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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1013.55600" ID-GBIF-Dataset="a967c90e-ea10-4a10-b43b-cd5328a1d6e6" ID-PMC="PMC8390796" ID-Pensoft-Pub="1313-2970-1013-1" ID-Pensoft-UUID="E4329863A39E5EEBA395938413BDD579" ID-PubMed="34512087" ID-ZooBank="CFDCEFBB523040339D46E302F66E9886" ModsDocID="1313-2970-1013-1" checkinTime="1612290696301" checkinUser="pensoft" docAuthor="Sharkey, Michael J., Janzen, Daniel H., Hallwachs, Winnie, Chapman, Eric G., Smith, M. Alex, Dapkey, Tanya, Brown, Allison, Ratnasingham, Sujeevan, Naik, Suresh, Manjunath, Ramya, Perez, Kate, Milton, Megan, Hebert, Paul, Shaw, Scott R., Kittel, Rebecca N., Solis, M. Alma, Metz, Mark A., Goldstein, Paul Z., Brown, John W., Quicke, Donald L. J., Achterberg, C. van, Brown, Brian V. &amp; Burns, John M." docDate="2021" docId="2FA7B141616E8A16A7D84C2C9E0D74C6" docLanguage="en" docName="ZooKeys 1013: 1-665" docOrigin="ZooKeys 1013" docPubDate="2021-02-02" docSource="http://dx.doi.org/10.3897/zookeys.1013.55600" docTitle="Macrocentrus harisridhari Sharkey &amp; van Achterberg 2021, sp. nov." docType="treatment" docUuid="27944FF6-D485-4D92-8A04-0813E4B866D2" docUuidSource="ZooBank" docVersion="4" id="E4329863A39E5EEBA395938413BDD579" lastPageNumber="1" masterDocId="E4329863A39E5EEBA395938413BDD579" masterDocTitle="Minimalist revision and description of 403 new species in 11 subfamilies of Costa Rican braconid parasitoid wasps, including host records for 219 species" masterLastPageNumber="665" masterPageNumber="1" pageNumber="1" updateTime="1668149748640" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Minimalist revision and description of 403 new species in 11 subfamilies of Costa Rican braconid parasitoid wasps, including host records for 219 species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sharkey, Michael J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6201-7340</mods:nameIdentifier>
<mods:affiliation>The Hymenoptera Institute, 116 Franklin Ave., Redlands, CA, 92373, USA</mods:affiliation>
<mods:nameIdentifier type="email">msharkey@uky.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Janzen, Daniel H.</mods:namePart>
<mods:affiliation>Department of Biology, University of Pennsylvania, Philadelphia, PA 19104 - 6018, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hallwachs, Winnie</mods:namePart>
<mods:affiliation>Department of Biology, University of Pennsylvania, Philadelphia, PA 19104 - 6018, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chapman, Eric G.</mods:namePart>
<mods:affiliation>Department of Entomology, University of Kentucky, Lexington, KY 40546 - 0091, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, M. Alex</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8650-2575</mods:nameIdentifier>
<mods:affiliation>Department of Integrative Biology, University of Guelph and Biodiversity Institute of Ontario, Guelph, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dapkey, Tanya</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brown, Allison</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ratnasingham, Sujeevan</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naik, Suresh</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Manjunath, Ramya</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perez, Kate</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Milton, Megan</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hebert, Paul</mods:namePart>
<mods:affiliation>Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaw, Scott R.</mods:namePart>
<mods:affiliation>Department of Ecosystem Science, University of Wyoming, 1000 East University Avenue, Laramie, Wyoming 82071, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kittel, Rebecca N.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0032-5764</mods:nameIdentifier>
<mods:affiliation>Museum Wiesbaden, Hessisches Landesmuseum fuer Kunst und Natur, Friedrich-Ebert-Allee 2, 65185 Wiesbaden, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Solis, M. Alma</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6379-1004</mods:nameIdentifier>
<mods:affiliation>Systematic Entomology Laboratory, Beltsville Agriculture Research Center, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum Natural History, MRC 168, Smithsonian Institution, P. O. Box 37012, Washington, DC, 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Metz, Mark A.</mods:namePart>
<mods:affiliation>Systematic Entomology Laboratory, Beltsville Agriculture Research Center, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum Natural History, MRC 168, Smithsonian Institution, P. O. Box 37012, Washington, DC, 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goldstein, Paul Z.</mods:namePart>
