204 lines
24 KiB
XML
204 lines
24 KiB
XML
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<mods:title id="A3EB92DDBDB471DD622E78FA9263ADED">A new species of Notarius (Siluriformes: Ariidae) from the Colombian Pacific.</mods:title>
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<mods:name id="8F72D0E4A030CB78C84B2395ED699212" type="personal">
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<mods:roleTerm id="9096650D5F93C9E18BBD62E4A6C40240">Author</mods:roleTerm>
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<mods:namePart id="67E0FF8FFD25A02B3398D2385AF6466A">Ricardo Betancur-R.</mods:namePart>
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</mods:name>
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<mods:name id="7B056E41C4E54F6E4A2DD6858ED18318" type="personal">
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<mods:namePart id="595A39F347185B2DE4386E6632FC938D">Arturo Acero P.</mods:namePart>
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<mods:title id="7BBB88310CA00CB0B5A77114D120E8EC">Zootaxa</mods:title>
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<mods:part id="5F839EAE7F14CB4E407B30F04563631E">
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<mods:date id="1F9D6332B7A8608C0FFD6B1C1B626CAA">2006</mods:date>
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<mods:number id="4FD6B31D0F4E089767A25876BA9D9777">1249</mods:number>
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<mods:url id="16ACBE08F07CB3FFA1639E2155E60EE8">http://www.zoobank.org/urn:lsid:zoobank.org:pub:55909877-7547-483F-8210-9F0522D81E0A</mods:url>
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<mods:classification id="3F59B86DDED08C6E2317C30DB97F2D3B">journal article</mods:classification>
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<mods:identifier id="592C36BAA919430050EE77B9FA522B87" type="Plazi-Custom">z01249p047</mods:identifier>
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<mods:identifier id="6FD8667AFAE46CA6E5F0D9C4F835F77A" type="ZooBank">55909877-7547-483F-8210-9F0522D81E0A</mods:identifier>
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<treatment id="8449D05AFE41D640777332E6DDA92DA8" ID-DOI="http://doi.org/10.5281/zenodo.6258311" ID-GBIF-Taxon="100122051" ID-Zenodo-Dep="6258311" LSID="urn:lsid:plazi:treatment:8449D05AFE41D640777332E6DDA92DA8" httpUri="http://treatment.plazi.org/id/8449D05AFE41D640777332E6DDA92DA8" lastPageNumber="56" pageNumber="50">
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<subSubSection id="E066C1E40942A2196119FAF27E434B4F" type="nomenclature">
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<paragraph id="81BE3D360A17425B0D9249E4579134B8" pageNumber="50">
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<taxonomicName id="A9E17DF83FB3A685479861366298D152" ID-CoL="47S79" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="50" phylum="Chordata" rank="species" species="armbrusteri">Notarius armbrusteri</taxonomicName>
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<taxonomicNameLabel id="FA0EBAFD5B33A4FCAFE33986B6F3D074" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="4512DC963880E08167A0C0C5D32572D2" type="description">
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<paragraph id="D2879CDA3A9033EA74BE3EE5D90ADEBF" pageNumber="50">Figs. 2-3</paragraph>
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</subSubSection>
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<subSubSection id="347D9B027C49BD1F21298A120F743824" type="reference_group">
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<paragraph id="107E2AB84710E057D5E3F36B39E1F544" pageNumber="50">
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<taxonomicName id="7D00E8797FF3DCA13304153E97602948" LSID-ZBK="urn:lsid:zoobank.org:act:3CD2092D-E0A4-4C39-806B-3BD5ABD5E67F" family="Ariidae" genus="Arius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Arius planiceps Steindachner 1877:576 [26]" order="Siluriformes" pageNumber="50" phylum="Chordata" rank="species" species="planiceps">Arius planiceps</taxonomicName>
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non Steindachner: Acero P. and Betancur-R. (2002): 9, Fig. 3.
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</paragraph>
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</subSubSection>
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<subSubSection id="D4335BF2FC7ECAAB659EEBCB9992BD81" type="nomenclature">
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<paragraph id="1D353161F02E1ECCFC80E187A4D4D27E" pageNumber="50">
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<materialsCitation id="D6CBB4C18FD1471E3D2917FAE982FFEE" ID-GBIF-Occurrence="923892181" country="Colombia" stateProvince="Valle Del Cauca">
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<typeStatus id="2880E29D4FEC245B55B2DDA76884C461">Holotype</typeStatus>
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.
