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<document id="E2AC8A252E5DC219DD41F5A181FBFF73" ID-CLB-Dataset="7905" ID-DOI="10.11646/zootaxa.4877.2.7" ID-GBIF-Dataset="8e61eca8-4bf7-4fba-80e9-adad7179d4f3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4576567" ID-ZooBank="212908EB-EC38-493E-BBA7-245D10AB5A06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610029642077" checkinUser="plazi" docAuthor="Carvajal-Cogollo, Juan E., Eguis-Avendaño, Jorge A. &amp; Meza-Joya, Fabio Leonardo" docDate="2020" docId="2C245E37FFFA023DFF00FF0AFD40FF28" docLanguage="en" docName="zootaxa.4877.2.7.pdf" docOrigin="Zootaxa 4877 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gonatodes castanae Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="351" masterDocId="D01D264FFFFE023AFF97FFD5FF98FFFD" masterDocTitle="A new diurnal gecko in the genus Gonatodes (Squamata: Sphaerodactylidae) from the foothills of the Serranía de San Lucas, Norosí-Colombia" masterLastPageNumber="360" masterPageNumber="345" pageNumber="349" updateTime="1698923493878" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1BC5681F9F501659C3691DF4BF35F573">A new diurnal gecko in the genus Gonatodes (Squamata: Sphaerodactylidae) from the foothills of the Serranía de San Lucas, Norosí-Colombia</mods:title>
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</mods:role>
<mods:namePart id="C6CB8AAF37F7C17EE59803B9679BB63A">Carvajal-Cogollo, Juan E.</mods:namePart>
<mods:nameIdentifier id="E9FB7DF35AC3250B5F77436653069332" type="ORCID">0000-0002-4542-6967</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="DFFD8EB159C58C1BA74E05CFABDE9464">Eguis-Avendaño, Jorge A.</mods:namePart>
<mods:nameIdentifier id="758B83849821BC01592AC0844909E4B4" type="ORCID">0000-0002-2620-199X</mods:nameIdentifier>
<mods:affiliation id="7CA09E38471BD8BE51D7671BC16D3199">https: // orcid. org / 0000 - 0002 - 2620 - 199 X</mods:affiliation>
</mods:name>
<mods:name id="5DBBF519D12A2E5AF2DD664A139EF871" type="personal">
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<mods:roleTerm id="8730C6837618CAADE9A33CBDD6F4A3CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6024847C5F67D6CE9CE62BB848F46CB8">Meza-Joya, Fabio Leonardo</mods:namePart>
<mods:nameIdentifier id="75F13A79FE3E418D9C78AFC04E3D9B62" type="ORCID">0000-0003-4465-4471</mods:nameIdentifier>
<mods:affiliation id="F677C6BD4E467AEE99300D7A6A9A1FF4">Colombia Endémica, Asociación para el Estudio y la Conservación de los Recursos Naturales, Bucaramanga, Santander, Colombia Grupo de Investigación en Biotecnología Industrial y Biología Molecular, Escuela de Biología, Universidad Industrial de Santander, Piedecuesta, Santander, Colombia. https: // orcid. org / 0000 - 0003 - 4465 - 4471</mods:affiliation>
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<mods:title id="972C95C30307E4C12CF7F335A33B817C">Zootaxa</mods:title>
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<mods:date id="1A1E2CF2DE22E3606E3C0DB6F0A49E61">2020</mods:date>
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<mods:number id="25EF094C61D90C6A606968A1470118BF">2020-11-10</mods:number>
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<mods:number id="B50B6F26C226F549AEBA6E5ABA558EEA">4877</mods:number>
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<mods:identifier id="DA509DBA5C833EA4406B97B933A44F33" type="DOI">10.11646/zootaxa.4877.2.7</mods:identifier>
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<treatment id="2C245E37FFFA023DFF00FF0AFD40FF28" ID-DOI="http://doi.org/10.5281/zenodo.4576631" ID-GBIF-Taxon="179987067" ID-Zenodo-Dep="4576631" LSID="urn:lsid:plazi:treatment:2C245E37FFFA023DFF00FF0AFD40FF28" httpUri="http://treatment.plazi.org/id/2C245E37FFFA023DFF00FF0AFD40FF28" lastPageId="7" lastPageNumber="351" pageId="4" pageNumber="349">
<subSubSection id="EC97BCAAFFFA023EFF00FF0AFE64FEE0" pageId="4" pageNumber="349" type="nomenclature">
<paragraph id="A432EF21FFFA023EFF00FF0AFE76FF04" blockId="4.[151,511,223,357]" box="[151,494,223,250]" pageId="4" pageNumber="349">
<heading id="FF7A584DFFFA023EFF00FF0AFE76FF04" bold="true" box="[151,494,223,250]" fontSize="11" level="1" pageId="4" pageNumber="349" reason="1">
<emphasis id="96F93333FFFA023EFF00FF0AFE76FF04" bold="true" box="[151,494,223,250]" pageId="4" pageNumber="349">
<taxonomicName id="638D94A2FFFA023EFF00FF0AFE11FF04" authority="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya, 2020" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[151,393,223,250]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae" status="sp. nov.">
<emphasis id="96F93333FFFA023EFF00FF0AFE11FF04" bold="true" box="[151,393,223,250]" italics="true" pageId="4" pageNumber="349">Gonatodes castanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DCA8E48FFFA023EFE06FF0AFE76FF04" box="[401,494,223,249]" pageId="4" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A432EF21FFFA023EFF00FED6FE64FEE0" blockId="4.[151,511,223,357]" box="[151,508,259,286]" pageId="4" pageNumber="349">English name: Castaño’s Gecko</paragraph>
</subSubSection>
<subSubSection id="EC97BCAAFFFA023EFF00FEF2FEB7FE98" pageId="4" pageNumber="349" type="description">
<paragraph id="A432EF21FFFA023EFF00FEF2FE67FEBC" blockId="4.[151,511,223,357]" box="[151,511,295,322]" pageId="4" pageNumber="349">Spanish name. Geco de Castaño</paragraph>
<paragraph id="A432EF21FFFA023EFF00FE9EFEB7FE98" blockId="4.[151,511,223,357]" box="[151,303,331,357]" pageId="4" pageNumber="349">
(
