treatments-xml/data/31/42/87/314287B0F625FFB4FF7BFC6708D8F9C3.xml

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<mods:title>A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>TONG, JUNBO</mods:namePart>
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<mods:namePart>CHEN, XIAOSHENG</mods:namePart>
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<mods:namePart>ZHANG, XIAONING</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>HUO, LIZHI</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>WU, GANG</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>PENG, ZHENGQIANG</mods:namePart>
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<mods:namePart>WANG, XINGMIN</mods:namePart>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-11-14</mods:number>
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<subSubSection box="[151,589,901,928]" pageId="20" pageNumber="21" type="nomenclature">
<paragraph blockId="20.[151,589,901,928]" box="[151,589,901,928]" pageId="20" pageNumber="21">
<heading bold="true" box="[151,589,901,928]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis bold="true" box="[151,589,901,928]" pageId="20" pageNumber="21">
<taxonomicName baseAuthorityName="Weise" baseAuthorityYear="1879" box="[151,414,901,928]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hareja">
<emphasis bold="true" box="[151,414,901,928]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus hareja</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
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<subSubSection pageId="20" pageNumber="21" type="discussion">
<paragraph blockId="20.[151,1437,973,1144]" pageId="20" pageNumber="21">
The
<taxonomicName baseAuthorityName="Weise" baseAuthorityYear="1879" box="[202,445,973,1000]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hareja">
<emphasis box="[202,445,973,1000]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus hareja</emphasis>
</taxonomicName>
species group is characterized by the stout penis with large penis capsule, apex of penis with membranous appendage, penis guide robust with prominent keel at base, interocular distance distinctly greater than eye width in frontal view, coxites elongate and triangular, styli well developed with terminal setae, and spermathecal with elongate nodulus.
</paragraph>
<paragraph blockId="20.[151,1437,973,1144]" box="[199,1419,1117,1144]" pageId="20" pageNumber="21">
34 species of
<taxonomicName baseAuthorityName="Weise" baseAuthorityYear="1879" box="[352,424,1117,1143]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hareja">
<emphasis box="[352,424,1117,1143]" italics="true" pageId="20" pageNumber="21">hareja</emphasis>
</taxonomicName>
group are recorded from
<collectingCountry box="[711,781,1117,1143]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
, including 12 new species and 1 newly recorded species.
</paragraph>
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<paragraph blockId="20.[151,1203,1225,1252]" box="[151,1203,1225,1252]" pageId="20" pageNumber="21">
<heading bold="true" box="[151,1203,1225,1252]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis bold="true" box="[151,1203,1225,1252]" pageId="20" pageNumber="21">
Key to the species of
<taxonomicName baseAuthorityName="Weise" baseAuthorityYear="1879" box="[413,491,1225,1252]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="hareja">
<emphasis bold="true" box="[413,491,1225,1252]" italics="true" pageId="20" pageNumber="21">hareja</emphasis>
</taxonomicName>
group within the genus
<taxonomicName authority="Vandenberg, 2004" authorityName="Vandenberg" authorityYear="2004" box="[793,1203,1225,1251]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[793,971,1225,1251]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus</emphasis>
<bibRefCitation author="Vandenberg, N. J." box="[977,1203,1225,1251]" pageId="20" pageNumber="21" pagination="483 - 484" refId="ref65879" refString="Vandenberg, N. J. (2004) Homonymy in the Coccinellidae (Coleoptera), or something fishy about Pseudoscymnus Chapin. Proceedings of the Entomological Society of Washington, 106 (2), 483 - 484." type="journal article" year="2004">Vandenberg, 2004</bibRefCitation>
</taxonomicName>
</emphasis>
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<key lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1294,1316]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1294,1316]" pageId="20" pageNumber="21">1. Parameres distinct..................................................................................... 2</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1437,1323,1346]" pageId="20" pageNumber="21">
<keyLead box="[151,1437,1323,1346]" pageId="20" pageNumber="21">
- Parameres inconspicuous (
<figureCitation box="[482,588,1324,1346]" captionStart="FIGURE 13" captionStartId="22.[151,250,1526,1550]" captionTargetBox="[151,1436,597,1503]" captionTargetId="figure-179@22.[151,1436,597,1503]" captionTargetPageId="22" captionText="FIGURE 13. S. gibbosus (Yu, 1997), holotype: a dorsal view, b lateral view, c frontal view, d abdomen, e–g male genitalia: e basal part of penis, f tegmen, ventral view, g tegmen, lateral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328114" httpUri="https://zenodo.org/record/7328114/files/figure.png" pageId="20" pageNumber="21">Figs 13f–g</figureCitation>
).........................................
<taxonomicName authority="(Yu, 1997)" baseAuthorityName="Yu" baseAuthorityYear="1997" box="[1088,1437,1323,1345]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis bold="true" box="[1088,1437,1323,1345]" pageId="20" pageNumber="21">
<emphasis bold="true" box="[1088,1326,1323,1345]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus gibbosus</emphasis>
(Yu, 1997)
</emphasis>
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<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1353,1375]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1353,1375]" pageId="20" pageNumber="21">2. Central part of penis smooth without small protuberances..................................................... 3</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1437,1382,1404]" pageId="20" pageNumber="21">
<keyLead box="[151,1437,1382,1404]" pageId="20" pageNumber="21">
- Central part of penis with several small protuberances (Fig. 14e)...............
<taxonomicName authority="(Pang, 1993)" baseAuthorityName="Pang" baseAuthorityYear="1993" box="[1021,1437,1382,1404]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="shixingiensis">
<emphasis bold="true" box="[1021,1437,1382,1404]" pageId="20" pageNumber="21">
<emphasis bold="true" box="[1021,1302,1382,1404]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus shixingiensis</emphasis>
(
<bibRefCitation author="Pang, H." box="[1315,1429,1382,1404]" pageId="20" pageNumber="21" pagination="493 - 494" refId="ref65161" refString="Pang, H. (1993) A new species of Pseudoscymnus (Coleoptera: Coccinellidae). Acta Entomologica Sinica, 36 (4), 493 - 494. [in Chinese with English summary]" type="journal article" year="1993">Pang, 1993</bibRefCitation>
)
</emphasis>
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<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1412,1434]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1412,1434]" pageId="20" pageNumber="21">3. Penis uniform without constricted part..................................................................... 4</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1441,1463]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1441,1463]" pageId="20" pageNumber="21">
- Penis strongly constricted near distal 1/3 (
<figureCitation box="[629,709,1441,1463]" captionStart="FIGURE 15" captionStartId="24.[151,250,1938,1962]" captionTargetBox="[160,1427,851,1915]" captionTargetId="figure-244@24.[151,1436,849,1915]" captionTargetPageId="24" captionText="FIGURE 15. S. striatus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328116" httpUri="https://zenodo.org/record/7328116/files/figure.png" pageId="20" pageNumber="21">Fig. 15e</figureCitation>
).....................
<emphasis bold="true" box="[980,1436,1441,1463]" pageId="20" pageNumber="21">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[980,1353,1441,1463]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis bold="true" box="[980,1205,1441,1462]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus striatus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1441,1462]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1470,1492]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1470,1492]" pageId="20" pageNumber="21">4. Penis with a large protuberance on inner side............................................................... 5</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1500,1522]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1500,1522]" pageId="20" pageNumber="21">- Penis with smooth inner side............................................................................ 6</keyLead>
</paragraph>
</keyStep>
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<paragraph blockId="20.[151,1437,1294,2020]" pageId="20" pageNumber="21">
<keyLead pageId="20" pageNumber="21">
5. Pronotum entirely yellow (Fig. 16c); parameres about 1/3 as long as penis guide (Fig.
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)......................................................................................
<emphasis bold="true" box="[925,1436,1558,1580]" pageId="20" pageNumber="21">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[925,1353,1558,1580]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="wuliangshan" status="sp. nov.">
<emphasis bold="true" box="[925,1205,1558,1580]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus wuliangshan</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1558,1579]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
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<paragraph blockId="20.[151,1437,1294,2020]" pageId="20" pageNumber="21">
<keyLead pageId="20" pageNumber="21">
- Pronotum black with yellow margins (
<figureCitation box="[594,677,1588,1610]" captionStart="FIGURE 17" captionStartId="27.[151,250,1250,1274]" captionTargetBox="[165,1426,168,1233]" captionTargetId="figure-18@27.[151,1436,166,1233]" captionTargetPageId="27" captionText="FIGURE 17. S. ocelliferus (Canepari, 1997): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328120" httpUri="https://zenodo.org/record/7328120/files/figure.png" pageId="20" pageNumber="21">Fig. 17c</figureCitation>
); parameres about 3/5 as long as penis guide (
<figureCitation box="[1119,1201,1588,1610]" captionStart="FIGURE 17" captionStartId="27.[151,250,1250,1274]" captionTargetBox="[165,1426,168,1233]" captionTargetId="figure-18@27.[151,1436,166,1233]" captionTargetPageId="27" captionText="FIGURE 17. S. ocelliferus (Canepari, 1997): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328120" httpUri="https://zenodo.org/record/7328120/files/figure.png" pageId="20" pageNumber="21">
Fig.
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</figureCitation>
)..................................................................................
<taxonomicName authority="(Canepari, 1997)" baseAuthorityName="Canepari" baseAuthorityYear="1997" box="[1003,1437,1616,1639]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ocelliferus">
<emphasis bold="true" box="[1003,1437,1616,1639]" pageId="20" pageNumber="21">
<emphasis bold="true" box="[1003,1258,1616,1638]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus ocelliferus</emphasis>
(
<bibRefCitation author="Canepari, C." box="[1271,1429,1617,1639]" pageId="20" pageNumber="21" pagination="1 - 65" refId="ref63702" refString="Canepari, C. (1997) Coccinellidae (Coleoptera) from the Nepal Himalayas. Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), 565 (65), 1 - 65." type="journal article" year="1997">Canepari, 1997</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1646,1668]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1646,1668]" pageId="20" pageNumber="21">6. Elytron black with an eye-spotted pattern.................................................................. 7</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1676,1698]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1676,1698]" pageId="20" pageNumber="21">- Elytron without an eye-spotted pattern..................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1704,1727]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1704,1727]" pageId="20" pageNumber="21">
7. Yellow area of the eye spot extending to the apex of elytra (
<figureCitation box="[798,907,1705,1727]" captionStart="FIGURE 18" captionStartId="28.[151,250,1415,1439]" captionTargetBox="[151,1436,561,1392]" captionTargetId="figure-183@28.[151,1436,561,1392]" captionTargetPageId="28" captionText="FIGURE 18. S. orbiculatus (Yang, 1971): a dorsal view, paratype, b lateral view, paratype, c frontal view, paratype, d abdomen, holotype, e penis, holotype, f apex of penis, holotype, g tegmen, holotype, h labels, paratype." figureDoi="http://doi.org/10.5281/zenodo.7328122" httpUri="https://zenodo.org/record/7328122/files/figure.png" pageId="20" pageNumber="21">Figs 18a–b</figureCitation>
)..........
<taxonomicName authority="(Yang, 1971)" baseAuthorityName="Yang" baseAuthorityYear="1971" box="[1039,1436,1704,1727]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="orbiculatus">
<emphasis bold="true" box="[1039,1436,1704,1727]" pageId="20" pageNumber="21">
<emphasis bold="true" box="[1039,1302,1704,1726]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus orbiculatus</emphasis>
(
<bibRefCitation author="Yang, C. T." box="[1315,1429,1705,1727]" pageId="20" pageNumber="21" pagination="85 - 96" refId="ref66136" refString="Yang, C. T. (1971) Notes on the species of genus Pseudoscymnus from Taiwan. Journal of Agriculture and Forestry, 20: 85 - 96." type="journal article" year="1971">Yang, 1971</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1437,1734,1756]" pageId="20" pageNumber="21">
<keyLead box="[151,1437,1734,1756]" pageId="20" pageNumber="21">
- Yellow area of the eye spot not extending to the apex of elytra (
<figureCitation box="[835,944,1734,1756]" captionStart="FIGURE 19" captionStartId="29.[151,250,1904,1928]" captionTargetBox="[158,1427,816,1876]" captionTargetId="figure-262@29.[151,1436,813,1880]" captionTargetPageId="29" captionText="FIGURE 19. S. ocellatus (Yu, 2000): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–g male genitalia: e penis, f tegmen, lateral view, g tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–g: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328128" httpUri="https://zenodo.org/record/7328128/files/figure.png" pageId="20" pageNumber="21">Figs 19a–b</figureCitation>
)...........
<taxonomicName authority="(Yu, 2000)" baseAuthorityName="Yu" baseAuthorityYear="2000" box="[1086,1437,1734,1756]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ocellatus">
<emphasis bold="true" box="[1086,1437,1734,1756]" pageId="20" pageNumber="21">
<emphasis bold="true" box="[1086,1326,1734,1756]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus ocellatus</emphasis>
(
<bibRefCitation author="Yu, G. Y. &amp; Montgomery, M. E. &amp; Yao, D. F." box="[1338,1429,1734,1756]" pageId="20" pageNumber="21" pagination="154 - 199" refId="ref66379" refString="Yu, G. Y., Montgomery, M. E. &amp; Yao, D. F. (2000) Lady beetles (Coleoptera: Coccinellidae) from Chinese hemlocks infected with the hemlock woolly adelgid, Adelges tsugae Annand (Homoptera: Adelgidae). The Coleopterists Bulletin, 54 (2), 154 - 199. https: // doi. org / 10.1649 / 0010 - 065 X (2000) 054 [0154: LBCCFC] 2.0. CO; 2" type="journal article" year="2000">Yu, 2000</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1764,1786]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1764,1786]" pageId="20" pageNumber="21">8. Ground color of elytra uniformly yellow or brown........................................................... 9</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1793,1815]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1793,1815]" pageId="20" pageNumber="21">- Ground color of elytra black with yellow marking or yellow with black marking.................................. 11</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1822,1844]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1822,1844]" pageId="20" pageNumber="21">9. Ventral surfaces yellow to black......................................................................... 10</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1437,1851,1874]" pageId="20" pageNumber="21">
<keyLead box="[151,1437,1851,1874]" pageId="20" pageNumber="21">
- Ventral surfaces yellow without any contrasting markings (Figs 20a–c).....
<taxonomicName authority="(Pang &amp; Huang, 1986)" baseAuthorityName="Pang &amp; Huang" baseAuthorityYear="1986" box="[971,1437,1851,1873]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="paltatus">
<emphasis bold="true" box="[971,1437,1851,1873]" pageId="20" pageNumber="21">
<emphasis bold="true" box="[971,1201,1851,1873]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus paltatus</emphasis>
(
<bibRefCitation author="Pang, X. F. &amp; Huang, B. K." box="[1213,1429,1851,1873]" pageId="20" pageNumber="21" pagination="61 - 73" refId="ref65289" refString="Pang, X. F. &amp; Huang, B. K. (1986) Six new species of Scymnus and four new species of Pseudoscymnus from Fujian (Coleoptera: Coccinellidae: Scymnini). Journal of Wuyi Science, 6, 61 - 73. [in Chinese with English summary]" type="journal article" year="1986">Pang &amp; Huang, 1986</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1881,1903]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1881,1903]" pageId="20" pageNumber="21">
10. Elytra uniformly yellow (
<figureCitation box="[471,551,1881,1903]" captionStart="FIGURE 21" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[163,1428,816,1883]" captionTargetId="figure-256@31.[151,1436,813,1884]" captionTargetPageId="31" captionText="FIGURE 21. S. aureus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genita-" figureDoi="http://doi.org/10.5281/zenodo.7328132" httpUri="https://zenodo.org/record/7328132/files/figure.png" pageId="20" pageNumber="21">Fig. 21a</figureCitation>
)....................................
<emphasis bold="true" box="[986,1436,1881,1903]" pageId="20" pageNumber="21">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[986,1353,1881,1903]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="aureus" status="sp. nov.">
<emphasis bold="true" box="[986,1205,1881,1902]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus aureus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1881,1902]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1910,1932]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1910,1932]" pageId="20" pageNumber="21">
- Elytra uniformly brown (
<figureCitation box="[467,548,1910,1932]" captionStart="FIGURE 22" captionStartId="32.[151,250,1763,1787]" captionTargetBox="[151,1437,1012,1739]" captionTargetId="figure-245@32.[151,1437,1012,1739]" captionTargetPageId="32" captionText="FIGURE 22. S. fuscus (Yang, 1971): a dorsal view, paratype, b ventral view, paratype, c lateral view, paratype, d frontal view, paratype, e abdomen, holotype, f penis, holotype, g apex of penis, holotype, h tegmen, ventral view, holotype, i labels, paratype." figureDoi="http://doi.org/10.5281/zenodo.7328134" httpUri="https://zenodo.org/record/7328134/files/figure.png" pageId="20" pageNumber="21">Fig. 22a</figureCitation>
)............................................
<taxonomicName authority="(Yang, 1971)" baseAuthorityName="Yang" baseAuthorityYear="1971" box="[1087,1436,1910,1932]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis bold="true" box="[1087,1436,1910,1932]" pageId="20" pageNumber="21">
<emphasis bold="true" box="[1087,1302,1910,1932]" italics="true" pageId="20" pageNumber="21">Sasajiscymnus fuscus</emphasis>
(
<bibRefCitation author="Yang, C. T." box="[1315,1429,1910,1932]" pageId="20" pageNumber="21" pagination="85 - 96" refId="ref66136" refString="Yang, C. T. (1971) Notes on the species of genus Pseudoscymnus from Taiwan. Journal of Agriculture and Forestry, 20: 85 - 96." type="journal article" year="1971">Yang, 1971</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="20" pageNumber="21">
<footnote box="[151,1436,1940,1962]" pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1940,1962]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1940,1962]" pageId="20" pageNumber="21">11. Parameres distinctly dilated in lateral view, with widest part about twice of basal width............................. 12</keyLead>
</paragraph>
</footnote>
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1969,1991]" pageId="20" pageNumber="21">
<pageTitle box="[151,1436,1969,1991]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1969,1991]" pageId="20" pageNumber="21">- Parameres slightly dilated or not........................................................................ 14</keyLead>
</pageTitle>
</paragraph>
</keyStep>
<keyStep lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<paragraph blockId="20.[151,1437,1294,2020]" box="[151,1436,1998,2020]" pageId="20" pageNumber="21">
<pageTitle box="[151,1436,1998,2020]" pageId="20" pageNumber="21">
<keyLead box="[151,1436,1998,2020]" pageId="20" pageNumber="21">12. Parameres shorter than 3/4 length of penis guide............................................................ 13</keyLead>
</pageTitle>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,151,173]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,151,173]" pageId="21" pageNumber="22">
- Parameres slightly shorter than penis guide (
<figureCitation box="[653,735,151,173]" captionStart="FIGURE 23" captionStartId="33.[151,250,1927,1951]" captionTargetBox="[164,1436,834,1904]" captionTargetId="figure-190@33.[151,1437,833,1904]" captionTargetPageId="33" captionText="FIGURE 23. S. parenthesis (Weise, 1923): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e" figureDoi="http://doi.org/10.5281/zenodo.7328138" httpUri="https://zenodo.org/record/7328138/files/figure.png" pageId="21" pageNumber="22">
Fig.
<quantity box="[698,735,152,173]" metricMagnitude="-2" metricUnit="kg" metricValue="2.3" pageId="21" pageNumber="22" unit="g" value="23.0">23g</quantity>
</figureCitation>
).......................
<taxonomicName authority="(Weise, 1923)" baseAuthorityName="Weise" baseAuthorityYear="1923" box="[1030,1437,151,173]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="parenthesis">
<emphasis bold="true" box="[1030,1437,151,173]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[1030,1294,151,173]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus parenthesis</emphasis>
(
<bibRefCitation author="Weise, J." box="[1307,1429,151,173]" pageId="21" pageNumber="22" pagination="182 - 188" refId="ref66064" refString="Weise, J. (1923) Sauter's H. Formosan-Ausbeute: Coccinellidae. Archiv fur Naturgeschichte, 89, 182 - 188." type="journal article" year="1923">Weise, 1923</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,180,203]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,180,203]" pageId="21" pageNumber="22">
13. Parameres are about 1/2 as long as penis guide (
<figureCitation box="[687,769,181,203]" captionStart="FIGURE 24" captionStartId="35.[151,250,1272,1296]" captionTargetBox="[151,1430,182,1247]" captionTargetId="figure-18@35.[151,1436,181,1248]" captionTargetPageId="35" captionText="FIGURE 24. S. longus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328140" httpUri="https://zenodo.org/record/7328140/files/figure.png" pageId="21" pageNumber="22">
Fig.
<quantity box="[732,769,181,202]" metricMagnitude="-2" metricUnit="kg" metricValue="2.4" pageId="21" pageNumber="22" unit="g" value="24.0">24g</quantity>
</figureCitation>
).................
<emphasis bold="true" box="[987,1436,180,203]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[987,1353,180,203]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="longus" status="sp. nov.">
<emphasis bold="true" box="[987,1205,180,202]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus longus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,181,202]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,210,232]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,210,232]" pageId="21" pageNumber="22">
- Parameres are about 3/4 as long as penis guide (
<figureCitation box="[687,757,210,232]" captionStart="FIGURE 2" captionStartId="6.[151,250,1902,1926]" captionTargetBox="[164,1414,181,1877]" captionTargetId="figure-29@6.[164,1423,181,1878]" captionTargetPageId="6" captionText="FIGURE 2. Main characters of Sasajiscymnus, S. quinquepunctatus (Weise, 1923): a head, frontal view, b mandible, left, dorsal view, c mandible, right, dorsal view, d labrum, e antenna, f maxilla, g labium, h prothorax, ventral view, i prosternum, ventral view, j hind leg, k abdomen, male, m–o male genitalia: m apex of penis, l penis, n tegmen, lateral view, o tegmen, ventral view. Scale bars: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.7328084" httpUri="https://zenodo.org/record/7328084/files/figure.png" pageId="21" pageNumber="22">Fig. 2n</figureCitation>
)................
<taxonomicName authority="(Weise, 1923)" baseAuthorityName="Weise" baseAuthorityYear="1923" box="[964,1437,210,232]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="quinquepunctatus">
<emphasis bold="true" box="[964,1437,210,232]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[964,1295,210,231]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus quinquepunctatus</emphasis>
(
<bibRefCitation author="Weise, J." box="[1307,1429,210,232]" pageId="21" pageNumber="22" pagination="182 - 188" refId="ref66064" refString="Weise, J. (1923) Sauter's H. Formosan-Ausbeute: Coccinellidae. Archiv fur Naturgeschichte, 89, 182 - 188." type="journal article" year="1923">Weise, 1923</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,239,261]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,239,261]" pageId="21" pageNumber="22">14. Parameres longer than 1/4 length of penis guide............................................................ 15</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,268,291]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,268,291]" pageId="21" pageNumber="22">
- Parameres about 1/4 as long as penis guide (
<figureCitation box="[652,728,269,291]" captionStart="FIGURE 26" captionStartId="39.[151,250,1725,1749]" captionTargetBox="[169,1418,183,1698]" captionTargetId="figure-18@39.[159,1429,181,1701]" captionTargetPageId="39" captionText="FIGURE 26. S. heijia (Yu &amp; Montgomery, 2000): a, d dorsal view, b, e lateral view, c, f frontal view, g abdomen, female, h–k male genitalia: h penis, i apex of penis, j tegmen, lateral view, k tegmen, ventral view. Scale bars: a–f: 0.5 mm; g–k: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328144" httpUri="https://zenodo.org/record/7328144/files/figure.png" pageId="21" pageNumber="22">Fig. 26j</figureCitation>
)..................
<taxonomicName authority="(Yu &amp; Montgomery, 2000)" baseAuthorityName="Yu &amp; Montgomery" baseAuthorityYear="2000" box="[954,1437,268,291]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="heijia">
<emphasis bold="true" box="[954,1437,268,291]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[954,1162,268,290]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus heijia</emphasis>
(
<bibRefCitation author="Yu, G. Y. &amp; Montgomery, M. E. &amp; Yao, D. F." box="[1174,1429,268,291]" pageId="21" pageNumber="22" pagination="154 - 199" refId="ref66379" refString="Yu, G. Y., Montgomery, M. E. &amp; Yao, D. F. (2000) Lady beetles (Coleoptera: Coccinellidae) from Chinese hemlocks infected with the hemlock woolly adelgid, Adelges tsugae Annand (Homoptera: Adelgidae). The Coleopterists Bulletin, 54 (2), 154 - 199. https: // doi. org / 10.1649 / 0010 - 065 X (2000) 054 [0154: LBCCFC] 2.0. CO; 2" type="journal article" year="2000">Yu &amp; Montgomery, 2000</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,298,320]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,298,320]" pageId="21" pageNumber="22">15. Penis guide widest at base or near the base in ventral view.................................................... 16</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,327,349]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,327,349]" pageId="21" pageNumber="22">
- Penis guide widest at 1/2 length in ventral view (Fig. 27h)..................
<emphasis bold="true" box="[1006,1436,327,349]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[1006,1353,327,349]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis bold="true" box="[1006,1205,327,349]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus latus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,328,349]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,357,379]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,357,379]" pageId="21" pageNumber="22">16. Parameres distinctly shorter than 1/2 length of penis guide.................................................... 17</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,386,408]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,386,408]" pageId="21" pageNumber="22">- Parameres as long as or much longer than 1/2 length of penis guide............................................. 24</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,415,437]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,415,437]" pageId="21" pageNumber="22">17. Scutellar shield yellow to yellowish brown................................................................ 18</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,444,467]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,444,467]" pageId="21" pageNumber="22">
- Scutellar shield black (
<figureCitation box="[446,526,445,467]" captionStart="FIGURE 28" captionStartId="42.[151,250,1666,1690]" captionTargetBox="[171,1413,599,1639]" captionTargetId="figure-203@42.[166,1422,597,1642]" captionTargetPageId="42" captionText="FIGURE 28. S. curvatus (Yu, 1999): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328146" httpUri="https://zenodo.org/record/7328146/files/figure.png" pageId="21" pageNumber="22">Fig. 28a</figureCitation>
)..............................................
<taxonomicName authority="(Yu, 1999)" baseAuthorityName="Yu" baseAuthorityYear="1999" box="[1089,1437,444,466]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="curvatus">
<emphasis bold="true" box="[1089,1437,444,466]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[1089,1325,445,466]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus curvatus</emphasis>
(
<bibRefCitation author="Yu, G. Y." box="[1338,1429,444,466]" pageId="21" pageNumber="22" pagination="17 - 20" refId="ref66261" refString="Yu, G. Y. (1999) New species of family Coccinellidae from Mt. Jigong in Henan (Coleoptera: Coccinellidae). In: Shen, X. C. &amp; Pei, H. C. (Eds.), The fauna and taxonomy of insects in Henan, Vol. 3, Insects of the Mountain Jigong Regions. China Agricultural Science and Technology Press, Beijing, pp. 17 - 20. [in Chinese with English summary]" type="book chapter" year="1999">Yu, 1999</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,474,496]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,474,496]" pageId="21" pageNumber="22">18. Humeral angles black................................................................................. 19</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,503,525]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,503,525]" pageId="21" pageNumber="22">- Humeral angles yellow to yellowish brown................................................................ 21</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,533,555]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,533,555]" pageId="21" pageNumber="22">19. Elytra black with yellow apices, and a large yellow spot presenting in the middle part.............................. 20</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" pageId="21" pageNumber="22">
<keyLead pageId="21" pageNumber="22">
- Elytra black with yellowish brown apices, sometimes with two brown spots close to suture (
<figureCitation box="[1110,1190,562,584]" captionStart="FIGURE 29" captionStartId="43.[151,250,1941,1965]" captionTargetBox="[161,1424,851,1917]" captionTargetId="figure-259@43.[151,1436,849,1918]" captionTargetPageId="43" captionText="FIGURE 29. S. ancistroides (Pang &amp; Huang, 1986): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328150" httpUri="https://zenodo.org/record/7328150/files/figure.png" pageId="21" pageNumber="22">Fig. 29a</figureCitation>
).............................................................................
<taxonomicName authority="(Pang &amp; Huang, 1986)" baseAuthorityName="Pang &amp; Huang" baseAuthorityYear="1986" box="[931,1437,591,613]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ancistroides">
<emphasis bold="true" box="[931,1437,591,613]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[931,1201,591,613]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus ancistroides</emphasis>
(
<bibRefCitation author="Pang, X. F. &amp; Huang, B. K." box="[1213,1429,591,613]" pageId="21" pageNumber="22" pagination="61 - 73" refId="ref65289" refString="Pang, X. F. &amp; Huang, B. K. (1986) Six new species of Scymnus and four new species of Pseudoscymnus from Fujian (Coleoptera: Coccinellidae: Scymnini). Journal of Wuyi Science, 6, 61 - 73. [in Chinese with English summary]" type="journal article" year="1986">Pang &amp; Huang, 1986</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" pageId="21" pageNumber="22">
<keyLead pageId="21" pageNumber="22">
20. Middle part of elytra with yellow spot extending to the lateral margins (
<figureCitation box="[913,1022,621,643]" captionStart="FIGURE 30" captionStartId="44.[151,250,1766,1790]" captionTargetBox="[176,1409,706,1741]" captionTargetId="figure-215@44.[170,1418,705,1742]" captionTargetPageId="44" captionText="FIGURE 30. S. obliquus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328152" httpUri="https://zenodo.org/record/7328152/files/figure.png" pageId="21" pageNumber="22">Figs 30a–b</figureCitation>
)..............................................................................................
<emphasis bold="true" box="[968,1436,650,672]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[968,1353,650,672]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="obliquus" status="sp. nov.">
<emphasis bold="true" box="[968,1205,650,672]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus obliquus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,650,671]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" pageId="21" pageNumber="22">
<keyLead pageId="21" pageNumber="22">
- Middle part of elytra with yellow spot not extending to the lateral margins (Figs 31a–b)..........................................................................................
<emphasis bold="true" box="[953,1436,708,731]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[953,1353,708,731]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="niganulus" status="sp. nov.">
<emphasis bold="true" box="[953,1205,708,730]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus niganulus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,709,730]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,738,760]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,738,760]" pageId="21" pageNumber="22">21. Elytral base entirely yellow............................................................................ 22</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,767,789]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,767,789]" pageId="21" pageNumber="22">- Elytral base mostly black.............................................................................. 23</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,796,819]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,796,819]" pageId="21" pageNumber="22">
22. Elytral outer margin black in the middle............................
<taxonomicName authority="(Yang &amp; Wu, 1972)" baseAuthorityName="Yang &amp; Wu" baseAuthorityYear="1972" box="[949,1437,796,819]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fulvihumerus">
<emphasis bold="true" box="[949,1437,796,819]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[949,1233,796,818]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus fulvihumerus</emphasis>
(
<bibRefCitation author="Yang, C. T. &amp; Wu, R. H." box="[1246,1429,796,819]" pageId="21" pageNumber="22" pagination="115 - 128" refId="ref66088" refString="Yang, C. T. &amp; Wu, R. H. (1972) Notes on some Coccinellidae of Taiwan. Journal of Agriculture and Forestry, 21, 115 - 128. https: // doi. org / 10.2337 / diab. 21.2.128 b" type="journal article" year="1972">Yang &amp; Wu, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,826,848]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,826,848]" pageId="21" pageNumber="22">
- Elytral outer margin entirely yellow (Figs 33a–b).....................
<emphasis bold="true" box="[951,1436,826,848]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[951,1353,826,848]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis bold="true" box="[951,1206,826,848]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus maculatus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,826,847]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" pageId="21" pageNumber="22">
<keyLead pageId="21" pageNumber="22">
23. Elytra black with yellow apices and humeral angles, and a large yellowish brown spot presenting in the middle part (
<figureCitation captionStart="FIGURE 34" captionStartId="49.[151,250,1782,1806]" captionTargetBox="[157,1429,239,1747]" captionTargetId="figure-52@49.[151,1436,236,1758]" captionTargetPageId="49" captionText="FIGURE 34. S. seboshii (Ohta, 1929): a, d dorsal view, b, e lateral view, c, f frontal view, g abdomen, male, h–k male genitalia: h penis, i apex of penis, j tegmen, lateral view, k tegmen, ventral view. Scale bars: a–f: 0.5 mm; g–k: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328158" httpUri="https://zenodo.org/record/7328158/files/figure.png" pageId="21" pageNumber="22">Figs 34a–b</figureCitation>
).................................................................
<taxonomicName authority="(Ohta, 1929)" baseAuthorityName="Ohta" baseAuthorityYear="1929" box="[1071,1437,884,906]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="seboshii">
<emphasis bold="true" box="[1071,1437,884,906]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[1071,1302,884,906]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus seboshii</emphasis>
(
<bibRefCitation author="Ohta, Y." box="[1315,1429,884,906]" pageId="21" pageNumber="22" pagination="1 - 16" refId="ref65137" refString="Ohta, Y. (1929) Scymninen Japans. Insecta Matsumurana, 4 (1 - 2), 1 - 16." type="journal article" year="1929">Ohta, 1929</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,914,936]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,914,936]" pageId="21" pageNumber="22">
- Elytra black with yellowish brown apices and humeral angles (Figs 35a–b).........
<taxonomicName authority="(Yu, 1997)" baseAuthorityName="Yu" baseAuthorityYear="1997" box="[1060,1437,914,936]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="truncatulus">
<emphasis bold="true" box="[1060,1437,914,936]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[1060,1326,914,936]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus truncatulus</emphasis>
(Yu, 1997)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,943,965]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,943,965]" pageId="21" pageNumber="22">24. Penis with a deep terminal emargination.................................................................. 25</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,973,995]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,973,995]" pageId="21" pageNumber="22">- Penis without emarginations............................................................................ 26</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,1002,1024]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,1002,1024]" pageId="21" pageNumber="22">
25. Elytra black with yellowish brown apices.............................
<taxonomicName authority="(Yang &amp; Wu, 1972)" baseAuthorityName="Yang &amp; Wu" baseAuthorityYear="1972" box="[980,1437,1002,1024]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lamellatus">
<emphasis bold="true" box="[980,1437,1002,1024]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[980,1234,1002,1024]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus lamellatus</emphasis>
(
<bibRefCitation author="Yang, C. T. &amp; Wu, R. H." box="[1246,1429,1002,1024]" pageId="21" pageNumber="22" pagination="115 - 128" refId="ref66088" refString="Yang, C. T. &amp; Wu, R. H. (1972) Notes on some Coccinellidae of Taiwan. Journal of Agriculture and Forestry, 21, 115 - 128. https: // doi. org / 10.2337 / diab. 21.2.128 b" type="journal article" year="1972">Yang &amp; Wu, 1972</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,1031,1053]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,1031,1053]" pageId="21" pageNumber="22">
- Elytra black with yellow to yellowish brown markings (
<figureCitation box="[762,870,1031,1053]" captionStart="FIGURE 37" captionStartId="53.[151,250,1742,1766]" captionTargetBox="[154,1428,184,1690]" captionTargetId="figure-18@53.[151,1436,181,1718]" captionTargetPageId="53" captionText="FIGURE 37. S. variabilis Tong &amp; Wang sp. nov.: a, d dorsal view, b, e lateral view, c, f frontal view, g abdomen, male, h–k male genitalia: h penis, i apex of penis, j tegmen, lateral view, k tegmen, ventral view. Scale bars: a–f: 0.5 mm; g–k: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328164" httpUri="https://zenodo.org/record/7328164/files/figure.png" pageId="21" pageNumber="22">Figs 37a–b</figureCitation>
).......