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<collectionCode id="7A34A48683B48862E6BDA89317CACCDE">INVEMAR-PEC</collectionCode>
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6677 (undeposited tissue tagged as 527), female, 194 mm SL, purchased by R. Betancur-R. and A. Acero P.,
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<date id="365BFE8446A02E8265E5B16133104796" day="28" month="05" value="2004-05-28" year="2004">28 May 2004</date>
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, fish market of Buenaventura,
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<collectingRegion id="C1A7EE9097271F6E29FEB2265C3569A5">Valle del Cauca</collectingRegion>
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(VC),
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<collectingCountry id="1A4E6F307C946805365E222637CF425B">Colombia</collectingCountry>
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(CO); cyt b, ATPase 8 and ATPase 6 sequences are available in GenBank, accession numbers DQ373045, DQ373041, and DQ373043, respectively.
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</materialsCitation>
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</paragraph>
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<paragraph id="6CB0D304E38E075ABCE6B7EB6A8AC1DB" pageNumber="50">
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<materialsCitation id="A391B87B577EBDAD84C69D6936DDC319" ID-GBIF-Occurrence="923892179" country="Colombia" stateProvince="Valle Del Cauca">
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<typeStatus id="38095E67CB3C2CE2055083D7BFD5DE61">Paratypes</typeStatus>
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.
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<collectionCode id="343376D95F0DDE5B19E5D5F72153F94E">INVEMAR-PEC</collectionCode>
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6678 (undeposited tissue tagged as 529), male, 177 mm SL, same collection data as holotype; cyt b, ATPase 8 and ATPase 6 sequences are available in GenBank, accession numbers DQ373046, DQ373042, and DQ373044, respectively.
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</materialsCitation>
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<materialsCitation id="BC6400E2FC1618E2B55E112B52EB10F3" ID-GBIF-Occurrence="923892178" country="Colombia" stateProvince="Valle Del Cauca">
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<collectionCode id="C097F542BC6A2ECE16CF2BFF8EE997E2">ICN-MHN</collectionCode>
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14829, female, 208 mm SL, unsexed specimen, same collection data as holotype.
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</materialsCitation>
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<materialsCitation id="77F8CA1767FCC42359AE59E7543A2684" ID-GBIF-Occurrence="923892180" country="Colombia" stateProvince="Valle Del Cauca">
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<collectionCode id="A07C97774B4AE5D1574A1A33B33B6C08" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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292738, female, 205 mm SL, purchased by W.R. Taylor,
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<date id="9C7246DBF79965D48B740FBDDF003BB8" day="17" month="10" value="1970-10-17" year="1970">17 October 1970</date>
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, fish market of Buenaventura,
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<collectingRegion id="BDB49A2D4E925CBBFCFA7D5FB134EEE4">VC</collectingRegion>
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,
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<collectingCountry id="8AE5F0FF44DAFFFCF088EC2EE4455F5D">CO</collectingCountry>
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.
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</materialsCitation>
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<materialsCitation id="A1D3E7C67FE7DFD2E093B1045CE5DEC3" ID-GBIF-Occurrence="923892182" country="Colombia" stateProvince="Valle Del Cauca">
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<collectionCode id="331C7BFA5D61943972F62F710985DD75" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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264834, skeletonized specimen, 69 mm skull length, purchased at fish market of Buenaventura,
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<collectingRegion id="F011B615CD00EA186563FB13E0DC722C">VC</collectingRegion>
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,
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<collectingCountry id="83EC1A82A5F5E183A98F384CC86B3332">CO</collectingCountry>
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, catalogued
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<date id="F136626160D20E2C2A25DD636C0B123E" day="25" month="01" value="1984-01-25" year="1984">25 January 1984</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DDD26B7CE5F34039380775BCAE9079D7" type="diagnosis">
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<paragraph id="04B09A42EAE134B62283C3DE88CC2BB2" pageNumber="50">
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Diagnosis.
