treatments-xml/data/31/42/87/314287B0F61CFF8FFF7BFE55087AFD04.xml

<document ID-DOI="10.11646/zootaxa.5207.1.1" ID-GBIF-Dataset="a1bc8125-ad00-4d07-b9a1-4e6420c6bd98" ID-ISSN="1175­5334" ID-Zenodo-Dep="7328080" ID-ZooBank="A57EB27B-8524-42D8-8B85-564F879A5938" checkinTime="1668423676913" checkinUser="plazi" docAuthor="TONG, JUNBO, CHEN, XIAOSHENG, ZHANG, XIAONING, HUO, LIZHI, WU, GANG, PENG, ZHENGQIANG &amp; WANG, XINGMIN" docDate="2022" docId="314287B0F61CFF8FFF7BFE55087AFD04" docLanguage="en" docName="zootaxa.5207.1.1.pdf" docOrigin="Zootaxa 5207 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5207.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.4:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="4" docTitle="Sasajiscymnus niganulus Tong &amp; Wang, sp. nov." docType="treatment" docVersion="2" lastPageNumber="47" masterDocId="CD7BFFC8F631FFA1FFECFFE20B40FF81" masterDocTitle="A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae)" masterLastPageNumber="104" masterPageNumber="1" pageNumber="46" updateTime="1668629162570" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>A review of the genus Sasajiscymnus Vandenberg, 2004 from China (Coleoptera, Coccinellidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>TONG, JUNBO</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CHEN, XIAOSHENG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ZHANG, XIAONING</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>HUO, LIZHI</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WU, GANG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>PENG, ZHENGQIANG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WANG, XINGMIN</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5207</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>104</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5207.1.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5207.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">a1bc8125-ad00-4d07-b9a1-4e6420c6bd98</mods:identifier>
<mods:identifier type="ISSN">1175­5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">7328080</mods:identifier>
<mods:identifier type="ZooBank">A57EB27B-8524-42D8-8B85-564F879A5938</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7328260" ID-Zenodo-Dep="7328260" LSID="urn:lsid:plazi:treatment:314287B0F61CFF8FFF7BFE55087AFD04" httpUri="http://treatment.plazi.org/id/314287B0F61CFF8FFF7BFE55087AFD04" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<subSubSection pageId="45" pageNumber="46" type="nomenclature">
<paragraph blockId="45.[151,741,438,537]" box="[151,741,438,465]" pageId="45" pageNumber="46">
<heading bold="true" box="[151,741,438,465]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<emphasis bold="true" box="[151,741,438,465]" pageId="45" pageNumber="46">
<taxonomicName authority="Tong &amp; Wang" authorityName="Tong &amp; Wang" box="[151,639,438,465]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="niganulus" status="sp. nov.">
<emphasis bold="true" box="[151,458,438,465]" italics="true" pageId="45" pageNumber="46">Sasajiscymnus niganulus</emphasis>
Tong &amp; Wang
</taxonomicName>
<taxonomicNameLabel box="[647,741,439,465]" pageId="45" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="45.[151,741,438,537]" box="[151,295,475,501]" pageId="45" pageNumber="46">(Figs 31, 71)</paragraph>
</subSubSection>
<subSubSection box="[151,463,511,537]" pageId="45" pageNumber="46" type="description">
<paragraph blockId="45.[151,741,438,537]" box="[151,463,511,537]" pageId="45" pageNumber="46">
Chinese name: 
<emphasis bold="true" box="[324,463,511,534]" pageId="45" pageNumber="46">DZẸḧ瓢•</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="diagnosis">
<paragraph blockId="45.[151,1436,582,897]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[151,274,582,608]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
This species is similar to 
<taxonomicName authority="(Ohta, 1929)" baseAuthorityName="Ohta" baseAuthorityYear="1929" box="[567,841,583,609]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="seboshii">
<emphasis box="[567,688,583,609]" italics="true" pageId="45" pageNumber="46">S. seboshii</emphasis>
(
<bibRefCitation author="Ohta, Y." box="[703,834,583,609]" pageId="45" pageNumber="46" pagination="1 - 16" refId="ref65137" refString="Ohta, Y. (1929) Scymninen Japans. Insecta Matsumurana, 4 (1 - 2), 1 - 16." type="journal article" year="1929">Ohta, 1929</bibRefCitation>
