east" orientation="longitude" precision="9" value="118.01667">118°1.00'E</geoCoordinate>
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): JJV 1830. Madai limestone hill, Tawau Province, Sabah (
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<geoCoordinate id="5E509F3B8B6DC04A8A57E1286B53043D" direction="north" orientation="latitude" precision="9" value="4.727667">04°43.66'N</geoCoordinate>
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,
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<geoCoordinate id="915EA8A112AE8B0D2BF8D888B3E6B5AC" direction="east" orientation="longitude" precision="9" value="118.1785">118°10.71'E</geoCoordinate>
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): RMNH/MOL 337817, RMNH/MOL 337827, RMNH/MOL 337834. Cave on Teck Guan estate, Lahad Datu, Sabah: ZMA/MOLL 315607, ZMA/MOLL 315608, ZMA/MOLL 315609, ZMA/MOLL 315622.
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</paragraph>
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<caption id="90490946783D2DA7FC40E90D1F0C6807" pageId="14" pageNumber="49">
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<paragraph id="388234057FACAC55BC6F4510C4FCEFB8" pageId="14" pageNumber="49">
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Figure 8.
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<taxonomicName id="ED0C068221832DFE71E0DBFD66C0EBEB" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa williamsi" order="Cycloneritida" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="williamsi">Georissa williamsi</taxonomicName>
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Godwin-Austen, 1889.
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<normalizedToken id="3DFE26AD0760746AC2838D5F2F158071" originalValue="A–I">A-I</normalizedToken>
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RMNH/MOL 333946 A, D shell apertural view B shell side view C shell rear view E, F shell cross-section from 3D model G shell top view H protoconch side view I close up of protoconch from top at 1000
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<normalizedToken id="2B621C419D7789A87A11AC7B7FFBE04D" originalValue="×">x</normalizedToken>
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magnification. Scale bars: 500
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||
<normalizedToken id="1796981403EF20A4B4D760FA54356F1B" originalValue="µm">µm</normalizedToken>
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(
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||
<normalizedToken id="64B54E3980FE322391555E0C75DCB087" originalValue="A–G">A-G</normalizedToken>
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); 200
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<normalizedToken id="86155D9B7EF8514128177198824C1814" originalValue="µm">µm</normalizedToken>
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(H); 10
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<normalizedToken id="1F5ADD0EFE1C606022CB5E17121AC602" originalValue="µm">µm</normalizedToken>
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(I).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="3AEC1753E1BC20D5FAF6F6E05D653363" pageId="14" pageNumber="49" type="description">
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<paragraph id="C5C8BEAA3CE86B2840A424A91E7F5356" pageId="14" pageNumber="49">Description.</paragraph>
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<paragraph id="5D4549039189B0A2B68B4E01168D95A9" pageId="14" pageNumber="49">
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Protoconch. Colour: orange to red. Sculpture: a mix of rounded, ellipsoidal to irregular sculptural shape. Mesh width: 2-6
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<normalizedToken id="51005B92E7F17819E74F67C47BD55C6B" originalValue="µm">µm</normalizedToken>
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. Teleoconch. Colour: orange to red. First whorl: convex. Subsequent whorls: convex. Suture: clearly impressed. Shoulder: narrow. Number of whorls: 3-3
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<normalizedToken id="55B046DA36EFF7400794E191D445BB3C" originalValue="¼">1/4</normalizedToken>
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. SH: 1.58-1.91 mm. SW: 1.17-1.42 mm. SI: 1.30-1.38. Shell sculpture. Radial sculpture: absent, only weak to strong growth lines are visible at irregular intervals. Spiral sculpture: present, thin, regularly spaced at the first whorl, appearing immediately after the protoconch, on later whorls the spiral sculpture weakens and becomes distorted by the growth lines; more than 20 spiral ribs on the body whorl of the adult individual. Aperture. Shape: semi-elliptic, straight to concave parietal side, palatal side rounded, palatal edge contiguous with the body whorl, basal side slightly convex. AH: 0.63- 0.81 mm. AW: 0.71-0.87 mm. AI: 0.89-0.95.