<figureCitation id="3CB6F3A4FFFA023EFF08FE9EFEBEFE98" box="[159,294,331,357]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1942,1966]" captionStartId-1="5.[151,250,1982,2006]" captionStartId-2="7.[151,250,1171,1195]" captionTargetBox-0="[183,1404,316,1913]" captionTargetBox-1="[198,1388,713,1948]" captionTargetBox-2="[151,1436,237,1148]" captionTargetId-0="figure-78@3.[175,1413,309,1919]" captionTargetId-1="figure-299@5.[190,1398,705,1958]" captionTargetId-2="figure-48@7.[151,1436,237,1148]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="FIGURE 1. Maximum likelihood tree of Gonatodes showing the phylogenetic position of G. castanae sp. nov. Support values for well-supported nodes correspond to ML ultrafast bootstrap (&gt;95) and Bayesian posterior probabilities (&gt;0.95), respectively. Picture of the male holotype (UPTC-Re 1010). Photo by Jorge Andrés Eguis-Avendaño." captionText-1="FIGURE 2. Scale pattern of Gonatodes castanae sp. nov. (male holotype, UPTC-Re 1010). Subcaudal scale pattern Type B (A); escutcheon (B); scale pattern in the gular area (C). Illustrations by Ariadna Valenzuela." captionText-2="FIGURE 3. Color pattern of Gonatodes castanae sp. nov. UPTC-Re 1010 male holotype (A); UPTC-Re 1011 male paratype (B); UPTC-Re 1012 female paratype (C); UPTC-Re 1013 male paratype (D). Photos by Jorge Andrés Eguis-Avendaño." figureDoi-0="http://doi.org/10.5281/zenodo.4576569" figureDoi-1="http://doi.org/10.5281/zenodo.4576571" figureDoi-2="http://doi.org/10.5281/zenodo.4576575" httpUri-0="https://zenodo.org/record/4576569/files/figure.png" httpUri-1="https://zenodo.org/record/4576571/files/figure.png" httpUri-2="https://zenodo.org/record/4576575/files/figure.png" pageId="4" pageNumber="349">Figures 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EC97BCAAFFFA023EFF00FE46FA0EFDE4" pageId="4" pageNumber="349" type="materials_examined">
<paragraph id="A432EF21FFFA023EFF00FE46FDD3FE0B" blockId="4.[151,1437,403,2014]" pageId="4" pageNumber="349">
<materialsCitation id="14E5E57CFFFA023EFF00FE46FDD3FE0B" ID-GBIF-Occurrence="3045014303" collectingDate="2019-05-01" collectorName="Jorge Eguis" country="Colombia" elevation="150" latitude="8.578611" location="Serrania de San Lucas." longLatPrecision="21" longitude="-74.12028" pageId="4" pageNumber="349" specimenCount="1" typeStatus="holotype">
<emphasis id="96F93333FFFA023EFF00FE46FE93FE50" bold="true" box="[151,267,403,429]" pageId="4" pageNumber="349">
<typeStatus id="7B365183FFFA023EFF00FE46FE9EFE50" box="[151,262,403,429]" pageId="4" pageNumber="349" type="holotype">Holotype</typeStatus>
.
</emphasis>
UPTC-Re 1010, adult male, collected by
<collectorName id="09788AF7FFFA023EFD49FE41FCFBFE50" box="[734,867,403,430]" pageId="4" pageNumber="349">Jorge Eguis</collectorName>
,
<date id="D033C9E1FFFA023EFCF9FE46FB9AFE50" box="[878,1026,403,430]" pageId="4" pageNumber="349" value="2019-05-01">
<collectingDate id="C0773009FFFA023EFCF9FE46FB9AFE50" box="[878,1026,403,430]" pageId="4" pageNumber="349" value="2019-05-01">01 May 2019</collectingDate>
</date>
at Hacienda IRHOPALMA, municipality of Norisí, department of Bolívar,
<collectingCountry id="DC9AAFB1FFFA023EFDC7FE62FD26FE2C" box="[592,702,439,465]" name="Colombia" pageId="4" pageNumber="349">Colombia</collectingCountry>
(
<geoCoordinate id="C1B989E6FFFA023EFD5AFE62FCCCFE2F" box="[717,852,439,466]" degrees="8" direction="north" minutes="34" orientation="latitude" pageId="4" pageNumber="349" precision="15" seconds="43" value="8.578611">8°34’43’’ N</geoCoordinate>
,
<geoCoordinate id="C1B989E6FFFA023EFCC9FE62FC60FE2F" box="[862,1016,439,466]" degrees="74" direction="west" minutes="07" orientation="longitude" pageId="4" pageNumber="349" precision="15" seconds="13" value="-74.12028">74°07’13’’ W</geoCoordinate>
;
<emphasis id="96F93333FFFA023EFB95FE6DFB84FE2C" box="[1026,1052,440,465]" italics="true" pageId="4" pageNumber="349">ca</emphasis>
.
<elevation id="2FA00812FFFA023EFBBDFE62FB33FE2F" box="[1066,1195,439,466]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="349" unit="m" value="150.0">
<quantity id="637542C4FFFA023EFBBDFE62FBE8FE2F" box="[1066,1136,439,466]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="349" unit="m" value="150.0">150 m</quantity>
a.s.l.
</elevation>
),
<quantity id="637542C4FFFA023EFB56FE62FB61FE2C" box="[1217,1273,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="349" unit="km" value="1.0">1 km</quantity>
apart from the main hill of the
<location id="A152B9FAFFFA023EFEDDFE0EFDD3FE0B" LSID="urn:lsid:plazi:treatment:2C245E37FFFA023DFF00FF0AFD40FF28:A152B9FAFFFA023EFEDDFE0EFDD3FE0B" box="[330,587,475,502]" country="Colombia" latitude="8.578611" longLatPrecision="21" longitude="-74.12028" name="Serrania de San Lucas." pageId="4" pageNumber="349">Serranía de San Lucas.</location>
</materialsCitation>
</paragraph>
<paragraph id="A432EF21FFFA023EFF50FE2AFA0EFDE4" blockId="4.[151,1437,403,2014]" box="[199,1430,511,538]" pageId="4" pageNumber="349">
<materialsCitation id="14E5E57CFFFA023EFF50FE2AFA0BFDE4" ID-GBIF-Occurrence="3045014302" box="[199,1427,511,538]" collectingDate="2019-05-01" collectorName="Jorge Eguis" country="Colombia" elevation="150" latitude="8.578611" location="Serrania de San Lucas." longLatPrecision="21" longitude="-74.12028" pageId="4" pageNumber="349" specimenCount="1" typeStatus="paratype">
<emphasis id="96F93333FFFA023EFF50FE2AFE15FDE4" bold="true" box="[199,397,511,537]" pageId="4" pageNumber="349">
<typeStatus id="7B365183FFFA023EFF50FE2AFEA7FDE4" box="[199,319,511,537]" pageId="4" pageNumber="349" type="paratype">Paratypes</typeStatus>
(n=3).
</emphasis>
UPTC-Re 1011–13 with same data as the holotype, collected by
<collectorName id="09788AF7FFFA023EFBFEFDD5FB76FDE4" box="[1129,1262,511,538]" pageId="4" pageNumber="349">Jorge Eguis</collectorName>
,
<date id="D033C9E1FFFA023EFB6DFE2AFA0BFDE4" box="[1274,1427,511,538]" pageId="4" pageNumber="349" value="2019-05-01">
<collectingDate id="C0773009FFFA023EFB6DFE2AFA0BFDE4" box="[1274,1427,511,538]" pageId="4" pageNumber="349" value="2019-05-01">01 May 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EC97BCAAFFFA023EFF50FDF6FD03F943" pageId="4" pageNumber="349" type="etymology">
<paragraph id="A432EF21FFFA023EFF50FDF6FC70FD78" blockId="4.[151,1437,403,2014]" pageId="4" pageNumber="349">
<emphasis id="96F93333FFFA023EFF50FDF6FED6FDC0" bold="true" box="[199,334,547,573]" pageId="4" pageNumber="349">Etymology.</emphasis>
The specific name
<taxonomicName id="638D94A2FFFA023EFDB9FDF1FD0AFDC0" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[558,658,548,573]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFDB9FDF1FD0AFDC0" box="[558,658,548,573]" italics="true" pageId="4" pageNumber="349">castanae</emphasis>
</taxonomicName>
is a patronym for Olga Victoria Castaño, to whom we dedicate with great honor this species in recognition of her multiple and meaningful contributions to the field of herpetology in
<collectingCountry id="DC9AAFB1FFFA023EFF00FDBEFE9DFD78" box="[151,261,619,645]" name="Colombia" pageId="4" pageNumber="349">Colombia</collectingCountry>
and who for many years was the mentor of the first author JECC.
</paragraph>
<paragraph id="A432EF21FFFA023EFF50FD5AFC9AFC58" blockId="4.[151,1437,403,2014]" pageId="4" pageNumber="349">
<emphasis id="96F93333FFFA023EFF50FD5AFE6BFD54" bold="true" box="[199,499,655,681]" pageId="4" pageNumber="349">Definition and diagnosis.</emphasis>
<materialsCitation id="14E5E57CFFFA023EFE6CFD5AFD66FC58" ID-GBIF-Occurrence="3045014304" collectionCode="SVL" collectorName="Blanco &amp; Schargel" pageId="4" pageNumber="349" specimenCount="1">
The new species is distinguished from other species of the genus
<taxonomicName id="638D94A2FFFA023EFB7EFD5AFAFAFD54" authorityName="Fitzinger" authorityYear="1843" box="[1257,1378,655,681]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="genus">
<emphasis id="96F93333FFFA023EFB7EFD5AFAFAFD54" box="[1257,1378,655,681]" italics="true" pageId="4" pageNumber="349">Gonatodes</emphasis>
</taxonomicName>
by a combination of the following characters: (1) body of moderate size (males from
<quantity id="637542C4FFFA023EFBA6FD66FA94FD33" box="[1073,1292,691,718]" metricMagnitude="-2" metricUnit="m" metricValue="4.305" metricValueMax="4.308" metricValueMin="4.302" pageId="4" pageNumber="349" unit="mm" value="43.05" valueMax="43.08" valueMin="43.02">43.02 to 43.08 mm</quantity>
and females from
<quantity id="637542C4FFFA023EFF43FD02FE13FD0F" box="[212,395,727,754]" metricMagnitude="-2" metricUnit="m" metricValue="4.175" metricValueMax="4.21" metricValueMin="4.14" pageId="4" pageNumber="349" unit="mm" value="41.75" valueMax="42.1" valueMin="41.4">41.4 to 42.1 mm</quantity>
<collectionCode id="C29C77E4FFFA023EFE06FD02FE52FD0C" box="[401,458,727,753]" pageId="4" pageNumber="349">SVL</collectionCode>
); (2) supraciliary spine absent, but with 3–7 conical supraciliary scales; (3) absence of a cluster of distinctly enlarged conical scales on the sides; (4) 102 to 109 scales around midbody; (5) 57 to 60 ventral scales counted in a longitudinal line back gular region to cloaca; (6) subcaudal scale pattern
<typeStatus id="7B365183FFFA023EFB37FCF5FB40FCC7" box="[1184,1240,800,826]" pageId="4" pageNumber="349">Type</typeStatus>
B (
<emphasis id="96F93333FFFA023EFA96FCF5FAA7FCC4" box="[1281,1343,800,825]" italics="true" pageId="4" pageNumber="349">sensu</emphasis>
Rivero-
<collectorName id="09788AF7FFFA023EFF00FC96FF7EFCA0" box="[151,230,835,861]" pageId="4" pageNumber="349">Blanco</collectorName>
&amp;
<collectorName id="09788AF7FFFA023EFE9EFC96FEF2FCA0" box="[265,362,835,861]" pageId="4" pageNumber="349">Schargel</collectorName>
2012,
<figureCitation id="3CB6F3A4FFFA023EFE22FC96FD96FCA0" box="[437,526,835,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1982,2006]" captionTargetBox="[198,1388,713,1948]" captionTargetId="figure-299@5.[190,1398,705,1958]" captionTargetPageId="5" captionText="FIGURE 2. Scale pattern of Gonatodes castanae sp. nov. (male holotype, UPTC-Re 1010). Subcaudal scale pattern Type B (A); escutcheon (B); scale pattern in the gular area (C). Illustrations by Ariadna Valenzuela." figureDoi="http://doi.org/10.5281/zenodo.4576571" httpUri="https://zenodo.org/record/4576571/files/figure.png" pageId="4" pageNumber="349">Fig. 2A</figureCitation>
); (7) males with escutcheon on belly and undersurface of thighs (
<figureCitation id="3CB6F3A4FFFA023EFB77FC96FAAFFCA0" box="[1248,1335,835,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1982,2006]" captionTargetBox="[198,1388,713,1948]" captionTargetId="figure-299@5.[190,1398,705,1958]" captionTargetPageId="5" captionText="FIGURE 2. Scale pattern of Gonatodes castanae sp. nov. (male holotype, UPTC-Re 1010). Subcaudal scale pattern Type B (A); escutcheon (B); scale pattern in the gular area (C). Illustrations by Ariadna Valenzuela." figureDoi="http://doi.org/10.5281/zenodo.4576571" httpUri="https://zenodo.org/record/4576571/files/figure.png" pageId="4" pageNumber="349">Fig. 2B</figureCitation>
); (8) two posmental scales (
<figureCitation id="3CB6F3A4FFFA023EFEF4FCB2FE23FC7C" box="[355,443,871,897]" captionStart="FIGURE 2" captionStartId="5.[151,250,1982,2006]" captionTargetBox="[198,1388,713,1948]" captionTargetId="figure-299@5.[190,1398,705,1958]" captionTargetPageId="5" captionText="FIGURE 2. Scale pattern of Gonatodes castanae sp. nov. (male holotype, UPTC-Re 1010). Subcaudal scale pattern Type B (A); escutcheon (B); scale pattern in the gular area (C). Illustrations by Ariadna Valenzuela." figureDoi="http://doi.org/10.5281/zenodo.4576571" httpUri="https://zenodo.org/record/4576571/files/figure.png" pageId="4" pageNumber="349">Fig. 2C</figureCitation>
); (9) two lateral rows of scales on distal parts of fingers and toes; (10) males and females with orange iris; (11) presence of sexual dichromatism
</materialsCitation>
.
</paragraph>
<paragraph id="A432EF21FFFA023EFF50FC7AFE87FACC" blockId="4.[151,1437,403,2014]" pageId="4" pageNumber="349">
<taxonomicName id="638D94A2FFFA023EFF50FC7AFE35FC34" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[199,429,943,969]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFF50FC7AFE35FC34" box="[199,429,943,969]" italics="true" pageId="4" pageNumber="349">Gonatodes castanae</emphasis>
</taxonomicName>
is found in the trans-Andean region of
<collectingCountry id="DC9AAFB1FFFA023EFCE4FC7AFC7CFC34" box="[883,996,943,969]" name="Colombia" pageId="4" pageNumber="349">Colombia</collectingCountry>
, in geographical proximity with
<emphasis id="96F93333FFFA023EFACEFC65FE16FC10" italics="true" pageId="4" pageNumber="349">
<taxonomicName id="638D94A2FFFA023EFACEFC65FE9DFC10" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="albogularis">G. albogularis</taxonomicName>
,
<taxonomicName id="638D94A2FFFA023EFE84FC01FE16FC10" authorityName="Meneses-Pelayo &amp; Ramirez" authorityYear="2020" box="[275,398,979,1006]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="chucuri">G. chucuri</taxonomicName>
</emphasis>
and
<emphasis id="96F93333FFFA023EFE5EFC01FDD0FC10" box="[457,584,980,1006]" italics="true" pageId="4" pageNumber="349">
<taxonomicName id="638D94A2FFFA023EFE5EFC01FDDCFC10" baseAuthorityName="Lichtenstein &amp; Martens" baseAuthorityYear="1856" box="[457,580,980,1006]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="vittatus">G. vittatus</taxonomicName>
.
</emphasis>
However,
<taxonomicName id="638D94A2FFFA023EFD5FFC01FCC9FC10" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[712,849,980,1006]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFD5FFC01FCC9FC10" box="[712,849,980,1006]" italics="true" pageId="4" pageNumber="349">G. castanae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="638D94A2FFFA023EFC7BFC01FBFFFC10" authorityName="Meneses-Pelayo &amp; Ramirez" authorityYear="2020" box="[1004,1127,979,1006]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="chucuri">
<emphasis id="96F93333FFFA023EFC7BFC01FBFFFC10" box="[1004,1127,979,1006]" italics="true" pageId="4" pageNumber="349">G. chucuri</emphasis>
</taxonomicName>
in the presence of marked sexual dichromatism (
<taxonomicName id="638D94A2FFFA023EFE05FC2DFD93FBEC" authorityName="Meneses-Pelayo &amp; Ramirez" authorityYear="2020" box="[402,523,1015,1042]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="chucuri">
<emphasis id="96F93333FFFA023EFE05FC2DFD93FBEC" box="[402,523,1015,1042]" italics="true" pageId="4" pageNumber="349">G. chucuri</emphasis>
</taxonomicName>
does not have sexual dichromatism) and in the subcaudal scale pattern, which is
<typeStatus id="7B365183FFFA023EFF00FBC9FF5FFBCB" box="[151,199,1052,1078]" pageId="4" pageNumber="349">type</typeStatus>
B, 1’1’1’’ without basal or distal changes in
<taxonomicName id="638D94A2FFFA023EFD5BFBC9FCCCFBC8" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[716,852,1052,1078]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFD5BFBC9FCCCFBC8" box="[716,852,1052,1078]" italics="true" pageId="4" pageNumber="349">G. castanae</emphasis>
</taxonomicName>
and
<typeStatus id="7B365183FFFA023EFC1AFBC9FC25FBCB" box="[909,957,1052,1078]" pageId="4" pageNumber="349">type</typeStatus>
C, 1’1’’or 1’2’’ basally, which changes to 1’1’’ distally in
<taxonomicName id="638D94A2FFFA023EFED8FB95FCF0FBA4" authority="(Meneses-Pelayo &amp; Ramirez 2020)" baseAuthorityName="Meneses-Pelayo &amp; Ramirez" baseAuthorityYear="2020" box="[335,872,1087,1114]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="chucuri">
<emphasis id="96F93333FFFA023EFED8FB95FE52FBA4" box="[335,458,1087,1114]" italics="true" pageId="4" pageNumber="349">G. chucuri</emphasis>
(
<bibRefCitation id="C01C92D0FFFA023EFE4DFBEAFCF9FBA4" author="Meneses-Pelayo, E. &amp; Ramirez, J. P." box="[474,865,1087,1113]" pageId="4" pageNumber="349" pagination="207 - 227" refId="ref7433" refString="Meneses-Pelayo, E. &amp; Ramirez, J. P. (2020) A new species of the genus Gonatodes (Squamata: Sauria: Sphaerodactylidae) from the western flank of the Cordillera Oriental in Colombia, with description of its hemipenial morphology. Zootaxa, 4729 (2), 207 - 227. https: // doi. org / 10.11646 / zootaxa. 4729.2.4" type="journal article" year="2020">Meneses-Pelayo &amp; Ramírez 2020</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="638D94A2FFFA023EFCE2FBEAFBC4FBA4" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[885,1116,1087,1113]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFCE2FBEAFBC4FBA4" box="[885,1116,1087,1113]" italics="true" pageId="4" pageNumber="349">Gonatodes castanae</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="638D94A2FFFA023EFB60FB95FA04FBA4" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[1271,1436,1087,1114]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="albogularis">
<emphasis id="96F93333FFFA023EFB60FB95FA04FBA4" box="[1271,1436,1087,1114]" italics="true" pageId="4" pageNumber="349">G. albogularis</emphasis>
</taxonomicName>
and
<taxonomicName id="638D94A2FFFA023EFF51FBB1FEA2FB80" baseAuthorityName="Lichtenstein &amp; Martens" baseAuthorityYear="1856" box="[198,314,1124,1150]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="vittatus">
<emphasis id="96F93333FFFA023EFF51FBB1FEA2FB80" box="[198,314,1124,1150]" italics="true" pageId="4" pageNumber="349">G. vittatus</emphasis>
</taxonomicName>
in the possession of subcaudal scale
<typeStatus id="7B365183FFFA023EFD4EFBB1FC91FB83" box="[729,777,1124,1150]" pageId="4" pageNumber="349">type</typeStatus>
B pattern, but differs from these in aspects of color pattern and size.
<taxonomicName id="638D94A2FFFA023EFE97FB52FE7CFB5C" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[256,484,1159,1185]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFE97FB52FE7CFB5C" box="[256,484,1159,1185]" italics="true" pageId="4" pageNumber="349">Gonatodes castanae</emphasis>
</taxonomicName>
has a dorsal color pattern in males, with conspicuous white and other gray ocelli and a moderate SVL (±
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), unlike
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<emphasis id="96F93333FFFA023EFDB8FB79FD4CFB38" box="[559,724,1195,1222]" italics="true" pageId="4" pageNumber="349">G. albogularis</emphasis>
</taxonomicName>
where males present a dorsal color that is most often uniform, in various shades (gray, black, brown) and is small in size (SVL &lt;
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;
<bibRefCitation id="C01C92D0FFFA023EFC4FFB1AFADBFB14" author="Rivero-Blanco, C. &amp; Schargel, W. E." box="[984,1347,1231,1257]" pageId="4" pageNumber="349" pagination="66 - 78" refId="ref7831" refString="Rivero-Blanco, C. &amp; Schargel, W. E. (2012) A strikingly polychromatic new species of Gonatodes (Squamata: Sphaerodactylidae) from northern Venezuela. Zootaxa, 3518 (1), 66 - 78. https: // doi. org / 10.11646 / zootaxa. 3518.1.4" type="journal article" year="2012">Rivero-Blanco &amp; Schargel 2012</bibRefCitation>
) and
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<emphasis id="96F93333FFFA023EFA16FB05FF71FAF0" italics="true" pageId="4" pageNumber="349">G. vittatus</emphasis>
</taxonomicName>
whose males have a distinctive color pattern with a pale mid-dorsal stripe and an SVL &lt;
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(
<bibRefCitation id="C01C92D0FFFA023EFAAEFB26FE8BFACC" author="Schargel, W. E. &amp; Rivas, G. A. &amp; Garcia-Perez, J. E. &amp; Rivero-Blanco, C. &amp; Chippindale, P. T. &amp; Fujita, M. K." pageId="4" pageNumber="349" pagination="549 - 562" refId="ref8254" refString="Schargel, W. E., Rivas, G. A., Garcia-Perez, J. E., Rivero-Blanco, C., Chippindale, P. T. &amp; Fujita, M. K. (2017) A new species of Gonatodes (Squamata: Sphaerodactylidae) from the western versant of the Cordillera de Merida, Venezuela. Zootaxa, 4291 (3), 549 - 562. https: // doi. org / 10.11646 / zootaxa. 4291.3.7" type="journal article" year="2017">
Schargel
<emphasis id="96F93333FFFA023EFF00FACDFF50FACC" box="[151,200,1303,1329]" italics="true" pageId="4" pageNumber="349">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
<paragraph id="A432EF21FFFA023EFF50FAEEFD03F943" blockId="4.[151,1437,403,2014]" pageId="4" pageNumber="349">
<taxonomicName id="638D94A2FFFA023EFF50FAEEFE37FAA8" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[199,431,1339,1365]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFF50FAEEFE37FAA8" box="[199,431,1339,1365]" italics="true" pageId="4" pageNumber="349">Gonatodes castanae</emphasis>
</taxonomicName>
shows similarities to the color pattern of
<taxonomicName id="638D94A2FFFA023EFC09FAEEFF4AFA84" authority="Rivero-Blanco &amp; Schargel, 2012" authorityName="Rivero-Blanco &amp; Schargel" authorityYear="2012" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="rozei">
<emphasis id="96F93333FFFA023EFC09FAEEFBC1FAA8" box="[926,1113,1339,1365]" italics="true" pageId="4" pageNumber="349">Gonatodes rozei</emphasis>
<bibRefCitation id="C01C92D0FFFA023EFBF3FAEEFF4AFA84" author="Rivero-Blanco, C. &amp; Schargel, W. E." pageId="4" pageNumber="349" pagination="66 - 78" refId="ref7831" refString="Rivero-Blanco, C. &amp; Schargel, W. E. (2012) A strikingly polychromatic new species of Gonatodes (Squamata: Sphaerodactylidae) from northern Venezuela. Zootaxa, 3518 (1), 66 - 78. https: // doi. org / 10.11646 / zootaxa. 3518.1.4" type="journal article" year="2012">Rivero-Blanco &amp; Schargel, 2012</bibRefCitation>
</taxonomicName>
, the latter found in both chains (Serranía del Litoral and Serranía del Interior) of the Cordillera de la Costa Central in north central
<collectingCountry id="DC9AAFB1FFFA023EFE39FA56FDB9FA60" box="[430,545,1411,1437]" name="Venezuela" pageId="4" pageNumber="349">Venezuela</collectingCountry>
(
<bibRefCitation id="C01C92D0FFFA023EFDA5FA56FC32FA60" author="Rivero-Blanco, C. &amp; Schargel, W. E." box="[562,938,1411,1437]" pageId="4" pageNumber="349" pagination="66 - 78" refId="ref7831" refString="Rivero-Blanco, C. &amp; Schargel, W. E. (2012) A strikingly polychromatic new species of Gonatodes (Squamata: Sphaerodactylidae) from northern Venezuela. Zootaxa, 3518 (1), 66 - 78. https: // doi. org / 10.11646 / zootaxa. 3518.1.4" type="journal article" year="2012">Rivero-Blanco &amp; Schargel 2012</bibRefCitation>
). The dorsal color pattern with ocelli in
<taxonomicName id="638D94A2FFFA023EFA16FA51FF63FA3C" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="castanae">
<emphasis id="96F93333FFFA023EFA16FA51FF63FA3C" italics="true" pageId="4" pageNumber="349">G. castanae</emphasis>
</taxonomicName>
is homogeneous and covers the dorsal region, the flanks, the anterior and posterior limbs and the tip of the tail, with a high concentration of ocelli in the posterior part of the body and in the posterior limbs. Unlike
<taxonomicName id="638D94A2FFFA023EFAABFA19FA0DFA18" authorityName="Rivero-Blanco &amp; Schargel" authorityYear="2012" box="[1340,1429,1484,1510]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="349" phylum="Chordata" rank="species" species="rozei">
<emphasis id="96F93333FFFA023EFAABFA19FA0DFA18" box="[1340,1429,1484,1510]" italics="true" pageId="4" pageNumber="349">G. rozei</emphasis>
</taxonomicName>
, where the ocelli are present on the back, flanks, anterior legs and thighs. Each ocellus has poorly defined black borders and the ocelli are more pronounced anteriorally towards the front of the body and become small and scattered posteriorly towards the rear and do not extend beyond the base of the tail (
<bibRefCitation id="C01C92D0FFFA023EFC4EF9E2FAD2F9AC" author="Rivero-Blanco, C. &amp; Schargel, W. E." box="[985,1354,1591,1617]" pageId="4" pageNumber="349" pagination="66 - 78" refId="ref7831" refString="Rivero-Blanco, C. &amp; Schargel, W. E. (2012) A strikingly polychromatic new species of Gonatodes (Squamata: Sphaerodactylidae) from northern Venezuela. Zootaxa, 3518 (1), 66 - 78. https: // doi. org / 10.11646 / zootaxa. 3518.1.4" type="journal article" year="2012">Rivero-Blanco &amp; Schargel 2012</bibRefCitation>
). Other differences between these two species are (
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<emphasis id="96F93333FFFA023EFDEDF989FD4BF988" box="[634,723,1628,1654]" italics="true" pageId="4" pageNumber="349">G. rozei</emphasis>
</taxonomicName>
characters in parentheses): SVL
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(SVL
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), scales around midbody 102–109 (78–100 scales around midbody), iris of orange in males and orange or yellow in females (males with blue iris, females blue or yellow).
</paragraph>
</subSubSection>
<subSubSection id="EC97BCAAFFFA023CFF50F912FBA7F8FB" lastPageId="6" lastPageNumber="351" pageId="4" pageNumber="349" type="description">
<paragraph id="A432EF21FFFA023FFF50F912FD83FE2C" blockId="4.[151,1437,403,2014]" lastBlockId="5.[151,1437,151,681]" lastPageId="5" lastPageNumber="350" pageId="4" pageNumber="349">
<emphasis id="96F93333FFFA023EFF50F912FD81F91C" bold="true" box="[199,537,1735,1761]" pageId="4" pageNumber="349">
Description of the
<typeStatus id="7B365183FFFA023EFE3BF912FD8DF91C" box="[428,533,1735,1761]" pageId="4" pageNumber="349" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male (
<figureCitation id="3CB6F3A4FFFA023EFD22F912FC88F91C" box="[693,784,1735,1761]" captionStart="FIGURE 3" captionStartId="7.[151,250,1171,1195]" captionTargetBox="[151,1436,237,1148]" captionTargetId="figure-48@7.[151,1436,237,1148]" captionTargetPageId="7" captionText="FIGURE 3. Color pattern of Gonatodes castanae sp. nov. UPTC-Re 1010 male holotype (A); UPTC-Re 1011 male paratype (B); UPTC-Re 1012 female paratype (C); UPTC-Re 1013 male paratype (D). Photos by Jorge Andrés Eguis-Avendaño." figureDoi="http://doi.org/10.5281/zenodo.4576575" httpUri="https://zenodo.org/record/4576575/files/figure.png" pageId="4" pageNumber="349">Fig. 3A</figureCitation>
) with SVL of
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. Tail length
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, complete, original. Head approximately 1.3 times longer than wide (head length:
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; head width:
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). Short snout, with eye-nostril distance of
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, 0.3 times the length of the head. Rostral moderately large, visible from above, with a longitudinal slit almost dividing it that extends to just below the middle of the rostral. Two postrostrals (supranasals), large and in contact with each other. Nasals bordered by the rostral and three postnasals (left) and three (right). Postnasal slightly larger than scales in loreal region. Scales on top of snout and loreal region round at base, sub-conical and slightly juxtaposed. Loreal scales approximately
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a line between the postnasal and the anterior margin of the eye socket. The scales slightly decrease in size from the postrostral to the back of the head. Scales on the supraorbital almost the same size and shape as those on the top of the head. Supraciliares with seven well-developed spines located above the center and sides of the eye. The supraciliary scales anterior to the spines are slightly conical and larger than the supraorbital ones; posterior to the spines, the supraciliary scales are smaller, slightly conical in the form of short, poorly developed spines. Supralabial 6/6 (left/right), the first is large (as longer as tall), the second, third and fourth smaller than the first and almost equal in size between them, the fifth and sixth are considerably smaller in relation to the others. The fourth scale is at the level of the central part of the eye. After the sixth supralabial, 9–11 (left/right) small granular scales follow the corner of the mouth. Scales in the temporal region similar in shape and size to those on the upper back of the head. Otic opening (
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) about half size of the eye (
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), obliquely oval.