)
</taxonomicName>
, but can be distinguished from the later by the elytra with black humeral angles, abdominal postcoxal lines recurved toward the base of ventrite and more elongate penis guide (Figs 31a–d, 
<quantity box="[367,409,655,681]" metricMagnitude="-2" metricUnit="kg" metricValue="1.9" pageId="45" pageNumber="46" unit="g" value="19.0">19g</quantity>
–h). In 
<taxonomicName baseAuthorityName="Ohta" baseAuthorityYear="1929" box="[490,610,655,681]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="seboshii">
<emphasis box="[490,610,655,681]" italics="true" pageId="45" pageNumber="46">S. seboshii</emphasis>
</taxonomicName>
, humeral angles of elytra are yellowish brown, abdominal postcoxal lines not recurved toward the base of ventrite and penis guide is shorter (
<figureCitation box="[906,1034,691,717]" captionStart="FIGURE 34" captionStartId="49.[151,250,1782,1806]" captionTargetBox="[157,1429,239,1747]" captionTargetId="figure-52@49.[151,1436,236,1758]" captionTargetPageId="49" captionText="FIGURE 34. S. seboshii (Ohta, 1929): a, d dorsal view, b, e lateral view, c, f frontal view, g abdomen, male, h–k male genitalia: h penis, i apex of penis, j tegmen, lateral view, k tegmen, ventral view. Scale bars: a–f: 0.5 mm; g–k: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328158" httpUri="https://zenodo.org/record/7328158/files/figure.png" pageId="45" pageNumber="46">Figs 34a–g</figureCitation>
, 
<figureCitation box="[1045,1113,691,717]" captionStart="FIGURE 19" captionStartId="29.[151,250,1904,1928]" captionTargetBox="[158,1427,816,1876]" captionTargetId="figure-262@29.[151,1436,813,1880]" captionTargetPageId="29" captionText="FIGURE 19. S. ocellatus (Yu, 2000): a dorsal view, b lateral view, c frontal view, d abdomen, male, e–g male genitalia: e penis, f tegmen, lateral view, g tegmen, ventral view. Scale bars: a–c: 0.5 mm; d–g: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7328128" httpUri="https://zenodo.org/record/7328128/files/figure.png" pageId="45" pageNumber="46">19j–k</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph blockId="45.[151,1436,582,897]" box="[199,1032,726,753]" pageId="45" pageNumber="46">
<emphasis bold="true" box="[199,345,726,752]" pageId="45" pageNumber="46">Description.</emphasis>
TL: 
<quantity box="[400,565,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="2.0949999999999998" metricValueMax="2.25" metricValueMin="1.94" pageId="45" pageNumber="46" unit="mm" value="2.0949999999999998" valueMax="2.25" valueMin="1.94">1.94–2.25 mm</quantity>
, TW: 
<quantity box="[633,798,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.505" metricValueMax="1.63" metricValueMin="1.38" pageId="45" pageNumber="46" unit="mm" value="1.505" valueMax="1.63" valueMin="1.38">1.38–1.63 mm</quantity>
, TH: 
<quantity box="[862,1028,727,753]" metricMagnitude="-4" metricUnit="m" metricValue="9.95" metricValueMax="10.9" metricValueMin="9.0" pageId="45" pageNumber="46" unit="mm" value="0.995" valueMax="1.09" valueMin="0.9">0.90–1.09 mm</quantity>
.
</paragraph>
<paragraph blockId="45.[151,1436,582,897]" pageId="45" pageNumber="46">Body oval, moderately convex, surfaces covered with short pubescence (Figs 31a–c). Head yellow with interocular distance distinctly greater than eye width in frontal view. Pronotum yellow. Scutellar shield yellowish brown. Elytra black with yellow apices and a large yellowish brown spot in the middle part. Ventral surfaces yellow to black. Legs yellow. Abdominal postcoxal lines incomplete, recurved toward the base of ventrite (Fig. 31d).</paragraph>
<paragraph blockId="46.[151,1437,151,645]" pageId="46" pageNumber="47">
Male genitalia: Penis stout, penis capsule with stout outer arm and long inner one, apex of penis constricted with membranous appendage (Figs 31e–f). Penis guide with prominent keel at base in lateral view, then tapering towards pointed apex; parameres about 1/3 as long as penis guide, with a tuft of long setae at apex; penis guide widest at base, then gradually narrowing to pointed apex (Figs 
<quantity box="[749,791,259,285]" metricMagnitude="-2" metricUnit="kg" metricValue="3.1" pageId="46" pageNumber="47" unit="g" value="31.0">31g</quantity>
–h).