<emphasis bold="true" box="[962,1436,1031,1053]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[962,1353,1031,1053]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis bold="true" box="[962,1205,1031,1053]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus variabilis</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1032,1053]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,1061,1083]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,1061,1083]" pageId="21" pageNumber="22">26. Apex of penis guide pointed in ventral view............................................................... 27</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,1090,1112]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,1090,1112]" pageId="21" pageNumber="22">- Apex of penis guide blunt in ventral view................................................................. 29</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,1119,1141]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,1119,1141]" pageId="21" pageNumber="22">27. Elytral base mostly black.............................................................................. 28</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,1148,1171]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,1148,1171]" pageId="21" pageNumber="22">
- Elytral base entirely yellow (Figs 38a–b).............................
<taxonomicName authority="(Yu &amp; Pang, 1993)" baseAuthorityName="Yu &amp; Pang" baseAuthorityYear="1993" box="[961,1437,1148,1171]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="seminigrinus">
<emphasis bold="true" box="[961,1437,1148,1171]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[961,1241,1149,1170]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus seminigrinus</emphasis>
(
<bibRefCitation author="Yu, G. Y. &amp; Pang, H. &amp; Pang, X. F." box="[1254,1429,1148,1171]" pageId="21" pageNumber="22" pagination="467 - 505" refId="ref66468" refString="Yu, G. Y., Pang, H. &amp; Pang, X. F. (1993) Coccinellidae collected from Chebaling National Nature Reserve. In: Xu, Y. (Ed.), Collected papers for investigation in National Chebaling Nature Reserve. Science and Technology press of Guangdong, Guangzhou, pp. 467 - 505. [in Chinese]" type="book chapter" year="1993">Yu &amp; Pang, 1993</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,1178,1200]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,1178,1200]" pageId="21" pageNumber="22">
28. Scutellar shield black (
<figureCitation box="[446,529,1178,1200]" captionStart="FIGURE 39" captionStartId="56.[151,250,1393,1417]" captionTargetBox="[203,1382,382,1370]" captionTargetId="figure-123@56.[198,1389,381,1370]" captionTargetPageId="56" captionText="FIGURE 39. S. bivalvis (Yu, 1997): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328168" httpUri="https://zenodo.org/record/7328168/files/figure.png" pageId="21" pageNumber="22">Fig. 39a</figureCitation>
); parameres short and stout (
<figureCitation box="[805,887,1178,1200]" captionStart="FIGURE 39" captionStartId="56.[151,250,1393,1417]" captionTargetBox="[203,1382,382,1370]" captionTargetId="figure-123@56.[198,1389,381,1370]" captionTargetPageId="56" captionText="FIGURE 39. S. bivalvis (Yu, 1997): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328168" httpUri="https://zenodo.org/record/7328168/files/figure.png" pageId="21" pageNumber="22">
Fig.
<quantity box="[850,887,1178,1199]" metricMagnitude="-2" metricUnit="kg" metricValue="3.9" pageId="21" pageNumber="22" unit="g" value="39.0">39g</quantity>
</figureCitation>
).................
<taxonomicName authority="(Yu, 1997)" baseAuthorityName="Yu" baseAuthorityYear="1997" box="[1100,1437,1178,1200]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bivalvis">
<emphasis bold="true" box="[1100,1437,1178,1200]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[1100,1326,1178,1200]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus bivalvis</emphasis>
(Yu, 1997)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" pageId="21" pageNumber="22">
<keyLead pageId="21" pageNumber="22">
- Scutellar shield yellowish brown (Fig. 40a); parameres long and slender (Fig.
<quantity box="[976,1013,1208,1229]" metricMagnitude="-2" metricUnit="kg" metricValue="4.0" pageId="21" pageNumber="22" unit="g" value="40.0">40g</quantity>
).
<emphasis bold="true" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[1042,1402,1207,1229]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="flexus" status="sp. nov.">
<emphasis bold="true" box="[1042,1253,1207,1229]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus flexus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,1266,1288]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,1266,1288]" pageId="21" pageNumber="22">29. Apex of penis expanded............................................................................... 30</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1436,1295,1317]" pageId="21" pageNumber="22">
<keyLead box="[151,1436,1295,1317]" pageId="21" pageNumber="22">- Apex of penis not expanded............................................................................ 31</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" pageId="21" pageNumber="22">
<keyLead pageId="21" pageNumber="22">
30. Penis guide long; apex of penis with distinct membranous appendage (
<figureCitation box="[904,982,1325,1347]" captionStart="FIGURE 41" captionStartId="59.[151,250,1652,1676]" captionTargetBox="[157,1431,563,1627]" captionTargetId="figure-165@59.[151,1436,561,1628]" captionTargetPageId="59" captionText="FIGURE 41. S. sagittalis Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328170" httpUri="https://zenodo.org/record/7328170/files/figure.png" pageId="21" pageNumber="22">Fig. 41f</figureCitation>
)..................................................................................................
<emphasis bold="true" box="[966,1436,1354,1376]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[966,1353,1354,1376]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sagittalis" status="sp. nov.">
<emphasis bold="true" box="[966,1205,1354,1376]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus sagittalis</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1354,1375]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" pageId="21" pageNumber="22">
<keyLead pageId="21" pageNumber="22">
- Penis guide short; apex of penis with inconspicuous membranous appendage (
<figureCitation box="[976,1054,1383,1405]" captionStart="FIGURE 42" captionStartId="60.[151,250,1652,1676]" captionTargetBox="[158,1426,562,1627]" captionTargetId="figure-188@60.[151,1436,560,1628]" captionTargetPageId="60" captionText="FIGURE 42. S. robustus Tong &amp; Wang sp. nov.: a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328174" httpUri="https://zenodo.org/record/7328174/files/figure.png" pageId="21" pageNumber="22">Fig. 42f</figureCitation>
)............................................................................................
<emphasis bold="true" box="[968,1436,1412,1435]" pageId="21" pageNumber="22">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[968,1353,1412,1435]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis bold="true" box="[968,1205,1412,1434]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus robustus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[1360,1436,1413,1434]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="21" pageNumber="22">
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,1442,1464]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,1442,1464]" pageId="21" pageNumber="22">
31. Apex of penis with membranous appendage on both sides (
<figureCitation box="[794,871,1442,1464]" captionStart="FIGURE 43" captionStartId="62.[151,250,1241,1265]" captionTargetBox="[184,1407,183,1215]" captionTargetId="figure-29@62.[172,1415,181,1217]" captionTargetPageId="62" captionText="FIGURE 43. S. lancetapicalis (Pang &amp; Gordon, 1986): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–h male genitalia: e penis, f apex of penis, g tegmen, lateral view, h tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–h: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328176" httpUri="https://zenodo.org/record/7328176/files/figure.png" pageId="21" pageNumber="22">Fig. 43f</figureCitation>
)..
<taxonomicName authority="(Pang &amp; Gordon, 1986)" baseAuthorityName="Pang &amp; Gordon" baseAuthorityYear="1986" box="[903,1437,1442,1464]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lancetapicalis">
<emphasis bold="true" box="[903,1437,1442,1464]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[903,1190,1442,1464]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus lancetapicalis</emphasis>
(
<bibRefCitation author="Pang, X. F. &amp; Gordon, R. D." box="[1203,1429,1442,1464]" pageId="21" pageNumber="22" pagination="157 - 199" refId="ref65250" refString="Pang, X. F. &amp; Gordon, R. D. (1986) The Scymnini (Coleoptera: Coccinellidae) of China. The Coleopterists Bulletin, 40 (2), 157 - 199." type="journal article" year="1986">Pang &amp; Gordon, 1986</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="21.[151,1437,151,1493]" box="[151,1437,1471,1493]" pageId="21" pageNumber="22">
<keyLead box="[151,1437,1471,1493]" pageId="21" pageNumber="22">
- Apex of penis with membranous appendage on lateral side (
<figureCitation box="[802,880,1471,1493]" captionStart="FIGURE 44" captionStartId="63.[151,250,1539,1563]" captionTargetBox="[197,1392,563,1511]" captionTargetId="figure-151@63.[194,1393,561,1516]" captionTargetPageId="63" captionText="FIGURE 44. S. disselasmatus (Pang &amp; Huang, 1986), holotype: a dorsal view, b lateral view, c frontal view, d labels, e abdomen, f–i male genitalia: f penis, g apex of penis, h tegmen, lateral view, i tegmen, ventral view. Scale bars: a–c: 0.5 mm; e–i: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328180" httpUri="https://zenodo.org/record/7328180/files/figure.png" pageId="21" pageNumber="22">Fig. 44f</figureCitation>
)..
<taxonomicName authority="(Pang &amp; Huang, 1986)" baseAuthorityName="Pang &amp; Huang" baseAuthorityYear="1986" box="[913,1437,1471,1493]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="disselasmatus">
<emphasis bold="true" box="[913,1437,1471,1493]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[913,1201,1471,1493]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus disselasmatus</emphasis>
(
<bibRefCitation author="Pang, X. F. &amp; Huang, B. K." box="[1213,1429,1471,1493]" pageId="21" pageNumber="22" pagination="61 - 73" refId="ref65289" refString="Pang, X. F. &amp; Huang, B. K. (1986) Six new species of Scymnus and four new species of Pseudoscymnus from Fujian (Coleoptera: Coccinellidae: Scymnini). Journal of Wuyi Science, 6, 61 - 73. [in Chinese with English summary]" type="journal article" year="1986">Pang &amp; Huang, 1986</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="discussion">
<paragraph blockId="21.[151,1436,1540,1602]" pageId="21" pageNumber="22">
<emphasis bold="true" box="[151,267,1540,1566]" pageId="21" pageNumber="22">Remarks.</emphasis>
<taxonomicName authority="(Weise, 1879)" baseAuthorityName="Weise" baseAuthorityYear="1879" box="[276,687,1540,1566]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="hareja">
<emphasis box="[276,520,1540,1566]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus hareja</emphasis>
(
<bibRefCitation author="Weise, J." box="[536,678,1540,1566]" pageId="21" pageNumber="22" pagination="147 - 152" refId="ref65987" refString="Weise, J. (1879) Beitrage zur kaferfauna von Japan. Deutsche Entomologische Zeitschrift, 23, 147 - 152. https: // doi. org / 10.1002 / mmnd. 48018790119" type="journal article" year="1879">Weise, 1879</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName authority="(Yang, 1971)" baseAuthorityName="Yang" baseAuthorityYear="1971" box="[745,1150,1540,1566]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="changi">
<emphasis box="[745,993,1540,1566]" italics="true" pageId="21" pageNumber="22">Sasajiscymnus changi</emphasis>
(
<bibRefCitation author="Yang, C. T." box="[1009,1142,1540,1566]" pageId="21" pageNumber="22" pagination="85 - 96" refId="ref66136" refString="Yang, C. T. (1971) Notes on the species of genus Pseudoscymnus from Taiwan. Journal of Agriculture and Forestry, 20: 85 - 96." type="journal article" year="1971">Yang, 1971</bibRefCitation>
)
</taxonomicName>
were not included in the above key since no specimens of these species were available for us.
</paragraph>
</subSubSection>
</treatment>
</document>