<mods:affiliation>Systematic Entomology Laboratory, Beltsville Agriculture Research Center, Agricultural Research Service, U. S. Department of Agriculture, c / o National Museum Natural History, MRC 168, Smithsonian Institution, P. O. Box 37012, Washington, DC, 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brown, John W.</mods:namePart>
<mods:affiliation>Division of Entomology, PO Box 37012 12. National Museum of Natural History E 515 MRC 127, Washington, DC 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Quicke, Donald L. J.</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Life Sciences, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Achterberg, C. van</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation>Naturalis Biodiversity Center, Postbus 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brown, Brian V.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6367-6057</mods:nameIdentifier>
<mods:affiliation>Department of Entomology, Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, CA, 90007, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burns, John M.</mods:namePart>
<mods:affiliation>Division of Entomology, PO Box 37012 12. National Museum of Natural History E 515 MRC 127, Washington, DC 20013 - 7012, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1013</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>665</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1013.55600</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1013.55600</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1013-1</mods:identifier>
<mods:identifier type="ZooBank">CFDCEFBB523040339D46E302F66E9886</mods:identifier>
<mods:identifier type="Pensoft-UUID">E4329863A39E5EEBA395938413BDD579</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="177205988" LSID="urn:lsid:zoobank.org:act:27944FF6-D485-4D92-8A04-0813E4B866D2" httpUri="http://treatment.plazi.org/id/2FA7B141616E8A16A7D84C2C9E0D74C6" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/27944FF6-D485-4D92-8A04-0813E4B866D2" authority="Sharkey &amp; van Achterberg" authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Macrocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocentrus harisridhari" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="harisridhari" status="sp. nov.">Macrocentrus harisridhari Sharkey &amp; van Achterberg</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 305" captionStartId="F305" captionText="Figure 305. Macrocentrus harisridhari, holotype." figureDoi="10.3897/zookeys.1013.55600.figure305" httpUri="https://binary.pensoft.net/fig/503760" pageId="0" pageNumber="1">Figure 305</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnostics">
<paragraph pageId="0" pageNumber="1">Diagnostics.</paragraph>
<paragraph pageId="0" pageNumber="1">
BOLD:ACK7467. Consensus barcode. TATATTGTATTTTTTWWTTGGYATRTGRTCRGGAGTATTAGGTTTATCATTAAGTTTAATTATTCGTATAGAATTAGGTCAAATTGGKTCATTTATTGGAAATGATCAGATTTATAATAGTATYGTTACTTCTCATGCTTTYATTATAATTTTTTTTATAGTTATACCTATTATAATTGGKGGATTTGGMAATTGATTAATTCCTTTRATRTTRGGAAGTGTWGATATRGCYTTYCCTCGAATAAATAATATAAGATTTTGATTATTAATTCCTTCTTTAATATTATTAATTTTAAGAGGWTTTATAAATATTGGTGTAGGTACAGGATGAACAGTTTATCCYCCYTTATCATTAAATATTAGTCATATAGGAATTTCTGTAGATATAGCTATTTTTTCATTACATTTGGCGGGTATTTCTTCAATTATAGGTGCTATTAATTTTATTGTTACTATTATAAATATACGAAATTATGGGGTATTAATAGATAAAATTAGATTATTATCATGATCAATTTTAATCACAGCTATTTTATTATTATTATCTTTACCTGTRTTAGCTGGTGCWATTACAATRTTGTTAACTGATCGTAATTTAAATACATCTTTYTTTGACCCYGCYGGAGGGGGTGACCCTRTTTTATAYCAACATTTATTT. There are two species in this BIN (
<taxonomicName lsidName="M. harisridhari" pageId="0" pageNumber="1" rank="species" species="harisridhari">
<emphasis italics="true" pageId="0" pageNumber="1">M. harisridhari</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="M. hiroshikidonoi" pageId="0" pageNumber="1" rank="species" species="hiroshikidonoi">
<emphasis italics="true" pageId="0" pageNumber="1">M. hiroshikidonoi</emphasis>
</taxonomicName>
) that differ by almost 2%, and both share having the metasoma dark brown medio-dorsally, vein 1Cua of forewing distinctly longer than vein cu-a, R1b paler than pterostigma, and vein cu-a curved and posteriorly widened. Their barcodes form two groups with no intermediates. The two species also differ in color:
<taxonomicName lsidName="M. harisridhari" pageId="0" pageNumber="1" rank="species" species="harisridhari">
<emphasis italics="true" pageId="0" pageNumber="1">M. harisridhari</emphasis>
</taxonomicName>
is darker than
<taxonomicName lsidName="M. hiroshikidonoi" pageId="0" pageNumber="1" rank="species" species="hiroshikidonoi">
<emphasis italics="true" pageId="0" pageNumber="1">M. hiroshikidonoi</emphasis>
</taxonomicName>
i.e.,
<taxonomicName lsidName="M. harisridhari" pageId="0" pageNumber="1" rank="species" species="harisridhari">
<emphasis italics="true" pageId="0" pageNumber="1">M. harisridhari</emphasis>
</taxonomicName>
is dark brown where
<taxonomicName lsidName="M. hiroshikidonoi" pageId="0" pageNumber="1" rank="species" species="hiroshikidonoi">
<emphasis italics="true" pageId="0" pageNumber="1">M. hiroshikidonoi</emphasis>