</paragraph>
<paragraph blockId="46.[151,1437,151,645]" pageId="46" pageNumber="47">Female genitalia: Coxites elongate, triangular; styli well developed with terminal setae; spermatheca distinctly curved, with basal ramus and elongate nodulus.</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="materials_examined">
<paragraph blockId="46.[151,1437,151,645]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[199,374,366,393]" pageId="46" pageNumber="47">Type material.</emphasis>
<materialsCitation collectingDate="2011-10-13" collectionCode="SCAU" collectorName="Huo LZ" country="China" county="Motuo County" elevation="3150" location="Zhamo Highroad" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis bold="true" box="[382,692,366,393]" pageId="46" pageNumber="47">
<typeStatus box="[382,495,367,393]" pageId="46" pageNumber="47">Holotype</typeStatus>
: 
<collectingCountry box="[508,585,366,392]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
: 
<collectingRegion box="[599,687,366,392]" country="China" name="Xizang" pageId="46" pageNumber="47">Xizang</collectingRegion>
:
</emphasis>
<specimenCount box="[700,723,367,393]" pageId="46" pageNumber="47" type="male">♂</specimenCount>
, 52K–65K of 
<location LSID="urn:lsid:plazi:treatment:314287B0F61CFF8FFF7BFE55087AFD04:BC34607DF61FFF8FFC9BFE8D0F7BFE08" box="[887,1083,367,393]" country="China" county="Motuo County" name="Zhamo Highroad" pageId="46" pageNumber="47" stateProvince="Xizang">Zhamo Highroad</location>
, 
<collectingCounty box="[1095,1262,367,393]" pageId="46" pageNumber="47">Motuo County</collectingCounty>
, 
<quantity box="[1273,1431,366,393]" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="46" pageNumber="47" unit="m" value="3150.0" valueMax="3500.0" valueMin="2800.0">
<elevation box="[1273,1431,366,393]" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="46" pageNumber="47" unit="m" value="3150.0" valueMax="3500.0" valueMin="2800.0">2800–3500 m</elevation>
</quantity>
, 
<date box="[151,271,403,429]" pageId="46" pageNumber="47" value="2011-10-13">
<collectingDate box="[151,271,403,429]" pageId="46" pageNumber="47" value="2011-10-13">13.X.2011</collectingDate>
</date>
, 
<collectorName box="[282,372,403,429]" pageId="46" pageNumber="47">Huo LZ</collectorName>
leg (
<collectionCode box="[428,509,403,429]" pageId="46" pageNumber="47">SCAU</collectionCode>
)
</materialsCitation>
. 
<materialsCitation collectingDate="2007-10-30" collectingDateMax="2007-10-31" collectingDateMin="2007-10-30" collectionCode="SCAU" collectorName="Wang XM" country="China" county="Bomi County" elevation="835" location="Zhamo Highroad" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
<emphasis bold="true" box="[528,949,402,429]" pageId="46" pageNumber="47">
<typeStatus box="[528,648,403,429]" pageId="46" pageNumber="47">Paratypes</typeStatus>
(
<collectionCode box="[663,746,402,428]" pageId="46" pageNumber="47">SCAU</collectionCode>
): 
<collectingCountry box="[766,843,402,428]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
: 
<collectingRegion box="[855,943,402,428]" country="China" name="Xizang" pageId="46" pageNumber="47">Xizang</collectingRegion>
:
</emphasis>
<specimenCount box="[956,991,403,429]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:314287B0F61CFF8FFF7BFE55087AFD04:BC34607DF61FFF8FFC01FE710FEDFE2C" box="[1005,1197,403,429]" country="China" county="Bomi County" name="Zhamo Highroad" pageId="46" pageNumber="47" stateProvince="Xizang">Zhamo Highroad</location>
113K, 
<collectingCounty box="[1279,1433,403,429]" pageId="46" pageNumber="47">Bomi County</collectingCounty>
, 
<quantity box="[151,227,438,465]" metricMagnitude="2" metricUnit="m" metricValue="8.35" pageId="46" pageNumber="47" unit="m" value="835.0">
<elevation box="[151,227,438,465]" metricMagnitude="2" metricUnit="m" metricValue="8.35" pageId="46" pageNumber="47" unit="m" value="835.0">835 m</elevation>
</quantity>
, 
<date box="[241,404,439,465]" pageId="46" pageNumber="47" value="2007-10-30" valueMax="2007-10-31" valueMin="2007-10-30">
<collectingDate box="[241,404,439,465]" pageId="46" pageNumber="47" value="2007-10-30" valueMax="2007-10-31" valueMin="2007-10-30">30–31.X.2007</collectingDate>
</date>
, 
<collectorName box="[417,536,439,465]" pageId="46" pageNumber="47">Wang XM</collectorName>
leg
</materialsCitation>
; 
<materialsCitation box="[597,1431,439,465]" collectingDate="2009-10-13" collectorName="Wang XM" country="China" county="Pailong Town" location="Linzhi City" municipality="Bayi District" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
<specimenCount box="[597,634,439,465]" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
, 
<collectingCounty box="[649,809,439,465]" pageId="46" pageNumber="47">Pailong Town</collectingCounty>
, 