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</paragraph>
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</subSubSection>
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<subSubSection id="EB2C143E642777D27135AF3806C4B907" lastPageId="15" lastPageNumber="50" pageId="14" pageNumber="49" type="diagnosis">
|
||
<paragraph id="256CC8A07D5FA0A969E9D31349BBCC9A" pageId="14" pageNumber="49">Diagnosis.</paragraph>
|
||
<paragraph id="FF779F0F06BDB956E58C9F5A7BE61E55" lastPageId="15" lastPageNumber="50" pageId="14" pageNumber="49">
|
||
<taxonomicName id="74ECEC2DD1188249994BDEC87A1598EA" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa williamsi" order="Cycloneritida" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="williamsi">Georissa williamsi</taxonomicName>
|
||
has a broad final whorl, in which it is similar to
|
||
<taxonomicName id="EF64E098002114DBB42891376F39668E" lsidName="G. gomantonensis" pageId="14" pageNumber="49" rank="species" species="gomantonensis">G. gomantonensis</taxonomicName>
|
||
and
|
||
<taxonomicName id="AFABC06EB332DCADF228C1CD7F979A5B" lsidName="G. everetti" pageId="14" pageNumber="49" rank="species" species="everetti">G. everetti</taxonomicName>
|
||
. However, these three species are all distinctly sculptured, where
|
||
<taxonomicName id="2C191B3B4D996B2EBC5D55F3FB63C40E" lsidName="G. gomantonensis" pageId="14" pageNumber="49" rank="species" species="gomantonensis">G. gomantonensis</taxonomicName>
|
||
has raised spiral sculpture,
|
||
<taxonomicName id="F84BF79D09E3355406FA3F79B6491D38" lsidName="G. everetti" pageId="14" pageNumber="49" rank="species" species="everetti">G. everetti</taxonomicName>
|
||
has oblique spiral sculp
|
||
<pageBreakToken id="ECD9D7FAF1A401681FE9F1A46E7B56B8" pageId="15" pageNumber="50" start="start">ture</pageBreakToken>
|
||
, but
|
||
<taxonomicName id="4EBC7F4964F5325045A45410F0C6C129" lsidName="G. williamsi" pageId="15" pageNumber="50" rank="species" species="williamsi">G. williamsi</taxonomicName>
|
||
has thin, hardly raised, and densely arranged spiral sculpture (4-6 ribs in every 0.1 mm), despite the similar shell habitus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC2848B4C9474AE54622A9D916AAE04B" pageId="15" pageNumber="50" type="distribution">
|
||
<paragraph id="FE8127BDDEA4375091B3ED20D8476416" pageId="15" pageNumber="50">Distribution.</paragraph>
|
||
<paragraph id="01B0F11DE8F1FF53C9609DCF5F2408CE" pageId="15" pageNumber="50">
|
||
<taxonomicName id="C53F6E22D9AF6784709D0F64DDBC7C3E" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa williamsi" order="Cycloneritida" pageId="15" pageNumber="50" phylum="Mollusca" rank="species" species="williamsi">Georissa williamsi</taxonomicName>
|
||
occurs over a large part of Sabah from the Sepulut valley in the west-central to Tawau and Lahad Datu in the east.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="43EAD9341CBDBF7F91899019A4DD77E9" pageId="16" pageNumber="51" type="discussion">
|
||
<paragraph id="93855C9753C4D82343AB99C05BF986D9" pageId="16" pageNumber="51">
|
||
<pageBreakToken id="D381BAA2CB880F8900F16F49FBE45A01" pageId="16" pageNumber="51" start="start">Discussion</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="38922CFAF40CD51FD7D6114BA252E497" pageId="16" pageNumber="51">
|
||
The type locality of
|
||
<taxonomicName id="19C7A9F34F8A97C03903C587718EADA8" lsidName="G. williamsi" pageId="16" pageNumber="51" rank="species" species="williamsi">G. williamsi</taxonomicName>
|
||
is
|
||
<normalizedToken id="C376F791C6EC691251FB0D4CDDBC9A30" originalValue="‘Borneo’">'Borneo'</normalizedToken>
|
||
, with no specific location stated by
|
||
<bibRefCitation id="34E02203A50769873B06888B0FB8FE96" author="Godwin-Austen, HH" journalOrPublisher="Molecular Marine Biology and Biotechnology" pageId="39" pageNumber="74" url="https://doi.org/10.1111/j.1469-7998.1891.tb06805.x" year="1889">Godwin-Austen (1889)</bibRefCitation>
|
||
. Saul (1966) in her note on "Shell collecting in the limestone cave of Borneo" mentioned that during her trip to Lahad Datu, Sabah, they collected
|
||
<taxonomicName id="A1548687C976221DBE7ADD3FA89890DC" lsidName="G. williamsi" pageId="16" pageNumber="51" rank="species" species="williamsi">G. williamsi</taxonomicName>
|
||
(syn.
