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<document id="1BECF56C8E852A8DC0EAE7CF93576DA9" ID-DOI="10.5281/zenodo.13643699" ID-ISSN="1732-2421" ID-Zenodo-Dep="13643699" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142205826" checkinUser="felipe" docAuthor="Soriano, Carmen &amp; Delclòs, Xavier" docDate="2006" docId="455787BCFF9D2450FC94F8066A69FC5B" docLanguage="en" docName="ActaPalaeontolPol.51.1.185-200.pdf" docOrigin="Acta Palaeontologica Polonica 51 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Zygadenia diazromerali Soriano &amp; Delclòs, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="191" masterDocId="B96EFFC4FF982456FFB8FF876E36FFDB" masterDocTitle="New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family" masterLastPageNumber="200" masterPageNumber="185" pageNumber="190" updateTime="1725365340826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C4CD60DEC1447DDCA4F8EEF6300A039C">New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family</mods:title>
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<mods:namePart id="D46A31F7548C6796A8561CEB97E7D6EE">Soriano, Carmen</mods:namePart>
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<mods:namePart id="CF3AE84468893A11E740EA9623338518">Delclòs, Xavier</mods:namePart>
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<mods:title id="577F6D4CE6E8DEFA969926216948940E">Acta Palaeontologica Polonica</mods:title>
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<mods:date id="5B292AB7CCCC142C1EA9EF2194E20E14">2006</mods:date>
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<mods:number id="11EC1E59B75ED9CBE782221859FD1890">51</mods:number>
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<mods:identifier id="30DB241A28FF76FAD10500B6653529E1" type="ISSN">1732-2421</mods:identifier>
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<treatment id="455787BCFF9D2450FC94F8066A69FC5B" LSID="urn:lsid:plazi:treatment:455787BCFF9D2450FC94F8066A69FC5B" httpUri="http://treatment.plazi.org/id/455787BCFF9D2450FC94F8066A69FC5B" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190">
<subSubSection id="85E46521FF9D2453FC94F8066AFBF846" box="[812,1229,1921,1949]" pageId="5" pageNumber="190" type="nomenclature">
<paragraph id="CD4136AAFF9D2453FC94F8066AFBF846" blockId="5.[812,1229,1921,1984]" box="[812,1229,1921,1949]" pageId="5" pageNumber="190">
<heading id="960981C6FF9D2453FC94F8066AFBF846" box="[812,1229,1921,1949]" fontSize="12" level="2" pageId="5" pageNumber="190" reason="9">
<taxonomicName id="0AFE4D29FF9D2453FC94F8066A56F846" authorityName="Soriano &amp; Delclòs" authorityYear="2006" box="[812,1120,1921,1949]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="190" phylum="Arthropoda" rank="species" species="diazromerali" status="sp. nov.">
<emphasis id="FF8AEAB8FF9D2453FC94F8066A56F846" box="[812,1120,1921,1949]" italics="true" pageId="5" pageNumber="190">Zygadenia diazromerali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B957C3FF9D2453FBD1F8066AFBF846" box="[1129,1229,1921,1949]" pageId="5" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="85E46521FF9D2450FC94F82C6A69FC5B" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190" type="description">
<paragraph id="CD4136AAFF9D2453FC94F82C6DA8F81B" blockId="5.[812,1229,1921,1984]" box="[812,926,1963,1984]" pageId="5" pageNumber="190">
<figureCitation id="55C52A2FFF9D2453FC94F82C6D59F81B" box="[812,879,1963,1984]" captionStart="Fig" captionStartId="6.[102,135,605,625]" captionTargetBox="[107,1483,232,568]" captionTargetId="figure-704@6.[102,1486,239,571]" captionTargetPageId="6" captionText="Fig. 5. Camera lucida drawings of Barremian (Early Cretaceos) ommatin Zygadenia diazromerali sp. nov. from Las Hoyas, Cuenca Province. A. Holotype MCCM LH 12001a, part, dorsal (A1) and ventral (A2) aspects. B. Paratype MCCM LH 23851, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643711" httpUri="https://zenodo.org/record/13643711/files/figure.png" pageId="5" pageNumber="190">Figs. 5</figureCitation>
,
<figureCitation id="55C52A2FFF9D2453FCC2F82C6DACF81B" box="[890,922,1963,1984]" captionStart="Fig" captionStartId="7.[102,134,1379,1399]" captionTargetBox="[160,1427,225,1352]" captionTargetId="figure-233@7.[160,1427,225,1352]" captionTargetPageId="7" captionText="Fig. 6. Barremian (Early Cretaceous) ommatins Brochocoleus (Brochocoleini) and Zygadenia (Notocupedini), and cupedins Priacma (Priacmini) and Anaglyphites (Mesocupedini) from Spain. A. Zygadenia diazromerali sp. nov. Holotype MCCM LH 12001a, part, Las Hoyas, Cuenca Province. B. Zygadenia martinclosas sp. nov. Holotype IEI LC 2663a, part, La Cabrúa, Lleida Province. C. Zygadenia longicoxa sp. nov. Holotype MCCM LH 908030a, part, Las Hoyas, Cuenca Province, Lower Cretaceous. D. Zygadenia siniestri sp. nov. Holotype MCCM LH 23515a, part, Las Hoyas, Cuenca Province. E. Priacma sanzii sp. nov. Holotype MCCM LH 24524, part, Las Hoyas, Cuenca Province. F. Anaglyphites zherikhini sp. nov. Holotype MCCM LH 15122a, part, Las Hoyas, Cuenca Province. G.?Anaglyphites pluricavus sp. nov. Holotype IEI LP 97Ga, part, La Pedrera, Lleida Province. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643715" httpUri="https://zenodo.org/record/13643715/files/figure.png" pageId="5" pageNumber="190">6A</figureCitation>
.
</paragraph>
<paragraph id="CD4136AAFF9D2450FC94F8536F65FD2C" blockId="5.[812,1293,2004,2025]" lastBlockId="6.[102,775,710,1281]" lastPageId="6" lastPageNumber="191" pageId="5" pageNumber="190">
<materialsCitation id="7D963CF7FF9D2450FC94F8536F79FD2C" collectorName="La Cierva" country="Spain" lastPageId="6" lastPageNumber="191" location="Serrania de Cuenca" municipality="Cuenca Province" pageId="5" pageNumber="190" specimenCount="1">
1988
<taxonomicName id="0AFE4D29FF9D2453FCDAF8536A8CF832" authority=", Sanz et al." authorityName="Sanz" box="[866,1210,2004,2025]" class="Magnoliopsida" family="Malvaceae" genus="Coleoptera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="190" phylum="Tracheophyta" rank="subGenus" subGenus="Cupedidae">Coleoptera (Cupedidae), Sanz et al.</taxonomicName>
: pl. 1: 5.
<emphasis id="FF8AEAB8FF9E2450FFDEFD416ED3FD00" box="[102,229,710,731]" italics="true" pageId="6" pageNumber="191">
<typeStatus id="12458808FF9E2450FFDEFD416EA3FD00" box="[102,149,710,731]" pageId="6" pageNumber="191">Type</typeStatus>
locality
</emphasis>
: Las Hoyas fossil site,
<collectorName id="600B537CFF9E2450FE69FD416C03FD00" box="[465,565,710,731]" pageId="6" pageNumber="191">La Cierva</collectorName>
,
<location id="C8216071FF9E2450FDF9FD416D35FD00" LSID="urn:lsid:plazi:treatment:455787BCFF9D2450FC94F8066A69FC5B:C8216071FF9E2450FDF9FD416D35FD00" box="[577,771,710,731]" country="Spain" municipality="Cuenca Province" name="Serrania de Cuenca" pageId="6" pageNumber="191">Serranía de Cuenca</location>
,
<collectingMunicipality id="2D25ACD0FF9E2450FFDEFD656F3AFD2C" box="[102,268,738,759]" pageId="6" pageNumber="191">Cuenca Province</collectingMunicipality>
,
<collectingCountry id="B5E9763AFF9E2450FEADFD656F79FD2C" box="[277,335,738,759]" name="Spain" pageId="6" pageNumber="191">Spain</collectingCountry>
</materialsCitation>
.