</paragraph>
<paragraph id="A432EF21FFFB023FFF50FE0EFB26FD54" blockId="5.[151,1437,151,681]" pageId="5" pageNumber="350">
Mental large, slightly hexagonal, rounded on the front and triangular (convex) on the back. Two postmentals, clearly larger than the adjacent posterior scales (
<figureCitation id="3CB6F3A4FFFB023FFD3AFE2AFC9CFDE4" box="[685,772,511,537]" captionStart="FIGURE 2" captionStartId="5.[151,250,1982,2006]" captionTargetBox="[198,1388,713,1948]" captionTargetId="figure-299@5.[190,1398,705,1958]" captionTargetPageId="5" captionText="FIGURE 2. Scale pattern of Gonatodes castanae sp. nov. (male holotype, UPTC-Re 1010). Subcaudal scale pattern Type B (A); escutcheon (B); scale pattern in the gular area (C). Illustrations by Ariadna Valenzuela." figureDoi="http://doi.org/10.5281/zenodo.4576571" httpUri="https://zenodo.org/record/4576571/files/figure.png" pageId="5" pageNumber="350">Fig. 2C</figureCitation>
). Mental region immediately behind the posmentals and between the infralabials with a series of moderately large scales of variable and juxtaposed shapes. The rest of the chin scales are small, granular and conical. Infralabials 3/3 at the level of the center of the eye, decreasing in size towards the back. The first infralabial is very large and projecting on the chin, the second is moderately large, slightly less than half the size of the first infralabial, the third located below the center of the eye is smaller.
</paragraph>
<caption id="F0F2BFA9FFFB023FFF00F86BFB9FF807" ID-DOI="http://doi.org/10.5281/zenodo.4576571" ID-Zenodo-Dep="4576571" httpUri="https://zenodo.org/record/4576571/files/figure.png" pageId="5" pageNumber="350" startId="5.[151,250,1982,2006]" targetBox="[198,1388,713,1948]" targetPageId="5">
<paragraph id="A432EF21FFFB023FFF00F86BFB9FF807" blockId="5.[151,1436,1982,2042]" pageId="5" pageNumber="350">
<emphasis id="96F93333FFFB023FFF00F86BFE8AF82B" bold="true" box="[151,274,1982,2006]" pageId="5" pageNumber="350">FIGURE 2.</emphasis>
Scale pattern of
<taxonomicName id="638D94A2FFFB023FFE2EF86BFD1DF82B" authority="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya, 2020" authorityName="Carvajal-Cogollo &amp; Eguis-Avendaño &amp; Meza-Joya" authorityYear="2020" box="[441,645,1982,2006]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="350" phylum="Chordata" rank="species" species="castanae" status="sp. nov.">
<emphasis id="96F93333FFFB023FFE2EF86BFD1DF82B" box="[441,645,1982,2006]" italics="true" pageId="5" pageNumber="350">Gonatodes castanae</emphasis>
</taxonomicName>
<emphasis id="96F93333FFFB023FFD1CF86BFD42F82B" bold="true" box="[651,730,1982,2006]" pageId="5" pageNumber="350">
<taxonomicNameLabel id="8DCA8E48FFFB023FFD1CF86BFD42F82B" box="[651,730,1982,2006]" pageId="5" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male holotype, UPTC-Re 1010). Subcaudal scale pattern Type B (A); escutcheon (B); scale pattern in the gular area (C). Illustrations by Ariadna Valenzuela.
</paragraph>
</caption>
<paragraph id="A432EF21FFF8023CFF50FF42FC05FE74" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="351">
Granular scales on the nape and on the sides of the neck, continuous with those of the head and body. Scales on the throat smooth, imbricate, with a round posterior margin, with a short transitional area of granular scales on the chin and gular area. Granular semi-conical vertebral ridges, in size similar to the scales on the snout, but larger than those on the top of the head. Neck
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wide (at the narrowest part), narrower than the head and body. Body wider than tall, sub-cylindrical, axillar-groin distance
<quantity id="637542C4FFF8023CFD65FEF2FCC1FEBC" box="[754,857,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="6" pageNumber="351" unit="mm" value="16.2">16.2 mm</quantity>
. Well-developed limbs with moderately long digits, length of the fourth toe
<quantity id="637542C4FFF8023CFE0CFE9FFE6AFE98" box="[411,498,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="6" pageNumber="351" unit="mm" value="5.6">5.6 mm</quantity>
, 0.96 times the length of the “shaft” (
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, from the knee to the ankle-point where the foot is bent). Round tail in cross section, tapering towards the tip.
</paragraph>
<paragraph id="A432EF21FFF8023CFF50FE46FBC1FD9C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="351">
Ventral scales much larger than dorsal, smooth, imbricate, with round posterior margins, arranged mostly in oblique rows. There are 44 scales between the armpit and the groin area. Scales around mid-body about 109 of which 24 are ventral. Scales in the precloacal region similar to the ventral ones, except proximally to the cloacal opening, which has tiny scales arranged in several rows. Escutcheon present in the lower abdomen and on the lower surface of the thighs (
<figureCitation id="3CB6F3A4FFF8023CFE04FDF6FE73FDC0" box="[403,491,547,573]" captionStart="FIGURE 2" captionStartId="5.[151,250,1982,2006]" captionTargetBox="[198,1388,713,1948]" captionTargetId="figure-299@5.[190,1398,705,1958]" captionTargetPageId="5" captionText="FIGURE 2. Scale pattern of Gonatodes castanae sp. nov. (male holotype, UPTC-Re 1010). Subcaudal scale pattern Type B (A); escutcheon (B); scale pattern in the gular area (C). Illustrations by Ariadna Valenzuela." figureDoi="http://doi.org/10.5281/zenodo.4576571" httpUri="https://zenodo.org/record/4576571/files/figure.png" pageId="6" pageNumber="351">Fig. 2B</figureCitation>
) with 16 scales rows wide and 18 scales long. On the ventral surfaces of the hind limbs, the escutcheon is 4 rows of scales wide and extends medially across the thighs.
</paragraph>
<paragraph id="A432EF21FFF8023CFF50FDBEFA18FCE8" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="351">
Small, conical and juxtaposed scales on the forelimbs, on the back of the forearm and become larger, flatter, imbricate and with a rounded posterior edge, towards the anterior margin of the forearm, as well as on the arm. Small, semi-conical, and juxtaposed scales on the posterior half of the hind limbs, both dorsal and ventral. In the ventral anterior half of the limb the scales are flat and overlapping. On the ventral surface of the hind limbs there is an abrupt change between two
<typeStatus id="7B365183FFF8023CFE64FD2EFDB6FCE8" box="[499,558,763,789]" pageId="6" pageNumber="351">types</typeStatus>
of scales, the ventrals and the 4 rows that form the escutcheon on the thigh.
