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<document id="C7CACE37414340B24AC758AD88D4152B" ID-DOI="10.5252/geodiversitas2018v40a6" ID-ISSN="1638-9395" ID-Zenodo-Dep="5745694" ID-ZooBank="urn:lsid:zoobank.org:pub:06EB756D-EE16-4B28-A09C-EA983B758397" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630284245038" checkinUser="marcus" docAuthor="Lambert, Olivier, Muizon, Christian de, Duhamel, Guy &amp; Plicht, Johannes Van Der" docDate="2018" docId="2C4687B9BD517A64FC90A9F5FA81FA1B" docLanguage="en" docName="Geodiversitas.40.6.135-160.pdf.imf" docOrigin="Geodiversitas 40 (6)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Izikoziphius rossi Bianucci, Lambert &amp; Post 2007" docType="treatment" docVersion="3" lastPageNumber="151" masterDocId="D07FFFC1BD417A76FFCCAF67FF84FFB6" masterDocTitle="Neogene and Quaternary fossil remains of beaked whales (Cetacea, Odontoceti, Ziphiidae) from deep-sea deposits off Crozet and Kerguelen islands, Southern Ocean" masterLastPageNumber="160" masterPageNumber="135" pageNumber="149" updateTime="1699213896873" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="CE03A77AACC1EE3D4458C5496F43D74C">Neogene and Quaternary fossil remains of beaked whales (Cetacea, Odontoceti, Ziphiidae) from deep-sea deposits off Crozet and Kerguelen islands, Southern Ocean</mods:title>
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<mods:namePart id="2056EBC1481B0AF8D74E831EA0ACEED8">Lambert, Olivier</mods:namePart>
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<mods:namePart id="5FB480601ECF1AEEA55E427979AB18F8">Muizon, Christian de</mods:namePart>
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<mods:namePart id="943BD5E021BD82477FD27B6E848BE9D7">Duhamel, Guy</mods:namePart>
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<mods:namePart id="3F2CC4ABC2EE786640DE1620A2CA0727">Plicht, Johannes Van Der</mods:namePart>
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<mods:title id="D464E1F98C9C532B22BCDCD5CAC6A580">Geodiversitas</mods:title>
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<mods:date id="1C8CC34F20AC84C56269DE3B5D0474F0">2018</mods:date>
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<mods:number id="78D535CA053E9B35ABC45F05E2625EC1">2018-03-29</mods:number>
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<paragraph id="A45036AFBD517A66FC90A9F5FA04F91A" blockId="16.[860,1408,1682,1709]" box="[860,1408,1682,1709]" pageId="16" pageNumber="149">
<heading id="FF1881C3BD517A66FC90A9F5FA04F91A" box="[860,1408,1682,1709]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="149" reason="2">
<taxonomicName id="63EF4D2CBD517A66FC90A9F5FA04F91A" authority="Bianucci, Lambert &amp; Post, 2007" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[860,1408,1682,1709]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="16" pageNumber="149" phylum="Chordata" rank="species" species="rossi">
<emphasis id="969BEABDBD517A66FC90A9F5FB93F91A" bold="true" box="[860,1047,1682,1709]" italics="true" pageId="16" pageNumber="149">Izikoziphius rossi</emphasis>
<bibRefCitation id="C07E4B5EBD517A66FBD2A9F5FA04F91A" author="BIANUCCI G. &amp; LAMBERT O. &amp; POST K." box="[1054,1408,1682,1709]" pageId="16" pageNumber="149" pagination="5 - 62" refId="ref15041" refString="BIANUCCI G., LAMBERT O. &amp; POST K. 2007. - A high diversity in fossil beaked whales (Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa. Geodiversitas 29: 5 - 62." type="journal article" year="2007">Bianucci, Lambert &amp; Post, 2007</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="A45036AFBD517A66FCE1A9ABFAA8F885" blockId="16.[811,1456,1739,1843]" pageId="16" pageNumber="149">
REFERRED SPECIMEN AND LOCALITY. — Partial craniumMNHN.F.COI8 including rostrum, premaxillary sac fossae, and vertex (
<figureCitation id="3CD42A2ABD517A66FAFDA982FAF7F948" box="[1329,1395,1765,1790]" captionStart="FIG" captionStartId="17.[132,143,2000,2018]" captionTargetBox="[266,1330,202,1987]" captionTargetId="figure-21@17.[793,1336,765,1334]" captionTargetPageId="17" captionText="FIG. 10. — Partial cranium of Izikoziphius rossi (MNHN.F.COI8): A, dorsal view; B, same view with interpretive line drawing; C, detail of the vertex in dorsal view; D, anterodorsal view; E, right lateral view; F, ventral view. Scale bars: A, B, D-F, 100 mm; C, 50 mm." figureDoi="http://doi.org/10.5281/zenodo.5745724" httpUri="https://zenodo.org/record/5745724/files/figure.png" pageId="16" pageNumber="149">Fig. 10</figureCitation>
);
<materialsCitation id="14873CF2BD517A66FA4AA981FBC4F884" latitude="-49.845665" location="Skiff Bank" longLatPrecision="1" longitude="63.324333" pageId="16" pageNumber="149" specimenCount="1" stateProvince="Iles Kerguelen">
geographic coordinates
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,
<geoCoordinate id="C1DB5068BD517A66FBA3A867FB74F8AE" box="[1135,1264,1792,1816]" degrees="63" direction="east" minutes="19" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="27.6" value="63.324333">63°1927.6”E</geoCoordinate>