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<taxonomicName id="FB0C6318840C88A5AEC361AC73EA85C6" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="50" phylum="Chordata" rank="species" species="armbrusteri">Notarius armbrusteri</taxonomicName>
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is distinguished from other EP species of
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<taxonomicName id="687D2184690761BF6B2B99D0BA32F270" LSID-ZBK="urn:lsid:zoobank.org:act:086E7B0F-D030-44BD-9DCC-8CAEB178CBAA" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Notarius Gill 1863:171" order="Siluriformes" pageNumber="50" phylum="Chordata" rank="genus">Notarius</taxonomicName>
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by the following combination of features: mouth rather small, its width 11.1-11.8% SL; eye large, diameter 4.3-4.9% SL; distance between anterior nostrils 6.1-6.9% SL, distance between posterior nostrils 5.9-6.9% SL; short maxillary barbels, barely reaching pectoralfin bases, their length 20.5-22.2% SL; and gill rakers on first arch 3-4+8-9 (total 11-13). Meristic and morphometric data of the type series are summarized in Tables 2 and 3, respectively. Selected features distinguishing
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<taxonomicName id="26FB89467D900A486541E09E119094B1" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="50" phylum="Chordata" rank="species" species="armbrusteri">N. armbrusteri</taxonomicName>
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from the seven other EP species of
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<taxonomicName id="64D2FCDC4780C5F792A904FB77AFFB9D" LSID-ZBK="urn:lsid:zoobank.org:act:086E7B0F-D030-44BD-9DCC-8CAEB178CBAA" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Notarius Gill 1863:171" order="Siluriformes" pageNumber="50" phylum="Chordata" rank="genus">Notarius</taxonomicName>
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are summarized in Table 4. See key section at the end for diagnostic characters distinguishing the new species from other EP ariid taxa.
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</paragraph>
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</subSubSection>
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<subSubSection id="64F8C3050566F222C1119CBD9F48C587" type="description">
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<paragraph id="3EC0E448D6BECA318DFA95987F8EABF3" pageNumber="51">Description (based on combined data from type series). Body depth 5.0-5.4 in SL; body width 4.7-5.2 in SL. Head not elongated, moderately depressed anteriorly, posterior profile slightly convex, length 3.7-4.1 in SL, width 1.1-1.2 in HL, depth 1.5-1.7 in HL. Snout rounded, length 3.1-3.4 in HL. Mouth inferior to subterminal, width 2.1-2.4 in HL. Lips thin to moderately thick, upper lip width 9.8-18.5 in HL. Maxillary barbels 1.2-1.3 in HL; mandibulary barbels 1.6-1.8 in HL, passing gill membrane; mental barbels 2.2-2.4 in HL, not reaching gill membrane. Distance between anterior nostrils 3.9-4.0 in HL; distance between posterior nostrils 3.9-4.2 in HL, no fleshy furrow connecting nostrils. Interorbital distance 1.9-2.2 in HL. Eyes lateral, diameter 5.1-6.2 in HL, 2.6-3.0 in interorbital distance. Postorbital length 1.8-2.0 in HL. Head shield exposed, covered posteriorly with dense layer of tiny granules extending anteriorly to opposite eyes (granules sometimes not very apparent). Frontal depression broad, with large pale spot on anterior margin immediately after interorbital space. Supraoccipital process keeled, rather short, slightly convex; length 3.6-4.7 in HL; width 3.6-4.0 in HL, 0.8-1.0 in length. E pioccipital bones (= epiotics) not invading or slightly invading skull surface (Fig. 1B). Predorsal plate narrow, crescent-shaped. Premaxillary teeth villiform, arranged in two subrectangular bands forming wide arrow point (Fig. 4). Palate with villiform teeth, arranged in four patches: inner patches (vomerine) united medially, longer than wide; lateral patches broader, triangular, projected posteriorly, with an indentation into which fit inner patches (Fig. 4).</paragraph>
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<paragraph id="3E01B97CA6CD69502768138BD6B7B14E" lastPageNumber="53" pageNumber="52">Predorsal fin length 2.6-3.1 in SL. Dorsal-fin base 8.3-9.0 in SL; dorsal-fin spine long, rather thin, longer than pectoral-fin spine, with small serrations along external margin, inner margin slightly serrated only in distal end; height 4.4-4.7 in SL. Distance between dorsal fin and adipose fin 3.5-3.9 in SL. Base of adipose fin 8.0-8.6 in SL, as long as or somewhat longer than base of dorsal fin, 0.9-1.0 in dorsal-fin base; height of adipose fin 9.1-10.2 in SL. Pectoral-fin base 15.5-16.7 in SL; pectoral-fin spine long, thin, slightly serrated along external margin, serrations on inner margin smaller and absent near proximal end; length 4.6-5.0 in SL. Pelvic-fin base 20.0-21.3 in SL; pelvic-fin length 5.3-5.7 in SL in females and 7.1 in SL in one male. Anal-fin base 6.0-6.5 in SL; anal-fin height 5.2-5.7 in SL. Caudal peduncle depth 13.5-14.3 in SL. Caudal fin deeply forked, lower lobe shorter than upper lobe. Lateral line originating below predorsal plate, tilting ventrally to mid-body line at about level of pelvic-fin origin, running posteriorly, bending dorsally before caudal fin. Gill rakers on second arch 3-4+9 (total 12-13). Dorsal fin elements I,7; pectoral fin elements I,11-12; pelvic fin elements 6; anal fin elements 19-22.</paragraph>
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</subSubSection>
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<subSubSection id="C48E7460CB28E267D67405D2B0BA6969" type="description">
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<paragraph id="CE1E2B2586D5875B94C41B58431E95B7" pageNumber="56">Coloration. In life, dorsum brownish grey with metallic tinges, flanks and venter whitish. Lower caudal fin lobe and tip of anal fin dark; pectoral and pelvic fins dark, distal tips pale. In alcohol, coloration fades, metallic tinges disappear.</paragraph>
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<paragraph id="7798300DE31AB88A2DE5FFB6AEFD8E4E" pageNumber="56">
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Size. Largest female specimen examined 208 mm SL (ICN-MHN 14829, paratype), largest male 177 mm SL (INVEMAR-PEC 6678, paratype). Female ariids have a greater development of pelvic fins than have males, and this dimorphism exhibits a positive allometry becoming more evident in mature specimens (Acero P. et al., 2005). Given that there is a clear difference in the size of pelvics between female (17.5-18.5% SL) and male (14.1% SL) specimens, it seems likely that
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<taxonomicName id="2ABDBE028EE8D590AD8C09AD7A1E30AF" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">Notarius armbrusteri</taxonomicName>
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reaches maturity below 200 mm SL and is probably the smallest known species of
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<taxonomicName id="4F394337B5EB28521BB5C427D03F4D53" LSID-ZBK="urn:lsid:zoobank.org:act:086E7B0F-D030-44BD-9DCC-8CAEB178CBAA" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Notarius Gill 1863:171" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Notarius</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="A219D2CE0047E349C7B1FAC2DD7ACCF3" type="distribution">
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<paragraph id="94A4867B2BEA8C14A30932D0D2CF06A2" pageNumber="56">
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Distribution and habitat.
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<taxonomicName id="5399D933F0916B496D54B9F065D9FF9E" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">Notarius armbrusteri</taxonomicName>
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is known only from Buenaventura, Valle del Cauca, Colombian Pacific. The species has not been recorded from Tumaco,
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<normalizedToken id="57CD43D91F64DA91B1E5B315DC9D8DEE" originalValue="Nariño">Narino</normalizedToken>
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, CO, about 300 km southwestwards from Buenaventura (pers. obs.). Habitat preferences are unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="A133A4F52F408A05D865CB22F68796C0" type="etymology">
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<paragraph id="FD463C69A6A215CAC94D7E034382E188" pageNumber="56">
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Etymology. Named for Dr. Jonathan W. Armbruster, curator of the Auburn University Museum, in recognition of his important contributions to the taxonomy of neotropical catfishes. We suggest the common names of “
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<normalizedToken id="F9D1442B143084FCE7451547BA3A38EC" originalValue="Ñato">Nato</normalizedToken>
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Sea Catfish”, “bagre
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<normalizedToken id="65256F0986F18AD05E68C9469E80D26A" originalValue="ñato">nato</normalizedToken>
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” (Spanish, name given by fishermen in the area), and “
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<normalizedToken id="7CACD66BB71DA3070A6AADC508D035CB" originalValue="mâchoiron">machoiron</normalizedToken>
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<normalizedToken id="2AE32016509D629874A2531CA0021D52" originalValue="ñato">nato</normalizedToken>
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” (French).