<collectingMunicipality box="[823,972,439,465]" pageId="46" pageNumber="47">Bayi District</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:314287B0F61CFF8FFF7BFE55087AFD04:BC34607DF61FFF8FFC36FE550F1DFE50" box="[986,1117,439,465]" country="China" county="Pailong Town" municipality="Bayi District" name="Linzhi City" pageId="46" pageNumber="47" stateProvince="Xizang">Linzhi City</location>
, 
<date box="[1131,1251,439,465]" pageId="46" pageNumber="47" value="2009-10-13">
<collectingDate box="[1131,1251,439,465]" pageId="46" pageNumber="47" value="2009-10-13">13.X.2009</collectingDate>
</date>
, 
<collectorName box="[1265,1385,439,465]" pageId="46" pageNumber="47">Wang XM</collectorName>
leg
</materialsCitation>
; 
<materialsCitation box="[151,940,474,501]" collectingDate="2011-10-05" collectingDateMax="2011-10-08" collectingDateMin="2011-10-05" collectorName="Huo LZ" country="China" county="Muotuo County" elevation="1450" location="Hanmi" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
<specimenCount box="[151,186,475,501]" pageId="46" pageNumber="47" type="female">1♀</specimenCount>
, 
<location LSID="urn:lsid:plazi:treatment:314287B0F61CFF8FFF7BFE55087AFD04:BC34607DF61FFF8FFF27FE390A5AFE74" box="[203,282,475,501]" country="China" county="Muotuo County" name="Hanmi" pageId="46" pageNumber="47" stateProvince="Xizang">Hanmi</location>
, 
<collectingCounty box="[296,479,475,501]" pageId="46" pageNumber="47">Muotuo County</collectingCounty>
, 
<quantity box="[492,638,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="2.1" metricValueMin="0.8" pageId="46" pageNumber="47" unit="m" value="1450.0" valueMax="2100.0" valueMin="800.0">
<elevation box="[492,638,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="2.1" metricValueMin="0.8" pageId="46" pageNumber="47" unit="m" value="1450.0" valueMax="2100.0" valueMin="800.0">800–2100 m</elevation>
</quantity>
, 
<date box="[653,786,474,501]" pageId="46" pageNumber="47" value="2011-10-05" valueMax="2011-10-08" valueMin="2011-10-05">
<collectingDate box="[653,786,474,501]" pageId="46" pageNumber="47" value="2011-10-05" valueMax="2011-10-08" valueMin="2011-10-05">5–8.X.2011</collectingDate>
</date>
, 
<collectorName box="[801,893,475,501]" pageId="46" pageNumber="47">Huo LZ</collectorName>
leg
</materialsCitation>
; 
<materialsCitation collectingDate="2011-10-19" collectorName="Huo LZ" country="China" elevation="2200" location="Linzhi City" municipality="Chilongzangbu" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
<specimenCount box="[955,992,475,501]" pageId="46" pageNumber="47" type="male">1♂</specimenCount>
, 
<collectingMunicipality box="[1007,1181,475,501]" pageId="46" pageNumber="47">Chilongzangbu</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:314287B0F61CFF8FFF7BFE55087AFD04:BC34607DF61FFF8FFB47FE390E6FFE74" box="[1195,1327,475,501]" country="China" municipality="Chilongzangbu" name="Linzhi City" pageId="46" pageNumber="47" stateProvince="Xizang">Linzhi City</location>
, 
<quantity box="[1341,1431,475,501]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="46" pageNumber="47" unit="m" value="2200.0">
<elevation box="[1341,1431,475,501]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="46" pageNumber="47" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, 
<date box="[151,270,511,537]" pageId="46" pageNumber="47" value="2011-10-19">
<collectingDate box="[151,270,511,537]" pageId="46" pageNumber="47" value="2011-10-19">19.X.2011</collectingDate>
</date>
, 
<collectorName box="[282,372,511,537]" pageId="46" pageNumber="47">Huo LZ</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[199,540,546,573]" pageId="46" pageNumber="47" type="distribution">
<paragraph blockId="46.[151,1437,151,645]" box="[199,540,546,573]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[199,353,546,572]" pageId="46" pageNumber="47">Distribution.</emphasis>
<collectingCountry box="[359,426,547,573]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
(
<collectingRegion box="[441,528,547,573]" country="China" name="Xizang" pageId="46" pageNumber="47">Xizang</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="etymology">
<paragraph blockId="46.[151,1437,151,645]" pageId="46" pageNumber="47">
<emphasis bold="true" box="[199,334,583,609]" pageId="46" pageNumber="47">Etymology.</emphasis>
The specific epithet 
<taxonomicName box="[568,678,583,609]" class="Insecta" family="Coccinellidae" genus="Sasajiscymnus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="niganulus">
<emphasis box="[568,678,583,609]" italics="true" pageId="46" pageNumber="47">niganulus</emphasis>
</taxonomicName>
derives from the Latin words “niger” (= black) and “anulus” (= annular), and refers to the black ring-spot on the disc of elytra.
</paragraph>
</subSubSection>
</treatment>
</document>