(
<location id="A1306074BD517A66FB35A867FADBF8AE" LSID="urn:lsid:plazi:treatment:2C4687B9BD517A64FC90A9F5FA81FA1B:A1306074BD517A66FB35A867FADBF8AE" box="[1273,1375,1792,1816]" latitude="-49.845665" longLatPrecision="1" longitude="63.324333" name="Skiff Bank" pageId="16" pageNumber="149" stateProvince="Iles Kerguelen">Skiff Bank</location>
,
<quantity id="63179B4ABD517A66FAAAA867FA2BF8AE" box="[1382,1455,1792,1816]" metricMagnitude="5" metricUnit="m" metricValue="3.9" pageId="16" pageNumber="149" unit="km" value="390.0">390 km</quantity>
SWW to
<collectingRegion id="662BF84DBD517A66FC44A87DFBBDF884" box="[904,1081,1818,1842]" country="French Southern Territories" name="Iles Kerguelen" pageId="16" pageNumber="149">Kerguelen Islands</collectingRegion>
)
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; depth
<quantity id="63179B4ABD517A66FB45A87CFB52F884" box="[1161,1238,1819,1843]" metricMagnitude="3" metricUnit="m" metricValue="2.041" pageId="16" pageNumber="149" unit="m" value="2041.0">2041 m</quantity>
(
<figureCitation id="3CD42A2ABD517A66FB2AA87DFA9AF885" box="[1254,1310,1818,1843]" captionStart="FIG" captionStartId="4.[132,143,1125,1143]" captionTargetBox="[323,1264,214,1081]" captionTargetId="graphics-412@4.[324,1264,214,1081]" captionTargetPageId="4" captionText="FIG. 1. — Locality maps: A, schematic map providing the position of Crozet and Kerguelen islands in the Southern Ocean; B, schematic map of the Crozet and Kerguelen islands area, including the main localities where fossil ziphiid specimens («) described in this work were found via longline fishing,as well as two ODP sites (l). Light grey shading for areas with a water depth lower than 1000 m; dark grey shading for islands and continents." figureDoi="http://doi.org/10.5281/zenodo.5745696" httpUri="https://zenodo.org/record/5745696/files/figure.png" pageId="16" pageNumber="149">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="ECF56524BD517A64FCE1A835FA81FA1B" lastPageId="18" lastPageNumber="151" pageId="16" pageNumber="149" type="description">
<paragraph id="A45036AFBD517A66FCE1A835FB46F8DC" blockId="16.[810,1456,1874,2028]" box="[813,1218,1874,1900]" pageId="16" pageNumber="149">BRIEF DESCRIPTION AND COMPARISON</paragraph>
<paragraph id="A45036AFBD517A66FCE1A816FA2AF85A" blockId="16.[810,1456,1874,2028]" pageId="16" pageNumber="149">
The cross section of the long and robust rostrum (not complete apically) is considerably wider than high for most of the rostral length (
<tableCitation id="E96D0314BD517A66FCB2A8D6FC4FF87A" box="[894,971,1969,1996]" captionStart="TABLE" captionStartId="18.[131,142,219,237]" captionText="TABLE 7. — Measurements (in mm) on the partial crania of Izikoziphius rossi (MNHN.F.COI8, PEM N 3265, and SAM PQ 2086 [from Bianucci et al. 2007]). Abbreviations: e, estimate; +, not complete; , no data." pageId="16" pageNumber="149">Table 7</tableCitation>
). A marked dorsal and lateral expansion of the thickened lateral margin of the rostrum has a maximum extent
</paragraph>
<caption id="F0906627BD507A67FF48A8B7FC46F84E" ID-DOI="http://doi.org/10.5281/zenodo.5745724" ID-Zenodo-Dep="5745724" httpUri="https://zenodo.org/record/5745724/files/figure.png" pageId="17" pageNumber="150" startId="17.[132,143,2000,2018]" targetBox="[266,1330,202,1987]" targetPageId="17">
<paragraph id="A45036AFBD507A67FF48A8B7FC46F84E" blockId="17.[132,1457,2000,2040]" pageId="17" pageNumber="150">
FIG. 10. — Partial cranium of
<taxonomicName id="63EF4D2CBD507A67FEBBA8B7FD85F854" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[375,513,2000,2018]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="17" pageNumber="150" phylum="Chordata" rank="species" species="rossi">
<emphasis id="969BEABDBD507A67FEBBA8B7FD85F854" box="[375,513,2000,2018]" italics="true" pageId="17" pageNumber="150">Izikoziphius rossi</emphasis>
</taxonomicName>
(MNHN.F.COI8):
<emphasis id="969BEABDBD507A67FD5CA8B7FD19F854" bold="true" box="[656,669,2000,2018]" pageId="17" pageNumber="150">A</emphasis>
, dorsal view;
<emphasis id="969BEABDBD507A67FCDCA8B7FC99F854" bold="true" box="[784,797,2000,2018]" pageId="17" pageNumber="150">B</emphasis>
, same view with interpretive line drawing;
<emphasis id="969BEABDBD507A67FBB3A8B7FB09F854" bold="true" box="[1151,1165,2000,2018]" pageId="17" pageNumber="150">C</emphasis>
, detail of the vertex in dorsal view;
<emphasis id="969BEABDBD507A67FF48A881FF16F84E" bold="true" box="[132,146,2022,2040]" pageId="17" pageNumber="150">D</emphasis>
, anterodorsal view;
<emphasis id="969BEABDBD507A67FEF6A881FEC2F84E" bold="true" box="[314,326,2022,2040]" pageId="17" pageNumber="150">E</emphasis>
, right lateral view;
<emphasis id="969BEABDBD507A67FE2FA881FE6AF84E" bold="true" box="[483,494,2022,2040]" pageId="17" pageNumber="150">F</emphasis>
, ventral view. Scale bars: A, B, D-F, 100 mm; C, 50 mm.
</paragraph>
</caption>
<caption id="F0906627BD537A64FF4FAFBCFDCBFEAC" pageId="18" pageNumber="151" startId="18.[131,142,219,237]">
<paragraph id="A45036AFBD537A64FF4FAFBCFDCBFEAC" blockId="18.[127,776,219,282]" pageId="18" pageNumber="151">
TABLE 7. — Measurements (in mm) on the partial crania of
<taxonomicName id="63EF4D2CBD537A64FDB5AFBCFC83FF5B" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[633,775,219,237]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="151" phylum="Chordata" rank="species" species="rossi">
<emphasis id="969BEABDBD537A64FDB5AFBCFC83FF5B" box="[633,775,219,237]" italics="true" pageId="18" pageNumber="151">Izikoziphius rossi</emphasis>
</taxonomicName>
(MNHN.F.COI8, PEM N 3265, and SAM PQ 2086 [from
<bibRefCitation id="C07E4B5EBD537A64FD9EAF95FD7CFEB2" author="BIANUCCI G. &amp; LAMBERT O. &amp; POST K." box="[594,760,242,260]" pageId="18" pageNumber="151" pagination="5 - 62" refId="ref15041" refString="BIANUCCI G., LAMBERT O. &amp; POST K. 2007. - A high diversity in fossil beaked whales (Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa. Geodiversitas 29: 5 - 62." type="journal article" year="2007">
Bianucci
<emphasis id="969BEABDBD537A64FD6CAF95FD4EFEB2" box="[672,714,242,260]" italics="true" pageId="18" pageNumber="151">et al.</emphasis>
2007
</bibRefCitation>
]). Abbreviations:
<emphasis id="969BEABDBD537A64FECDAE6FFE88FEAC" bold="true" box="[257,268,264,282]" pageId="18" pageNumber="151">e</emphasis>
, estimate;
<emphasis id="969BEABDBD537A64FEABAE6FFEF6FEAC" bold="true" box="[359,370,264,282]" pageId="18" pageNumber="151">+</emphasis>
, not complete;
<emphasis id="969BEABDBD537A64FE39AE6FFE7AFEAC" bold="true" box="[501,510,264,282]" pageId="18" pageNumber="151"></emphasis>
, no data.
</paragraph>
</caption>
<paragraph id="A45036AFBD537A64FD9BAE33FD5BFCF2" pageId="18" pageNumber="151">
<table id="D6EFC40FBD538589FF4AAE33FC87FCF2" box="[134,771,340,836]" gridcols="4" gridrows="20" pageId="18" pageNumber="151">
<tr id="1ADF34EDBD538589FF4AAE33FC87FEDE" box="[134,771,340,360]" gridrow="0" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAE33FC87FEDE" box="[134,771,340,360]" colspan="4" colspanRight="3" gridcol="0" gridrow="0" pageId="18" pageNumber="151">
<emphasis id="969BEABDBD537A64FD9BAE33FD18FEDE" bold="true" box="[599,668,340,360]" pageId="18" pageNumber="151">PEM N</emphasis>
</th>
</tr>
<tr id="1ADF34EDBD538589FF4AAE0AFC87FE37" box="[134,771,365,385]" gridrow="1" pageId="18" pageNumber="151" rowspan-0="1">
<td id="590E5D91BD538589FE23AE0AFD1DFE37" box="[495,665,365,385]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="18" pageNumber="151">
<emphasis id="969BEABDBD537A64FE23AE0AFD15FE37" bold="true" box="[495,657,365,385]" pageId="18" pageNumber="151">MNHN.F. 3265</emphasis>
</td>
<td id="590E5D91BD538589FD62AE0AFC87FE37" box="[686,771,365,385]" gridcol="3" gridrow="1" pageId="18" pageNumber="151">
<emphasis id="969BEABDBD537A64FD62AE0AFC87FE37" bold="true" box="[686,771,365,385]" pageId="18" pageNumber="151">SAM PQ</emphasis>
</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAEE0FC87FE2D" box="[134,771,391,411]" gridrow="2" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAEE0FE64FE2D" box="[134,480,391,411]" gridcol="0" gridrow="2" pageId="18" pageNumber="151">
<taxonomicName id="63EF4D2CBD537A64FF4AAEE0FEB7FE2D" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[134,307,391,411]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="151" phylum="Chordata" rank="species" species="rossi">
<emphasis id="969BEABDBD537A64FF4AAEE0FEB7FE2D" bold="true" box="[134,307,391,411]" italics="true" pageId="18" pageNumber="151">Izikoziphius rossi</emphasis>
</taxonomicName>
</th>
<td id="590E5D91BD538589FE23AEE0FDB0FE2D" box="[495,564,391,411]" gridcol="1" gridrow="2" pageId="18" pageNumber="151">
<emphasis id="969BEABDBD537A64FDCDAEE0FDB0FE2D" bold="true" box="[513,564,391,411]" pageId="18" pageNumber="151">COI8</emphasis>
</td>
<td id="590E5D91BD538589FD9CAEE0FD1DFE2D" box="[592,665,391,411]" gridcol="2" gridrow="2" pageId="18" pageNumber="151">
<emphasis id="969BEABDBD537A64FD96AEE0FD1DFE2D" bold="true" box="[602,665,391,411]" pageId="18" pageNumber="151">(holo.)</emphasis>
</td>
<td id="590E5D91BD538589FD62AEE0FC87FE2D" box="[686,771,391,411]" gridcol="3" gridrow="2" pageId="18" pageNumber="151">
<emphasis id="969BEABDBD537A64FD0CAEE0FD6BFE2D" bold="true" box="[704,751,391,411]" pageId="18" pageNumber="151">2086</emphasis>
</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAECDFC87FE08" box="[134,771,426,446]" gridrow="3" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAECDFE64FE08" box="[134,480,426,446]" gridcol="0" gridrow="3" pageId="18" pageNumber="151">Rostrum length</th>
<td id="590E5D91BD538589FE23AECDFDB0FE08" box="[495,564,426,446]" gridcol="1" gridrow="3" pageId="18" pageNumber="151">e495</td>
<td id="590E5D91BD538589FD9CAECDFD1DFE08" box="[592,665,426,446]" gridcol="2" gridrow="3" pageId="18" pageNumber="151">+490</td>
<td id="590E5D91BD538589FD62AECDFC87FE08" box="[686,771,426,446]" gridcol="3" gridrow="3" pageId="18" pageNumber="151">e530</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAEA5FC87FE61" box="[134,771,450,471]" gridrow="4" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAEA5FE64FE61" box="[134,480,450,471]" gridcol="0" gridrow="4" pageId="18" pageNumber="151">Height of rostrum at mid-length</th>
<td id="590E5D91BD538589FE23AEA5FDB0FE61" box="[495,564,450,471]" gridcol="1" gridrow="4" pageId="18" pageNumber="151">72</td>
<td id="590E5D91BD538589FD9CAEA5FD1DFE61" box="[592,665,450,471]" gridcol="2" gridrow="4" pageId="18" pageNumber="151">+100</td>
<td id="590E5D91BD538589FD62AEA5FC87FE61" box="[686,771,450,471]" gridcol="3" gridrow="4" pageId="18" pageNumber="151">84</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAEBCFC87FE59" box="[134,771,475,495]" gridrow="5" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAEBCFE64FE59" box="[134,480,475,495]" gridcol="0" gridrow="5" pageId="18" pageNumber="151">Width of rostrum at mid-length</th>
<td id="590E5D91BD538589FE23AEBCFDB0FE59" box="[495,564,475,495]" gridcol="1" gridrow="5" pageId="18" pageNumber="151">106</td>
<td id="590E5D91BD538589FD9CAEBCFD1DFE59" box="[592,665,475,495]" gridcol="2" gridrow="5" pageId="18" pageNumber="151">115</td>
<td id="590E5D91BD538589FD62AEBCFC87FE59" box="[686,771,475,495]" gridcol="3" gridrow="5" pageId="18" pageNumber="151">94</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAE94FC87FDB1" box="[134,771,499,519]" gridrow="6" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAE94FE64FDB1" box="[134,480,499,519]" gridcol="0" gridrow="6" pageId="18" pageNumber="151">Maximum width of rostrum at</th>
<td id="590E5D91BD538589FE23AE94FDB0FDB1" box="[495,564,499,519]" gridcol="1" gridrow="6" pageId="18" pageNumber="151">155</td>
<td id="590E5D91BD538589FD9CAE94FD1DFDB1" box="[592,665,499,519]" gridcol="2" gridrow="6" pageId="18" pageNumber="151"></td>
<td id="590E5D91BD538589FD62AE94FC87FDB1" box="[686,771,499,519]" gridcol="3" gridrow="6" pageId="18" pageNumber="151"></td>
</tr>
<tr id="1ADF34EDBD538589FF4AAD6BFC87FD96" box="[134,771,524,544]" gridrow="7" pageId="18" pageNumber="151" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="590E5D91BD538589FF4AAD6BFE64FD96" box="[134,480,524,544]" gridcol="0" gridrow="7" pageId="18" pageNumber="151">dorsolateral expansion</th>
</tr>
<tr id="1ADF34EDBD538589FF4AAD43FC87FD8E" box="[134,771,548,568]" gridrow="8" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAD43FE64FD8E" box="[134,480,548,568]" gridcol="0" gridrow="8" pageId="18" pageNumber="151">Width of premaxillary sac fossae</th>
<td id="590E5D91BD538589FE23AD43FDB0FD8E" box="[495,564,548,568]" gridcol="1" gridrow="8" pageId="18" pageNumber="151">131</td>
<td id="590E5D91BD538589FD9CAD43FD1DFD8E" box="[592,665,548,568]" gridcol="2" gridrow="8" pageId="18" pageNumber="151">145</td>
<td id="590E5D91BD538589FD62AD43FC87FD8E" box="[686,771,548,568]" gridcol="3" gridrow="8" pageId="18" pageNumber="151">155</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAD5BFC87FDE7" box="[134,771,572,593]" gridrow="9" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAD5BFE64FDE7" box="[134,480,572,593]" gridcol="0" gridrow="9" pageId="18" pageNumber="151">Width of right premaxillary sac fossa</th>
<td id="590E5D91BD538589FE23AD5BFDB0FDE7" box="[495,564,572,593]" gridcol="1" gridrow="9" pageId="18" pageNumber="151">72</td>
<td id="590E5D91BD538589FD9CAD5BFD1DFDE7" box="[592,665,572,593]" gridcol="2" gridrow="9" pageId="18" pageNumber="151">75</td>
<td id="590E5D91BD538589FD62AD5BFC87FDE7" box="[686,771,572,593]" gridcol="3" gridrow="9" pageId="18" pageNumber="151">80</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAD32FC87FDDF" box="[134,771,597,617]" gridrow="10" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAD32FE64FDDF" box="[134,480,597,617]" gridcol="0" gridrow="10" pageId="18" pageNumber="151">Width of left premaxillary sac fossa</th>
<td id="590E5D91BD538589FE23AD32FDB0FDDF" box="[495,564,597,617]" gridcol="1" gridrow="10" pageId="18" pageNumber="151">58</td>
<td id="590E5D91BD538589FD9CAD32FD1DFDDF" box="[592,665,597,617]" gridcol="2" gridrow="10" pageId="18" pageNumber="151">55</td>
<td id="590E5D91BD538589FD62AD32FC87FDDF" box="[686,771,597,617]" gridcol="3" gridrow="10" pageId="18" pageNumber="151">62</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAD0AFC87FD37" box="[134,771,621,641]" gridrow="11" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAD0AFE64FD37" box="[134,480,621,641]" gridcol="0" gridrow="11" pageId="18" pageNumber="151">Width of bony nares</th>
<td id="590E5D91BD538589FE23AD0AFDB0FD37" box="[495,564,621,641]" gridcol="1" gridrow="11" pageId="18" pageNumber="151">60</td>
<td id="590E5D91BD538589FD9CAD0AFD1DFD37" box="[592,665,621,641]" gridcol="2" gridrow="11" pageId="18" pageNumber="151">56</td>
<td id="590E5D91BD538589FD62AD0AFC87FD37" box="[686,771,621,641]" gridcol="3" gridrow="11" pageId="18" pageNumber="151">73</td>
</tr>
<tr id="1ADF34EDBD538589FF4AADE2FC87FD2F" box="[134,771,645,665]" gridrow="12" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AADE2FE64FD2F" box="[134,480,645,665]" gridcol="0" gridrow="12" pageId="18" pageNumber="151">Minimum width of right ascending</th>
<td id="590E5D91BD538589FE23ADE2FDB0FD2F" box="[495,564,645,665]" gridcol="1" gridrow="12" pageId="18" pageNumber="151">48</td>
<td id="590E5D91BD538589FD9CADE2FD1DFD2F" box="[592,665,645,665]" gridcol="2" gridrow="12" pageId="18" pageNumber="151"></td>
<td id="590E5D91BD538589FD62ADE2FC87FD2F" box="[686,771,645,665]" gridcol="3" gridrow="12" pageId="18" pageNumber="151">45</td>
</tr>
<tr id="1ADF34EDBD538589FF4AADF9FC87FD04" box="[134,771,670,690]" gridrow="13" pageId="18" pageNumber="151" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="590E5D91BD538589FF4AADF9FE64FD04" box="[134,480,670,690]" gridcol="0" gridrow="13" pageId="18" pageNumber="151">process of premaxilla</th>
</tr>
<tr id="1ADF34EDBD538589FF4AADD1FC87FD7C" box="[134,771,694,714]" gridrow="14" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AADD1FE64FD7C" box="[134,480,694,714]" gridcol="0" gridrow="14" pageId="18" pageNumber="151">Width of premaxillary crests</th>
<td id="590E5D91BD538589FE23ADD1FDB0FD7C" box="[495,564,694,714]" gridcol="1" gridrow="14" pageId="18" pageNumber="151">147</td>
<td id="590E5D91BD538589FD9CADD1FD1DFD7C" box="[592,665,694,714]" gridcol="2" gridrow="14" pageId="18" pageNumber="151">129</td>
<td id="590E5D91BD538589FD62ADD1FC87FD7C" box="[686,771,694,714]" gridcol="3" gridrow="14" pageId="18" pageNumber="151">+126</td>
</tr>
<tr id="1ADF34EDBD538589FF4AADA9FC87FD55" box="[134,771,718,739]" gridrow="15" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AADA9FE64FD55" box="[134,480,718,739]" gridcol="0" gridrow="15" pageId="18" pageNumber="151">Width of right premaxillary crest</th>
<td id="590E5D91BD538589FE23ADA9FDB0FD55" box="[495,564,718,739]" gridcol="1" gridrow="15" pageId="18" pageNumber="151">45</td>
<td id="590E5D91BD538589FD9CADA9FD1DFD55" box="[592,665,718,739]" gridcol="2" gridrow="15" pageId="18" pageNumber="151">71</td>
<td id="590E5D91BD538589FD62ADA9FC87FD55" box="[686,771,718,739]" gridcol="3" gridrow="15" pageId="18" pageNumber="151"></td>
</tr>