|
||
<taxonomicName id="E30B30D2DBA3CDA80256B5118C5BF0E6" class="Gastropoda" family="Hydrocenidae" genus="Hydrocena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrocena williamsi" order="Cycloneritida" pageId="16" pageNumber="51" phylum="Mollusca" rank="species" species="williamsi">Hydrocena williamsi</taxonomicName>
|
||
). Based on the characters of
|
||
<taxonomicName id="66B77A0BE08E9041C49DB936F4E76D2A" lsidName="G. williamsi" pageId="16" pageNumber="51" rank="species" species="williamsi">G. williamsi</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="08A3C903FC20281EEA9EBA1A6BF245E2" author="Godwin-Austen, HH" journalOrPublisher="Molecular Marine Biology and Biotechnology" pageId="39" pageNumber="74" url="https://doi.org/10.1111/j.1469-7998.1891.tb06805.x" year="1889">Godwin-Austen (1889)</bibRefCitation>
|
||
and the type material we have examined, the species does not have very prominent spiral sculpture.
|
||
<taxonomicName id="EF35FC7176AFAC805F71AC6293E3B286" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa williamsi" order="Cycloneritida" pageId="16" pageNumber="51" phylum="Mollusca" rank="species" species="williamsi">Georissa williamsi</taxonomicName>
|
||
was previously misinterpreted as having highly raised spiral sculpture, and the name was therefore misapplied to forms like
|
||
<taxonomicName id="E3415472523B887135625B3F6A39E5ED" lsidName="G. hungerfordi" pageId="16" pageNumber="51" rank="species" species="hungerfordi">G. hungerfordi</taxonomicName>
|
||
,
|
||
<taxonomicName id="88D52EAE6102CACD93BD742EF1EC6A2D" lsidName="G. insulae" pageId="16" pageNumber="51" rank="species" species="insulae">G. insulae</taxonomicName>
|
||
, and
|
||
<taxonomicName id="3B846A6D94CFAA07D6C9828AA3FBCF8E" lsidName="G. javana" pageId="16" pageNumber="51" rank="species" species="javana">G. javana</taxonomicName>
|
||
(
|
||
<bibRefCitation id="644AE90FFF4836DBAC3EA1450911DB73" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C9065F79FA78E24A81693B5907E1F71A" author="Phung, CC" journalOrPublisher="ZooKeys" pageId="40" pageNumber="75" pagination="49 - 104" title="A checklist of land snails from the west coast islands of Sabah, Borneo (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.673.12422" volume="673" year="2017">Phung et al. 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C438729259262D593B461063B8C739AA" author="Vermeulen, JJ" journalOrPublisher="Backhuys Publishers, Leiden" pageId="42" pageNumber="77" title="Fauna Malesiana guide to the land snails of Bali." year="1998">Vermeulen and Whitten 1998</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |