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<document id="FAA1D9C917C073B81F39815538981D72" ID-DOI="10.5281/zenodo.13643699" ID-ISSN="1732-2421" ID-Zenodo-Dep="13643699" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142205826" checkinUser="felipe" docAuthor="Soriano, Carmen &amp; Delclòs, Xavier" docDate="2006" docId="455787BCFF992454FC94F9C66F41F880" docLanguage="en" docName="ActaPalaeontolPol.51.1.185-200.pdf" docOrigin="Acta Palaeontologica Polonica 51 (1)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Tetraphalerus penalveri Soriano &amp; Delclòs, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="187" masterDocId="B96EFFC4FF982456FFB8FF876E36FFDB" masterDocTitle="New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family" masterLastPageNumber="200" masterPageNumber="185" pageNumber="186" updateTime="1725365340826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BACADEE66A33E616C46722CC8BEFA34A">New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family</mods:title>
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<mods:namePart id="127E2FFA215E1D36B3C932935F3080BE">Soriano, Carmen</mods:namePart>
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<mods:namePart id="12421ED263AECAC9E64080380741E053">Delclòs, Xavier</mods:namePart>
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<mods:title id="FAC0FAC751E866CD356CEE442A91DFDC">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="2AD5B23DCA65FCFE1A77D6E31DC584D7">
<mods:date id="A2B18D0639D6964CB88461F3BBCC9046">2006</mods:date>
<mods:detail id="BE908572D6FBF752742F7B08E86D76FF" type="volume">
<mods:number id="2684E195EDC56975A7F118B7800EDD9E">51</mods:number>
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<mods:identifier id="3541E248D6E8370455E08E3CC312349D" type="DOI">10.5281/zenodo.13643699</mods:identifier>
<mods:identifier id="A47438AD6FE4A209E866A200B8678832" type="ISSN">1732-2421</mods:identifier>
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<treatment id="455787BCFF992454FC94F9C66F41F880" ID-DOI="http://doi.org/10.5281/zenodo.13643713" ID-Zenodo-Dep="13643713" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880" httpUri="http://treatment.plazi.org/id/455787BCFF992454FC94F9C66F41F880" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186">
<subSubSection id="85E46521FF992457FC94F9C66AF9F986" box="[812,1231,1601,1629]" pageId="1" pageNumber="186" type="nomenclature">
<paragraph id="CD4136AAFF992457FC94F9C66AF9F986" blockId="1.[812,1231,1601,1662]" box="[812,1231,1601,1629]" pageId="1" pageNumber="186">
<heading id="960981C6FF992457FC94F9C66AF9F986" box="[812,1231,1601,1629]" fontSize="12" level="2" pageId="1" pageNumber="186" reason="9">
<taxonomicName id="0AFE4D29FF992457FC94F9C66A55F986" authorityName="Soriano &amp; Delclòs" authorityYear="2006" box="[812,1123,1601,1629]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="186" phylum="Arthropoda" rank="species" species="penalveri" status="sp. nov.">
<emphasis id="FF8AEAB8FF992457FC94F9C66A55F986" box="[812,1123,1601,1629]" italics="true" pageId="1" pageNumber="186">Tetraphalerus penalveri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B957C3FF992457FBD2F9C66AF9F986" box="[1130,1231,1601,1629]" pageId="1" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="85E46521FF992454FC94F9EE6F41F880" lastPageId="2" lastPageNumber="187" pageId="1" pageNumber="186" type="description">
<paragraph id="CD4136AAFF992457FC94F9EE6D9AF9A5" blockId="1.[812,1231,1601,1662]" box="[812,940,1641,1662]" pageId="1" pageNumber="186">
<figureCitation id="55C52A2FFF992457FC94F9EE6D49F9A5" box="[812,895,1641,1662]" captionStart="Fig" captionStartId="2.[102,135,891,911]" captionTargetBox="[136,1459,230,863]" captionTargetId="figure-540@2.[124,1463,225,864]" captionTargetPageId="2" captionText="Fig. 1. Camera lucida drawings of Barremian (Early Cretaceous) ommatin Tetraphalerus from Spain. A. Tetraphalerus ponomarenkoi sp. nov. Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province, dorsal (A1) and ventral (A2) aspects. B. Tetraphalerus penalveri sp. nov. Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643701" httpUri="https://zenodo.org/record/13643701/files/figure.png" pageId="1" pageNumber="186">Figs. 1B</figureCitation>
,
<figureCitation id="55C52A2FFF992457FC32F9EE6D9EF9A5" box="[906,936,1641,1662]" captionStart="Fig" captionStartId="3.[102,134,1570,1590]" captionTargetBox="[206,1382,225,1543]" captionTargetId="figure-150@3.[206,1382,225,1543]" captionTargetPageId="3" captionText="Fig. 2. Barremian (Early Cretaceous) ommatins Tetraphalerus, Cionocoleus (both Ommatini), Brochocoleus (Brochocoleini), and Zygadenia (Notocupedini) from Spain. A. Tetraphalerus ponomarenkoi sp. nov. Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province. B. Tetraphalerus penalveri sp. nov. Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province. C. Cionocoleus longicapitis sp. nov. Holotype IEI LP 0164G, La Pedrera, Lleida Province. D. Brochocoleus indibili sp. nov. Holotype IEI LC 4636a, part, La Cabrúa, Lleida Province. E. Zygadenia viridis sp. nov. Holotype MCCM LH 1805a, part, Las Hoyas, Cuenca Province. F. Zygadenia oculata sp. nov. Holotype IEI LC 3675a, part, La Cabrúa, Lleida Province. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13643703" httpUri="https://zenodo.org/record/13643703/files/figure.png" pageId="1" pageNumber="186">2B</figureCitation>
.
</paragraph>
<paragraph id="CD4136AAFF992457FC94F9156A0FF918" blockId="1.[812,1485,1682,1886]" pageId="1" pageNumber="186">
<materialsCitation id="7D963CF7FF992457FC94F9156A03F918" country="Spain" county="Lleida Province" location="Quarry" municipality="Sierra del Montsec" pageId="1" pageNumber="186" specimenCount="1" typeStatus="holotype">
<emphasis id="FF8AEAB8FF992457FC94F9156D91F97C" box="[812,935,1682,1703]" italics="true" pageId="1" pageNumber="186">
<typeStatus id="12458808FF992457FC94F9156D6DF97C" box="[812,859,1682,1703]" pageId="1" pageNumber="186">Type</typeStatus>
locality
</emphasis>
:
<location id="C8216071FF992457FC0AF9156DC0F97C" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FC0AF9156DC0F97C" box="[946,1014,1682,1703]" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="Quarry" pageId="1" pageNumber="186">Quarry</location>
near
<location id="C8216071FF992457FB90F9156A5BF97C" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FB90F9156A5BF97C" box="[1064,1133,1682,1703]" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="Rubies" pageId="1" pageNumber="186">Rubies</location>
,
<location id="C8216071FF992457FBCDF9156B1EF97C" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FBCDF9156B1EF97C" box="[1141,1320,1682,1703]" country="Spain" county="Lleida Province" municipality="Sierra del Montsec" name="Sta. Maria de Meia" pageId="1" pageNumber="186">Sta. Maria de Meià</location>
,
<collectingMunicipality id="2D25ACD0FF992457FA8AF9156D7BF918" pageId="1" pageNumber="186">Sierra del Montsec</collectingMunicipality>
,
<collectingCounty id="24204E26FF992457FCEFF9296DC4F918" box="[855,1010,1710,1731]" pageId="1" pageNumber="186">Lleida Province</collectingCounty>
,
<collectingCountry id="B5E9763AFF992457FC43F9296A03F918" box="[1019,1077,1710,1731]" name="Spain" pageId="1" pageNumber="186">Spain</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="CD4136AAFF992457FC94F9556A29F8C4" blockId="1.[812,1485,1682,1886]" pageId="1" pageNumber="186">
<materialsCitation id="7D963CF7FF992457FC94F9556A2DF8C4" county="Lower Cretaceous" location="Calcaires" municipality="Barremian" pageId="1" pageNumber="186" specimenCount="1" typeStatus="holotype">
<emphasis id="FF8AEAB8FF992457FC94F9556D9AF93C" box="[812,940,1746,1767]" italics="true" pageId="1" pageNumber="186">
<typeStatus id="12458808FF992457FC94F9556D6DF93C" box="[812,859,1746,1767]" pageId="1" pageNumber="186">Type</typeStatus>
horizon
</emphasis>
:
<location id="C8216071FF992457FC04F9556A23F93C" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FC04F9556A23F93C" box="[956,1045,1746,1767]" county="Lower Cretaceous" municipality="Barremian" name="Calcaires" pageId="1" pageNumber="186">Calcaires</location>
lithographiques à
<location id="C8216071FF992457FB77F9556B22F93C" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FB77F9556B22F93C" box="[1231,1300,1746,1767]" county="Lower Cretaceous" municipality="Barremian" name="Plantes" pageId="1" pageNumber="186">Plantes</location>
et
<location id="C8216071FF992457FA8FF9556A3DF8D8" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FA8FF9556A3DF8D8" county="Lower Cretaceous" municipality="Barremian" name="Vertebres de la Pedrera de Rubies Fm." pageId="1" pageNumber="186">Vértébres de la Pedrera de Rubies Fm.</location>
,
<collectingMunicipality id="2D25ACD0FF992457FBADF9696A4BF8D8" box="[1045,1149,1774,1795]" pageId="1" pageNumber="186">Barremian</collectingMunicipality>
,
<collectingCounty id="24204E26FF992457FB3EF9696B00F8D8" box="[1158,1334,1774,1795]" pageId="1" pageNumber="186">Lower Cretaceous</collectingCounty>
(
<bibRefCitation id="A96F4B5BFF992457FAFAF9696A22F8C4" author="Martin-Closas, C. &amp; Lopez-Moron, N." pageId="1" pageNumber="186" pagination="29 - 31" refId="ref9705" refString="Martin-Closas, C. and Lopez-Moron, N. 1995. The Charophyte flora. In: X. Martinez-Delclos (ed.), Montsec and Montral-Alcover. Two Konservat- Lagerstatten, Catalonia, Spain. II International Symposium on Lithographic Limestones field trip guide book, 29 - 31. Institut d'Estudis Illerdens, Lleida." type="book chapter" year="1995">Martín−Closas and López−Morón 1995</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CD4136AAFF992457FC94F8AA6DBFF885" blockId="1.[812,1485,1682,1886]" pageId="1" pageNumber="186">
<emphasis id="FF8AEAB8FF992457FC94F8AA6A2AF899" box="[812,1052,1837,1858]" italics="true" pageId="1" pageNumber="186">Derivation of the name</emphasis>
: After the Spanish palaeoentomologist E. Peñalver.
</paragraph>
<paragraph id="CD4136AAFF992457FC94F8F56A98F831" blockId="1.[812,1485,1905,2026]" pageId="1" pageNumber="186">
<emphasis id="FF8AEAB8FF992457FC94F8F56DA6F851" box="[812,912,1906,1930]" italics="true" pageId="1" pageNumber="186">Material</emphasis>
.—
<materialsCitation id="7D963CF7FF992457FC0AF8F66A98F831" collectionCode="DEPGM" country="Spain" county="Exemplar" location="La Pedrera de Meia" municipality="Lleida" pageId="1" pageNumber="186" specimenCode="LP 0058" specimenCount="1" typeStatus="holotype">
<typeStatus id="12458808FF992457FC0AF8F66A29F851" box="[946,1055,1905,1930]" pageId="1" pageNumber="186" type="holotype">Holotype</typeStatus>
only (
<collectionCode id="ABEFAE6FFF992457FBCBF8F66AEEF851" box="[1139,1240,1905,1930]" pageId="1" pageNumber="186">DEPGM</collectionCode>
<specimenCode id="9D589ED1FF992457FB5DF8F56B66F851" box="[1253,1360,1905,1930]" collectionCode="DEPGM" pageId="1" pageNumber="186">LP 0058</specimenCode>
), part and counterpart, from
<location id="C8216071FF992457FBBEF8156AC3F871" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FBBEF8156AC3F871" box="[1030,1269,1937,1962]" country="Spain" county="Exemplar" municipality="Lleida" name="La Pedrera de Meia" pageId="1" pageNumber="186">La Pedrera de Meià</location>
fossil site, in the
<location id="C8216071FF992457FC94F8356DD6F811" LSID="urn:lsid:plazi:treatment:455787BCFF992454FC94F9C66F41F880:C8216071FF992457FC94F8356DD6F811" box="[812,992,1970,1994]" country="Spain" county="Exemplar" municipality="Lleida" name="Montsec Range" pageId="1" pageNumber="186">Montsec Range</location>
(
<collectingMunicipality id="2D25ACD0FF992457FC48F8366A08F811" box="[1008,1086,1969,1994]" pageId="1" pageNumber="186">Lleida</collectingMunicipality>
,
<collectingCountry id="B5E9763AFF992457FBF3F8366AA7F811" box="[1099,1169,1969,1994]" name="Spain" pageId="1" pageNumber="186">Spain</collectingCountry>
).
<collectingCounty id="24204E26FF992457FB1DF8366B22F811" box="[1189,1300,1969,1994]" pageId="1" pageNumber="186">Exemplar</collectingCounty>
well preserved, lacking part of antennae and legs.
</materialsCitation>
</paragraph>
<caption id="99816622FF9A2454FFDEFCFC6DFAFC1C" ID-DOI="http://doi.org/10.5281/zenodo.13643701" ID-Zenodo-Dep="13643701" httpUri="https://zenodo.org/record/13643701/files/figure.png" pageId="2" pageNumber="187" startId="2.[102,135,891,911]" targetBox="[136,1459,230,863]" targetPageId="2" targetType="figure">
<paragraph id="CD4136AAFF9A2454FFDEFCFC6DFAFC1C" blockId="2.[102,1485,891,971]" pageId="2" pageNumber="187">
Fig. 1. Camera lucida drawings of Barremian (Early Cretaceous) ommatin
<taxonomicName id="0AFE4D29FF9A2454FCB3FCFB6DBDFC4B" authorityName="Waterhouse" authorityYear="1901" box="[779,907,892,912]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="genus">
<emphasis id="FF8AEAB8FF9A2454FCB3FCFB6DBDFC4B" box="[779,907,892,912]" italics="true" pageId="2" pageNumber="187">Tetraphalerus</emphasis>
</taxonomicName>
from Spain.
<emphasis id="FF8AEAB8FF9A2454FBBAFCFC6A24FC54" bold="true" box="[1026,1042,891,911]" pageId="2" pageNumber="187">A</emphasis>
.
<taxonomicName id="0AFE4D29FF9A2454FBA5FCFB6B10FC4B" authorityName="Soriano &amp; Delclòs" authorityYear="2006" box="[1053,1318,892,912]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="ponomarenkoi" status="sp. nov.">
<emphasis id="FF8AEAB8FF9A2454FBA5FCFB6B10FC4B" box="[1053,1318,892,912]" italics="true" pageId="2" pageNumber="187">Tetraphalerus ponomarenkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B957C3FF9A2454FA94FCFB6B42FC54" box="[1324,1396,892,911]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province, dorsal (A
<subScript id="517A34EFFF9A2454FD34FC266CA2FC74" attach="left" box="[652,660,929,943]" fontSize="6" pageId="2" pageNumber="187">1</subScript>
) and ventral (A
<subScript id="517A34EFFF9A2454FC99FC266D1FFC74" attach="right" box="[801,809,929,943]" fontSize="6" pageId="2" pageNumber="187">2</subScript>
) aspects.
<emphasis id="FF8AEAB8FF9A2454FC38FC1F6DB9FC70" bold="true" box="[896,911,920,939]" pageId="2" pageNumber="187">B</emphasis>
.
<taxonomicName id="0AFE4D29FF9A2454FC22FC1F6A45FC77" authorityName="Soriano &amp; Delclòs" authorityYear="2006" box="[922,1139,920,940]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="penalveri" status="sp. nov.">
<emphasis id="FF8AEAB8FF9A2454FC22FC1F6A45FC77" box="[922,1139,920,940]" italics="true" pageId="2" pageNumber="187">Tetraphalerus penalveri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4B957C3FF9A2454FBC0FC1F6A89FC70" box="[1144,1215,920,939]" pageId="2" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province, dorsal (B
<subScript id="517A34EFFF9A2454FD8EFC3A6C08FC10" attach="right" box="[566,574,957,971]" fontSize="6" pageId="2" pageNumber="187">1</subScript>
) and ventral (B
<subScript id="517A34EFFF9A2454FD74FC3A6CE2FC10" attach="left" box="[716,724,957,971]" fontSize="6" pageId="2" pageNumber="187">2</subScript>
) aspects. Scale bars 2 mm.
</paragraph>
</caption>
<paragraph id="CD4136AAFF9A2454FFDEFC756FA7FB56" blockId="2.[102,775,1008,1165]" pageId="2" pageNumber="187">
<emphasis id="FF8AEAB8FF9A2454FFDEFC756EEEFBD2" box="[102,216,1010,1033]" italics="true" pageId="2" pageNumber="187">Diagnosis</emphasis>
.—One of the smaller species of the genus, from the
<taxonomicName id="0AFE4D29FF9A2454FF28FB956F49FBF1" authorityName="Heller" authorityYear="1913" box="[144,383,1042,1066]" class="Insecta" family="Ommatidae" genus="Tetraphalerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="187" phylum="Arthropoda" rank="species" species="bruchi">
<emphasis id="FF8AEAB8FF9A2454FF28FB956F49FBF1" box="[144,383,1042,1066]" italics="true" pageId="2" pageNumber="187">Tetraphalerus bruchi</emphasis>
</taxonomicName>
series (presence of two prominences in the vertex). From the smallest species of this series with similar ratio between last and penultimate ventrites, differs in the transverse head.
</paragraph>
<paragraph id="CD4136AAFF9A2454FFDEFB236F41F880" blockId="2.[102,775,1186,1883]" pageId="2" pageNumber="187">
<emphasis id="FF8AEAB8FF9A2454FFDEFB236ED1FB60" box="[102,231,1188,1211]" italics="true" pageId="2" pageNumber="187">Description</emphasis>
.—Measurements: length
<quantity id="0A069B4FFF9A2454FDBBFB256C6DFB61" box="[515,603,1186,1211]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="187" unit="mm" value="6.6">6.6 mm</quantity>
; width
<quantity id="0A069B4FFF9A2454FD13FB256D35FB61" box="[683,771,1186,1211]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="187" unit="mm" value="2.6">2.6 mm</quantity>
; elytra length
<quantity id="0A069B4FFF9A2454FF40FB456F67FB01" box="[248,337,1218,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="187" unit="mm" value="3.8">3.8 mm</quantity>
. Beetle small and flat. Head transverse, about 1.3 times as wide as long, narrowed roundly after eyes; temples as long as eyes, not projected laterally, vertex with two flat oval prominences, with posterior edge widened. Eyes comparatively big, about as long as mandibles. Scape 1.8 times longer than pedicel, equal in length to antennomeres 3–5, somewhat as wide as long. Neck−like constriction clear, two times shorter than the occipital part of head. Pronotum 1.5 times longer than wider, lateral edges flattened, anterior and posterior angles rounded; pronotal disc bearing two flat oval prominences. Procoxae rounded and contiguous. Mesosternum transverse, about 1.4 times wider than longer. Mesocoxae 1.4 times longer than wider, trochantin elongated. Metasternum nearly quadrate, almost as long as wide at its posterior edge, with longitudinal and transverse sutures clearly expressed. Metacoxae sub−triangular, roundly narrowed laterally. Abdomen with 5 visible ventrites, last ventrite about 1.6 times as long as previous one. Elytra about 3.5 times as long as wide, epipleural rim comparatively narrow. Pro− and mesofemora somewhat longer than metafemora. Body covered with shallow puntuation.
</paragraph>
</subSubSection>
</treatment>
</document>
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