</paragraph>
<caption id="99816622FF9E2450FFDEFDDA6B86FD56" ID-DOI="http://doi.org/10.5281/zenodo.13643711" ID-Zenodo-Dep="13643711" httpUri="https://zenodo.org/record/13643711/files/figure.png" pageId="6" pageNumber="191" startId="6.[102,135,605,625]" targetBox="[107,1483,232,568]" targetPageId="6" targetType="figure">
<paragraph id="CD4136AAFF9E2450FFDEFDDA6B86FD56" blockId="6.[102,1485,604,656]" pageId="6" pageNumber="191">
Fig. 5. Camera lucida drawings of Barremian (Early Cretaceos) ommatin
<taxonomicName id="0AFE4D29FF9E2450FD4FFDDA6DE6FDAA" authorityName="Soriano &amp; Delclòs" authorityYear="2006" box="[759,976,605,625]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="diazromerali" status="sp. nov.">
<emphasis id="FF8AEAB8FF9E2450FD4FFDDA6DE6FDAA" box="[759,976,605,625]" italics="true" pageId="6" pageNumber="191">Zygadenia diazromerali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B957C3FF9E2450FC6CFDD96A2AFDAA" box="[980,1052,606,625]" pageId="6" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
from Las Hoyas, Cuenca Province.
<emphasis id="FF8AEAB8FF9E2450FAE6FDDB6B58FDAB" bold="true" box="[1374,1390,604,624]" pageId="6" pageNumber="191">A</emphasis>
. Holotype MCCM LH 12001a, part, dorsal (A
<subScript id="517A34EFFF9E2450FE10FD056F86FD4B" attach="right" box="[424,432,642,656]" fontSize="6" pageId="6" pageNumber="191">1</subScript>
) and ventral (A
<subScript id="517A34EFFF9E2450FDF8FD056C7EFD4B" attach="right" box="[576,584,642,656]" fontSize="6" pageId="6" pageNumber="191">2</subScript>
) aspects.
<emphasis id="FF8AEAB8FF9E2450FD1AFDFE6C87FD57" bold="true" box="[674,689,633,652]" pageId="6" pageNumber="191">B</emphasis>
. Paratype MCCM LH 23851, dorsal (B
<subScript id="517A34EFFF9E2450FBA2FD056A14FD4B" attach="right" box="[1050,1058,642,656]" fontSize="6" pageId="6" pageNumber="191">1</subScript>
) and ventral (B
<subScript id="517A34EFFF9E2450FB09FD056A8FFD4B" attach="left" box="[1201,1209,642,656]" fontSize="6" pageId="6" pageNumber="191">2</subScript>
) aspects. Scale bars 2 mm.
</paragraph>
</caption>
<paragraph id="CD4136AAFF9E2450FFDEFC846FB8FC8B" blockId="6.[102,775,710,1281]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FFDEFC846F04FCC3" box="[102,306,771,792]" italics="true" pageId="6" pageNumber="191">Stratigraphic horizon</emphasis>
: Second lithosome of finely laminated limestones of La Huérguina Formation, Barremian, Lower Cretaceous (FregenalMartínez and Meléndez 2000).
</paragraph>
<paragraph id="CD4136AAFF9E2450FFDEFCDA6D34FCAA" blockId="6.[102,775,710,1281]" box="[102,770,860,882]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FFDEFCDA6F0AFCA9" box="[102,316,861,882]" italics="true" pageId="6" pageNumber="191">Derivation of the name</emphasis>
: Named after Armando Díaz Romeral (Cuenca).
</paragraph>
<paragraph id="CD4136AAFF9E2450FFDEFC056FA5FBE1" blockId="6.[102,775,710,1281]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FFDEFC056EFEFC41" box="[102,200,898,922]" italics="true" pageId="6" pageNumber="191">Material</emphasis>
.—
<materialsCitation id="7D963CF7FF9E2450FF50FC066FA5FBE1" collectionCode="MCCM" pageId="6" pageNumber="191" specimenCode="LH 12001, LH 23851, LH 20082" specimenCount="2" typeStatus="holotype">
<typeStatus id="12458808FF9E2450FF50FC066F64FC41" box="[232,338,897,922]" pageId="6" pageNumber="191" type="holotype">Holotype</typeStatus>
<collectionCode id="ABEFAE6FFF9E2450FEE6FC066F83FC41" box="[350,437,897,922]" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">MCCM</collectionCode>
<specimenCode id="9D589ED1FF9E2450FE07FC056C03FC41" box="[447,565,897,922]" collectionCode="MCCM" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">LH 12001</specimenCode>
part and counterpart, very well preserved specimen, lacking antennae and part of legs. From the same locality the
<typeStatus id="12458808FF9E2450FDA5FC456C4BFC01" box="[541,637,962,986]" pageId="6" pageNumber="191" type="paratype">paratype</typeStatus>
<collectionCode id="ABEFAE6FFF9E2450FD3BFC466CECFC01" box="[643,730,961,986]" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">MCCM</collectionCode>
<specimenCode id="9D589ED1FF9E2450FD59FC456E99FC21" collectionCode="MCCM" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">LH 23851</specimenCode>
, well preserved specimen without legs, and
<collectionCode id="ABEFAE6FFF9E2450FD17FC666D30FC21" box="[687,774,993,1018]" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">MCCM</collectionCode>
<specimenCode id="9D589ED1FF9E2450FFDEFB856EEDFBC1" box="[102,219,1025,1050]" collectionCode="MCCM" country="India" httpUri="http://grbio.org/cool/stc7-amit" name="Madras Christian College" pageId="6" pageNumber="191">LH 20082</specimenCode>
, only the part of an isolated elytra. All specimens from Las Hoyas fossil site.
</materialsCitation>
</paragraph>
<paragraph id="CD4136AAFF9E2450FFDEFBCD6D31FAF1" blockId="6.[102,775,710,1281]" lastBlockId="6.[102,776,1298,2026]" pageId="6" pageNumber="191">
<emphasis id="FF8AEAB8FF9E2450FFDEFBCD6EEEFBBA" box="[102,216,1098,1121]" italics="true" pageId="6" pageNumber="191">Diagnosis</emphasis>
.—From species with similar length of last ventrite and penultimate one (
<taxonomicName id="0AFE4D29FF9E2450FEE3FBEC6C8EFB59" authority="Ponomarenko, 2000" authorityName="Ponomarenko" authorityYear="2000" box="[347,696,1129,1154]" class="Insecta" family="Ommatidae" genus="Cionocoleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="sibirica">
<emphasis id="FF8AEAB8FF9E2450FEE3FBEC6FF8FB59" box="[347,462,1130,1154]" italics="true" pageId="6" pageNumber="191">Z. sibirica</emphasis>
Ponomarenko, 2000
</taxonomicName>
and
<taxonomicName id="0AFE4D29FF9E2450FD48FBEC6FF6FB79" authority="Ponomarenko, 1966" authorityName="Ponomarenko" authorityYear="1966" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="caudatus">
<emphasis id="FF8AEAB8FF9E2450FD48FBEC6EFDFB79" italics="true" pageId="6" pageNumber="191">Z. caudatus</emphasis>
<bibRefCitation id="A96F4B5BFF9E2450FF60FB0E6FF6FB79" author="Ponomarenko, A. G." box="[216,448,1161,1186]" pageId="6" pageNumber="191" pagination="138 - 143" refId="ref10028" refString="Ponomarenko, A. G. 1966. New beetles of the family Cupedidae (Coleoptera) from Mesozoic Deposits of Transbaikalia [in Russian]. Entomologiceskoe oboerenie 1: 138 - 143." type="journal article" year="1966">Ponomarenko, 1966</bibRefCitation>
</taxonomicName>
from Lower Cretaceous of Transbaikalia, and
<taxonomicName id="0AFE4D29FF9E2450FE81FB2C6CAEFB19" authority="Ponomarenko, 1969" authorityName="Ponomarenko" authorityYear="1969" box="[313,664,1193,1218]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="191" phylum="Arthropoda" rank="species" species="caducus">
<emphasis id="FF8AEAB8FF9E2450FE81FB2C6F84FB19" box="[313,434,1194,1218]" italics="true" pageId="6" pageNumber="191">Z. caducus</emphasis>
<bibRefCitation id="A96F4B5BFF9E2450FE00FB2E6CAEFB19" author="Ponomarenko, A. G." box="[440,664,1193,1218]" pageId="6" pageNumber="191" pagination="1 - 238" refId="ref10112" refString="Ponomarenko, A. G. 1969. Historical development of the Coleoptera-Archostemata [in Russian]. Trudy Paleontologiceskogo Instituta 125: 1 - 238." type="journal article" year="1969">Ponomarenko, 1969</bibRefCitation>
</taxonomicName>
from Upper Cretaceous of Mid Asia) differs in bigger size and wider pronotum and elytra.