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</paragraph>
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</subSubSection>
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<subSubSection id="F5B5E4FF102595AAAAFD44E9561407B2" type="discussion">
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<paragraph id="7E3FA216A3F803CBF0CDCF3E392374CC" pageNumber="56">Discussion</paragraph>
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<paragraph id="B2F3BC1A814EF3698A81B35BA97E062C" pageNumber="56">
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A new molecular phylogeny for the species of
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<taxonomicName id="15215F711F5BC5992543B4DF85E9FC4A" LSID-ZBK="urn:lsid:zoobank.org:act:086E7B0F-D030-44BD-9DCC-8CAEB178CBAA" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Notarius Gill 1863:171" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Notarius</taxonomicName>
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, inferred from the combined cyt b + ATPase 8/6 dataset, is presented in Fig. 5. The mitochondrial evidence suggests that
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<taxonomicName id="B2E64503490343159B73288F1EF7C993" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">N. armbrusteri</taxonomicName>
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is the sister species of
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<taxonomicName id="B67AA51C7626738C170BED92028FB432" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
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, from the Pacific coast of Panama; the bootstrap supporting this clade is strong (100%). The combined mitochondrial K2P distances between haplotypes of
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<taxonomicName id="ABE9E14D5D9A1825D2E97A38F098BF9F" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">N. armbrusteri</taxonomicName>
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and
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<taxonomicName id="DE29AE5330C42258FBF7E54589C056F2" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
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are 4.1-4.2%. Both species are also related to the
|
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<taxonomicName id="6C97C70531D5BFFF733A14B9897070F9" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="planiceps">N. planiceps</taxonomicName>
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/aff.
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<taxonomicName id="F9D9714395ACD5AF71C77B2C9BDD2712" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="planiceps">planiceps</taxonomicName>
|
|
group (bootstrap 94%), known from the Pacific Panama to Mexico. Despite the few specimens in the type series (five complete and one skeletonized), the fact that
|
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<taxonomicName id="15255DE8E26100544F990C972142F7CB" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">N. armbrusteri</taxonomicName>
|
|
is well discriminated as a separate and monophyletic entity among EP species of
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<taxonomicName id="BEE7ACC23B657712EE4D53B86F3D2E44" LSID-ZBK="urn:lsid:zoobank.org:act:086E7B0F-D030-44BD-9DCC-8CAEB178CBAA" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Notarius Gill 1863:171" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Notarius</taxonomicName>
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in the phylogram, gives strong support for its specific status.
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</paragraph>
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<paragraph id="A9CE1D904C5C666988BED0EB614C2756" pageNumber="56">
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<taxonomicName id="1940B6FEC3849DA1E88C0B8949726D7A" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">Notarius armbrusteri</taxonomicName>
|
|
can be separated from its sister species,
|
|
<taxonomicName id="7E2D6CAC8CD5A7714282FED49201E47F" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
|
|
, by the following features: epioccipitals not invading or only slightly invading skull surface (Fig. 1B) (vs. widely invading in
|
|
<taxonomicName id="33478D313EB2A6F23ADFCB5BA1857DF4" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
|
|
, Fig. 1A), complex process (formed by the epioccipitals and the supraoccipital) absent (Fig. 1B) (vs. present in
|
|
<taxonomicName id="F9A3012B694F91D6D286D5EAD1D42F65" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
|
|
, Fig. 1A); two pairs of palatal tooth patches present (vs. three pairs in
|
|
<taxonomicName id="16B5A44FB9ACA0DE3E13BCFD8B412353" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
|
|
); smaller mouth (11.1-11.8% SL in
|
|
<taxonomicName id="34B9417621410F95E7D0CDB0D4F55DF1" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">N. armbrusteri</taxonomicName>
|
|
vs. 12.3-14.4% SL in
|
|
<taxonomicName id="667AFABD734584551D92E141EB6C4EAC" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
|
|
); shorter maxillary barbels (20.5-22.2% SL in
|
|
<taxonomicName id="EF80269B289CB9882B73CC04AE13076A" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">N. armbrusteri</taxonomicName>
|
|
vs. 26.7-30.3% SL in
|
|
<taxonomicName id="B1A1EB013B694132E47DC6EDF50E129A" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
|
|
); and larger eyes (4.3-4.9% SL in
|
|
<taxonomicName id="60F51F814B477BBCDA2A7C92201F2102" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="armbrusteri">N. armbrusteri</taxonomicName>
|
|
vs. 3.8-4.3% SL in
|
|
<taxonomicName id="DF313F737B1E1B08C3D4A2B26FB3084F" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="insculptus">N. insculptus</taxonomicName>
|
|
). Other EP species of
|
|
<taxonomicName id="5F62751EB9A6CAC2D7004947F90FCB6E" LSID-ZBK="urn:lsid:zoobank.org:act:086E7B0F-D030-44BD-9DCC-8CAEB178CBAA" family="Ariidae" genus="Notarius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Notarius Gill 1863:171" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Notarius</taxonomicName>
|
|
are compared in Table 4.
|
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</paragraph>
|
|
</subSubSection>
|
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</treatment>
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</document> |