</taxonomicName>
is light reddish brown, and brownish yellow where
<taxonomicName lsidName="M. hiroshikidonoi" pageId="0" pageNumber="1" rank="species" species="hiroshikidonoi">
<emphasis italics="true" pageId="0" pageNumber="1">M. hiroshikidonoi</emphasis>
</taxonomicName>
is light yellow. Morphologically, both are very similar, but
<taxonomicName lsidName="M. harisridhari" pageId="0" pageNumber="1" rank="species" species="harisridhari">
<emphasis italics="true" pageId="0" pageNumber="1">M. harisridhari</emphasis>
</taxonomicName>
differs by having the first metasomal tergite slightly widened posteriorly and slightly shorter POL (0.7
<normalizedToken originalValue="×">x</normalizedToken>
diameter of posterior ocellus vs. 0.9
<normalizedToken originalValue="×">x</normalizedToken>
in
<taxonomicName lsidName="M. hiroshikidonoi" pageId="0" pageNumber="1" rank="species" species="hiroshikidonoi">
<emphasis italics="true" pageId="0" pageNumber="1">M. hiroshikidonoi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype ♀">
<paragraph pageId="0" pageNumber="1">Holotype ♀.</paragraph>
<paragraph pageId="0" pageNumber="1">
Alajuela, Sector Rincon Rain Forest, Flecha,
<geoCoordinate degrees="10.94741" direction="north" orientation="latitude" precision="1" value="10.94741">10.94741</geoCoordinate>
,
<geoCoordinate degrees="85.31501" direction="west" orientation="longitude" precision="1" value="-85.31501">-85.31501</geoCoordinate>
, 491 meters, caterpillar collection date: 21/i/2013, wasp eclosion date: 6/ii/2013. Depository: CNC.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Host data</emphasis>
.
</emphasis>
<taxonomicName authorityName="Lange" authorityYear="1956" class="Insecta" family="Crambidae" genus="Undulambia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Undulambia" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Undulambia</emphasis>
</taxonomicName>
Solis02 (
<taxonomicName class="Insecta" family="Crambidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Crambidae</taxonomicName>
) feeding on
<taxonomicName class="Filicopsida" family="Cyatheaceae" genus="Cyathea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cyathea multiflora" order="Cyatheales" pageId="0" pageNumber="1" phylum="Filicinophyta" rank="species" species="multiflora">
<emphasis italics="true" pageId="0" pageNumber="1">Cyathea multiflora</emphasis>
</taxonomicName>
(
<taxonomicName class="Filicopsida" family="Cyatheaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Cyatheales" pageId="0" pageNumber="1" phylum="Filicinophyta" rank="family">Cyatheaceae</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Caterpillar and holotype voucher codes</emphasis>
.
</emphasis>
13-SRNP-69137, DHJPAR0051314.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="paratypes">
<paragraph pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="1">
Hosts =
<taxonomicName authorityName="Lange" authorityYear="1956" class="Insecta" family="Crambidae" genus="Undulambia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Undulambia" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Undulambia</emphasis>
</taxonomicName>
Solis02,
<taxonomicName authorityName="Hampson" authorityYear="1895" class="Insecta" family="Crambidae" genus="Neurophyseta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Neurophyseta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Neurophyseta</emphasis>
</taxonomicName>
camptogrammalisDHJ01 (
<taxonomicName class="Insecta" family="Pyralidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Pyralidae</taxonomicName>
). DHJPAR0045384, DHJPAR0043243, DHJPAR0041469, DHJPAR0050021, DHJPAR0051386. Depository: CNC.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Macrocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocentrus harisridhari" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="harisridhari">
<emphasis italics="true" pageId="0" pageNumber="1">Macrocentrus harisridhari</emphasis>
</taxonomicName>
is named to honor Dr. Hari Sridhari of the Indian Institute of Science, Bangalore, India for his efforts to popularize and explain, as a science writer, the processes behind some of
<normalizedToken originalValue="DHJ’s">DHJ's</normalizedToken>
papers.
</paragraph>
<caption doi="10.3897/zookeys.1013.55600.figure305" httpUri="https://binary.pensoft.net/fig/503760" pageId="0" pageNumber="1" start="Figure 305" startId="F305">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 305.</emphasis>
<taxonomicName authorityName="Sharkey &amp; van Achterberg" authorityYear="2021" class="Insecta" family="Braconidae" genus="Macrocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocentrus harisridhari" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="harisridhari">
<emphasis italics="true" pageId="0" pageNumber="1">Macrocentrus harisridhari</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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