<tr id="1ADF34EDBD538589FF4AAD80FC87FD4D" box="[134,771,743,763]" gridrow="16" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAD80FE64FD4D" box="[134,480,743,763]" gridcol="0" gridrow="16" pageId="18" pageNumber="151">Width of left premaxillary crest</th>
<td id="590E5D91BD538589FE23AD80FDB0FD4D" box="[495,564,743,763]" gridcol="1" gridrow="16" pageId="18" pageNumber="151">36</td>
<td id="590E5D91BD538589FD9CAD80FD1DFD4D" box="[592,665,743,763]" gridcol="2" gridrow="16" pageId="18" pageNumber="151">32</td>
<td id="590E5D91BD538589FD62AD80FC87FD4D" box="[686,771,743,763]" gridcol="3" gridrow="16" pageId="18" pageNumber="151"></td>
</tr>
<tr id="1ADF34EDBD538589FF4AAD98FC87FCA5" box="[134,771,767,787]" gridrow="17" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAD98FE64FCA5" box="[134,480,767,787]" gridcol="0" gridrow="17" pageId="18" pageNumber="151">Minimum distance between</th>
<td id="590E5D91BD538589FE23AD98FDB0FCA5" box="[495,564,767,787]" gridcol="1" gridrow="17" pageId="18" pageNumber="151">54.5</td>
<td id="590E5D91BD538589FD9CAD98FD1DFCA5" box="[592,665,767,787]" gridcol="2" gridrow="17" pageId="18" pageNumber="151"></td>
<td id="590E5D91BD538589FD62AD98FC87FCA5" box="[686,771,767,787]" gridcol="3" gridrow="17" pageId="18" pageNumber="151">69</td>
</tr>
<tr id="1ADF34EDBD538589FF4AAC7FFC87FC9A" box="[134,771,792,812]" gridrow="18" pageId="18" pageNumber="151" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="590E5D91BD538589FF4AAC7FFE64FC9A" box="[134,480,792,812]" gridcol="0" gridrow="18" pageId="18" pageNumber="151">premaxillary crests</th>
</tr>
<tr id="1ADF34EDBD538589FF4AAC57FC87FCF2" box="[134,771,816,836]" gridrow="19" pageId="18" pageNumber="151">
<th id="590E5D91BD538589FF4AAC57FE64FCF2" box="[134,480,816,836]" gridcol="0" gridrow="19" pageId="18" pageNumber="151">Maximum width of nasals</th>
<td id="590E5D91BD538589FE23AC57FDB0FCF2" box="[495,564,816,836]" gridcol="1" gridrow="19" pageId="18" pageNumber="151">83</td>
<td id="590E5D91BD538589FD9CAC57FD1DFCF2" box="[592,665,816,836]" gridcol="2" gridrow="19" pageId="18" pageNumber="151"></td>
<td id="590E5D91BD538589FD62AC57FC87FCF2" box="[686,771,816,836]" gridcol="3" gridrow="19" pageId="18" pageNumber="151">e88</td>
</tr>
</table>
</paragraph>
<paragraph id="A45036AFBD537A64FF48ACF2FD35F99A" blockId="18.[130,777,917,2028]" pageId="18" pageNumber="151">
at about mid rostral length, resulting in the lateral margin of the rostrum being distinctly convex in dorsal view in that region. The mesorostral groove is filled by the thickened vomer until the level of the antorbital notches, and the dorsal surface of the vomer is transversely concave in the apical part of the rostrum. On the ventral surface of the rostrum, lateral to the ventral exposure of the vomer the maxillae are pierced by several large palatal foramina, anteriorly followed by deep sulci. The rostrum is broad at its base, with a depressed medial region laterally margined by the rostral maxillary crest. The latter progressively raises past the antorbital notch level, forming a high hump in the antorbital region, best seen in lateral view. Medial to the rostral maxillary crest, a large dorsal infraorbital foramen is followed anteriorly by a wide depression. Several smaller foramina pierce the rostral maxillary crests. The premaxillary foramen is roughly in line with the large dorsal infraorbital foramen, both being posterior to the estimated level of the antorbital notch. The two premaxillary sac fossae are moderately transversely concave, and the right fossa is distinctly wider than the left (
<tableCitation id="E96D0314BD537A64FF41AA94FF5FF9BB" box="[141,219,1523,1549]" captionStart="TABLE" captionStartId="18.[131,142,219,237]" captionText="TABLE 7. — Measurements (in mm) on the partial crania of Izikoziphius rossi (MNHN.F.COI8, PEM N 3265, and SAM PQ 2086 [from Bianucci et al. 2007]). Abbreviations: e, estimate; +, not complete; , no data." pageId="18" pageNumber="151">Table 7</tableCitation>
). The lateral margin of each ascending process of the premaxilla is moderately constricted in anterior view.