<emphasis id="FF8AEAB8FF9E2450FFDEFA946ED1FAF1" box="[102,231,1299,1322]" italics="true" pageId="6" pageNumber="191">Description</emphasis>
.—Measurements: length
<quantity id="0A069B4FFF9E2450FDB7FA966C8EFAF2" box="[527,696,1297,1322]" metricMagnitude="-2" metricUnit="m" metricValue="2.335" metricValueMax="2.34" metricValueMin="2.33" pageId="6" pageNumber="191" unit="mm" value="23.35" valueMax="23.4" valueMin="23.3">23.323.4 mm</quantity>
; width
</paragraph>
<paragraph id="CD4136AAFF9E2450FFDEFAB66A69FC5B" blockId="6.[102,776,1298,2026]" lastBlockId="6.[812,1486,711,896]" pageId="6" pageNumber="191">
11.612.0 mm; elytra length
<quantity id="0A069B4FFF9E2450FE12FAB66C66FA92" box="[426,592,1329,1354]" metricMagnitude="-2" metricUnit="m" metricValue="1.8049999999999997" metricValueMax="1.8199999999999998" metricValueMin="1.7899999999999998" pageId="6" pageNumber="191" unit="mm" value="18.049999999999997" valueMax="18.2" valueMin="17.9">17.918.2 mm</quantity>
. Body large and flattened. Head transverse, about 1.6 times as wide as long without mandibles, narrowed before eyes; cheeks and temples strongly shorter than eyes. Antennae filiform, scape and pedicel more than two times longer and wider than rest of antennomeres. Necklike constriction weak. Pronotum transverse, about 1.8 times as wide as long, widest in its medial length, and narrowed roundly posterior and anteriorly; anterior angles rounded and not incised, lateral edges somewhat flattened, posterior angles distinct. Disc of pronotum with a large rectangular elevation divided by a longitudinal furrow. Prosternum somewhat longer than broader. Mesosternum nearly quadrate. Mesocoxae 1.7 times as long as wide, with exposed trochantin. Metasternum strongly transverse, about two times as wide as long. Abdomen with 5 visible ventrites with sharp relief, their posterior half elevated above anterior half. Last ventrite about 3.3 times as long as previous one. Elytra about three times as long as wide, its apex not drawn out into “tail”, epipleura comparatively wide, narrowing posteriorly, and with a row of distinct cells extending to apex. Main veins of elytra sharply distinct from intercalaries, 25 rounded cells form a row. Legs quite developed and elongate. Pro and mesofemora about three times as long as wide, somewhat longer than metafemora. Metatibia strongly elongate, about 1.5 times as long as metafemora. Metatarsi tarsomeres subequal in length, almost as long as scape. Abdomen covered with small tubercles, larger on last abdominal ventrite and depressed parts of preceding ventrites.
</paragraph>
</subSubSection>
</treatment>
</document>