</paragraph>
<paragraph id="A45036AFBD537A64FF57A955FD35F95A" blockId="18.[130,777,917,2028]" pageId="18" pageNumber="151">The vertex of the cranium is elevated, with the anterior surface of the ascending process of the premaxilla reaching the vertical. This anterior surface is excavated on both sides by a deep fossa, extending somewhat on the lateral part of the corresponding nasal.Each premaxillary crest is directed anterolaterally, and the right crest is wider and dorsally higher than left crest.</paragraph>
<paragraph id="A45036AFBD537A64FF57A995FDF0F81D" blockId="18.[130,777,917,2028]" pageId="18" pageNumber="151">Both nasals are longer than wide, extending anteriorly beyond the level of the corresponding premaxillary crest. The right nasal is significantly longer posteriorly than the left. From their anterior apices, the lateral margins of the right and left nasals diverge abruptly until mid-length. The median region of the dorsal surface of the nasals is depressed.</paragraph>
<paragraph id="A45036AFBD537A64FF57A8D6FA0CFD26" blockId="18.[130,777,917,2028]" lastBlockId="18.[811,1458,215,1454]" pageId="18" pageNumber="151">
The combination of a mesorostral groove filled with the vomer, a high vertex with anterolaterally directed premaxil- lary crests, and anteriorly elongated nasals point to a ziphiine attribution (
<emphasis id="969BEABDBD537A64FC74AF9FFC6AFEA7" box="[952,1006,248,273]" italics="true" pageId="18" pageNumber="151">sensu</emphasis>
<bibRefCitation id="C07E4B5EBD537A64FC34AF90FB5EFEA7" author="BIANUCCI G. &amp; DI CELMA C. &amp; URBINA M. &amp; LAMBERT O." box="[1016,1242,247,273]" pageId="18" pageNumber="151" pagination="2479" refId="ref15205" refString="BIANUCCI G., DI CELMA C., URBINA M. &amp; LAMBERT O. 2016. - New beaked whales from the late Miocene of Peru and evidence for convergent evolution in stem and crown Ziphiidae (Cetacea, Odontoceti). PeerJ 4: e 2479. https: // doi. org / 10.7717 / peerj. 2479" type="journal article" year="2016">
Bianucci
<emphasis id="969BEABDBD537A64FBADAF9FFB1CFEA7" box="[1121,1176,247,273]" italics="true" pageId="18" pageNumber="151">et al.</emphasis>
2016
</bibRefCitation>
). Other characters (e.g. the long rostrum, the absence of a prenarial basin, the fossa on the anterior surface of the ascending process of the premaxilla, and the relatively long contact between nasal and premaxilla) contrast with
<taxonomicName id="63EF4D2CBD537A64FB85AE10FB1CFE27" authorityName="Cuvier" authorityYear="1823" box="[1097,1176,375,401]" class="Mammalia" family="Hyperoodontidae" genus="Ziphius" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="151" phylum="Chordata" rank="genus">
<emphasis id="969BEABDBD537A64FB85AE10FB1CFE27" box="[1097,1176,375,401]" italics="true" pageId="18" pageNumber="151">Ziphius</emphasis>
</taxonomicName>
(adult males of the latter for the prenarial basin) and unambiguously indicate affinities with
<taxonomicName id="63EF4D2CBD537A64FCA8AED1FC66FE66" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[868,994,438,464]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="151" phylum="Chordata" rank="genus">
<emphasis id="969BEABDBD537A64FCA8AED1FC66FE66" box="[868,994,438,464]" italics="true" pageId="18" pageNumber="151">Izikoziphius</emphasis>
</taxonomicName>
. Within the latter genus, MNHN.F.COI8 shares several characters with
<taxonomicName id="63EF4D2CBD537A64FB93AEB1FA8FFE46" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[1119,1291,470,496]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="151" phylum="Chordata" rank="species" species="rossi">
<emphasis id="969BEABDBD537A64FB93AEB1FA8FFE46" box="[1119,1291,470,496]" italics="true" pageId="18" pageNumber="151">Izikoziphius rossi</emphasis>
</taxonomicName>
: rostrum wider than high at mid-length, rostral maxillary crest extending on rostrum base, large fossa on the anterior surface of the ascending process of the premaxilla extending on the corresponding nasal, median depression of the dorsal surface of the nasals, and right premaxillary crest much higher than left crest.
</paragraph>
<paragraph id="A45036AFBD537A64FC8FADF1FAD9FC19" blockId="18.[811,1458,215,1454]" pageId="18" pageNumber="151">
However, MNHN.F.COI8 differs from the two currently known specimens of
<taxonomicName id="63EF4D2CBD537A64FBCBADD1FBC3FD79" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[1031,1095,694,719]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="151" phylum="Chordata" rank="species" species="rossi">
<emphasis id="969BEABDBD537A64FBCBADD1FBC3FD79" box="[1031,1095,694,719]" italics="true" pageId="18" pageNumber="151">I. rossi</emphasis>
</taxonomicName>
in the marked dorsolateral expansion of the lateral margin of the rostrum at mid-length. Pending the discovery of additional specimens this conspicuous difference is interpreted here as intraspecific variation, maybe indicating sexual dimorphism as recorded in several extant ziphiids and inferred in a few extinct species (
<bibRefCitation id="C07E4B5EBD537A64FAC0AC32FA2CFCD9" author="HEYNING J. E." box="[1292,1448,853,879]" pageId="18" pageNumber="151" pagination="1 - 64" refId="ref15852" refString="HEYNING J. E. 1989. - Comparative facial anatomy of beaked whales (Ziphiidae) and a systematic revision among the families of extant Odontoceti. Contributions in Science, Natural History Museum of Los Angeles County 405: 1 - 64." type="journal article" year="1989">Heyning 1989</bibRefCitation>
;
<bibRefCitation id="C07E4B5EBD537A64FCE1AC12FC31FC39" author="MEAD J. G." box="[813,949,885,911]" pageId="18" pageNumber="151" pagination="321 - 348" refId="ref16417" refString="MEAD J. G. 1989 b. - Bottlenose whales Hyperoodon ampullatus (Forster, 1770) and Hyperoodon planifrons Flower, 1882, in RIDGWAY S. H. &amp; HARRISON R. (eds), Handbook of Marine Mammals, Vol. 4: River Dolphins and the Larger Toothed Whales. Academic Press, London: 321 - 348." type="book chapter" year="1989">Mead 1989b</bibRefCitation>
;
<bibRefCitation id="C07E4B5EBD537A64FC0EAC12FB17FC39" author="BIANUCCI G. &amp; MIJAN I. &amp; LAMBERT O. &amp; POST K. &amp; MATEUS O." box="[962,1171,885,911]" pageId="18" pageNumber="151" pagination="105 - 153" refId="ref15141" refString="BIANUCCI G., MIJAN I., LAMBERT O., POST K. &amp; MATEUS O. 2013. - Bizarre fossil beaked whales (Odontoceti, Ziphiidae) fished from the Atlantic Ocean floor off the Iberian Peninsula. Geodiversitas 35: 105 - 153. https: // doi. org / 10.5252 / g 2013 n 1 a 6" type="journal article" year="2013">
Bianucci
<emphasis id="969BEABDBD537A64FBE9AC11FBD2FC39" box="[1061,1110,885,911]" italics="true" pageId="18" pageNumber="151">et al.</emphasis>
2013
</bibRefCitation>
). It could nevertheless also indicate a closely related new species of
<taxonomicName id="63EF4D2CBD537A64FB15ACF2FAD3FC19" authorityName="Bianucci, Lambert &amp; Post" authorityYear="2007" box="[1241,1367,917,943]" class="Mammalia" family="Hyperoodontidae" genus="Izikoziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="151" phylum="Chordata" rank="genus">
<emphasis id="969BEABDBD537A64FB15ACF2FAD3FC19" box="[1241,1367,917,943]" italics="true" pageId="18" pageNumber="151">Izikoziphius</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="A45036AFBD537A64FC8FACD2FA81FA1B" blockId="18.[811,1458,215,1454]" pageId="18" pageNumber="151">
Interestingly, although much less developed this expansion of the lateral margin of the rostrum is to some degree reminiscent of the huge rostral maxillary crests observed in SAM PQ 2717 and 2719,
<specimenCount id="B2E9FD26BD537A64FBC2AB72FB28FB98" box="[1038,1196,1044,1071]" pageId="18" pageNumber="151" type="generic">two specimens</specimenCount>
from the South African fauna(s) previously attributed to
<taxonomicName id="63EF4D2CBD537A64FB55AB53FA93FBF8" authorityName="Flower" authorityYear="1867" box="[1177,1303,1076,1102]" class="Mammalia" kingdom="Animalia" order="Cetacea" pageId="18" pageNumber="138" phylum="Chordata" rank="subOrder" subOrder="Odontoceti">Odontoceti</taxonomicName>
indet. due to their fragmentary state of preservation (
<bibRefCitation id="C07E4B5EBD537A64FB02AB33FA1BFBD8" author="BIANUCCI G. &amp; LAMBERT O. &amp; POST K." box="[1230,1439,1108,1134]" pageId="18" pageNumber="151" pagination="5 - 62" refId="ref15041" refString="BIANUCCI G., LAMBERT O. &amp; POST K. 2007. - A high diversity in fossil beaked whales (Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa. Geodiversitas 29: 5 - 62." type="journal article" year="2007">
Bianucci
<emphasis id="969BEABDBD537A64FAFDAB32FAE7FBD8" box="[1329,1379,1108,1134]" italics="true" pageId="18" pageNumber="151">et al.</emphasis>
2007
</bibRefCitation>
). In addition to rostral maxillary crests with the same position and orientation as the expansion of the lateral margin of the rostrum in MNHN.F.COI8, SAM PQ2717 and 2719 share with the latter a thickened vomer in the mesorostral groove, the dorsal surface of the vomer being transversely concave in the apical region of the rostrum, and several large palatal foramina.Together with the high compactness of rostral bones, all these similarities point to ziphiid (and maybe even ziphiine) rather than physeterid affinities for these two mysterious specimens (G. Bianucci pers. comm. 2017).
</paragraph>
</subSubSection>
</treatment>
</document>