</paragraph>
<paragraph id="A432EF21FFF8023CFF50FCCAFAB1FC58" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="351">
Lamellae under the first (I) to the fifth (V) finger (infraproximal in parentheses, right/left;
<figureCitation id="3CB6F3A4FFF8023CFB45FCCAFAB5FCC4" box="[1234,1325,799,825]" captionStart="FIGURE 4" captionStartId="7.[151,250,1929,1953]" captionTargetBox="[155,1433,1260,1902]" captionTargetId="figure-85@7.[151,1436,1256,1906]" captionTargetPageId="7" captionText="FIGURE 4. Ventral view of the hand and feet, illustrating the lamellae of Gonatodes castanae sp. nov. Left hand fingers (A), left foot fingers (B). Illustrations by Ariadna Valenzuela." figureDoi="http://doi.org/10.5281/zenodo.4576577" httpUri="https://zenodo.org/record/4576577/files/figure.png" pageId="6" pageNumber="351">Fig. 4A</figureCitation>
): I: 12/12 (4/4), II: 15/15 (7/6), III: 17/17 (8/8), IV: 18/17 (7/7), and V: 15/15 (5/5), respectively. Subdigital lamellae from first to fifth toe (infraproximal in parentheses;
<figureCitation id="3CB6F3A4FFF8023CFDFCFCB2FD5AFC7C" box="[619,706,871,897]" captionStart="FIGURE 4" captionStartId="7.[151,250,1929,1953]" captionTargetBox="[155,1433,1260,1902]" captionTargetId="figure-85@7.[151,1436,1256,1906]" captionTargetPageId="7" captionText="FIGURE 4. Ventral view of the hand and feet, illustrating the lamellae of Gonatodes castanae sp. nov. Left hand fingers (A), left foot fingers (B). Illustrations by Ariadna Valenzuela." figureDoi="http://doi.org/10.5281/zenodo.4576577" httpUri="https://zenodo.org/record/4576577/files/figure.png" pageId="6" pageNumber="351">Fig. 4B</figureCitation>
): I:? (no data)/10 (?/4), II: 13/14 (5/5), III: 17/19 (7/6), IV: 20/20 (8/9) and V: 19/19 (7/8), respectively. Fore and hind limb fingers with two lateral rows of scales distally.
</paragraph>
<paragraph id="A432EF21FFF8023CFF50FC7BFD91FA60" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="351">
<emphasis id="96F93333FFF8023CFF50FC7BFEC4FC35" bold="true" box="[199,348,942,968]" pageId="6" pageNumber="351">Color in life.</emphasis>
In live males (
<figureCitation id="3CB6F3A4FFF8023CFD96FC7AFDD9FC34" box="[513,577,943,969]" captionStart="FIGURE 3" captionStartId="7.[151,250,1171,1195]" captionTargetBox="[151,1436,237,1148]" captionTargetId="figure-48@7.[151,1436,237,1148]" captionTargetPageId="7" captionText="FIGURE 3. Color pattern of Gonatodes castanae sp. nov. UPTC-Re 1010 male holotype (A); UPTC-Re 1011 male paratype (B); UPTC-Re 1012 female paratype (C); UPTC-Re 1013 male paratype (D). Photos by Jorge Andrés Eguis-Avendaño." figureDoi="http://doi.org/10.5281/zenodo.4576575" httpUri="https://zenodo.org/record/4576575/files/figure.png" pageId="6" pageNumber="351">Fig. 3</figureCitation>
A–B) the dorsal and lateral surface of the body and the anterior and posterior limbs, like the anterolateral and dorsal portion of the tail, have a black background color. The background color turns brown on the distal, dorsal and lateral parts of the tail and hind legs. The back, flanks, front legs, thighs and the tip of the tail are covered with white ocelli. The dorsal and lateral surface of the body also features less conspicuous gray ocelli, interspersed with white ocelli. White ocelli exhibit different shapes (circular, irregular) and sizes. The largest ocelli are about one third the size of the eye and are located on the mid-dorsal region. White ocelli are smaller and more numerous on the low-dorsal region and become more dispersed in the tail. The head is covered, both dorsally and ventrally, by a reddish orange hood, which extends to the nape and neck. The ventral surface of the body, except the area of the shield, macroscopically exhibits a slightly mottled gray coloration. Under magnification, each ventral scale has multiple black dots homogeneously scattered throughout the scale. Females show a dorsal coloration with a greenish-yellow background, with brown to black reticulations and scattered white spots on the head, vertebral region and flanks. The fore and hind limbs have a grayish background color with small greenish-yellow stripes (
<figureCitation id="3CB6F3A4FFF8023CFEFCFA8AFE34FA84" box="[363,428,1375,1401]" captionStart="FIGURE 3" captionStartId="7.[151,250,1171,1195]" captionTargetBox="[151,1436,237,1148]" captionTargetId="figure-48@7.[151,1436,237,1148]" captionTargetPageId="7" captionText="FIGURE 3. Color pattern of Gonatodes castanae sp. nov. UPTC-Re 1010 male holotype (A); UPTC-Re 1011 male paratype (B); UPTC-Re 1012 female paratype (C); UPTC-Re 1013 male paratype (D). Photos by Jorge Andrés Eguis-Avendaño." figureDoi="http://doi.org/10.5281/zenodo.4576575" httpUri="https://zenodo.org/record/4576575/files/figure.png" pageId="6" pageNumber="351">Fig. 3</figureCitation>
C–D). Venter with yellowish tones, more accentuated in the lower part of the belly, turning whitish towards the chest region.
</paragraph>
<paragraph id="A432EF21FFF8023CFF50FA73FD88FA18" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="351">
<emphasis id="96F93333FFF8023CFF50FA73FE53FA3D" bold="true" box="[199,459,1446,1472]" pageId="6" pageNumber="351">Color in preservative.</emphasis>
The preserved color has not changed significantly, except for the head in males, which has taken on a grayish coloration.
</paragraph>
<paragraph id="A432EF21FFF8023CFF50FA3BFBA7F8FB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="351">
<emphasis id="96F93333FFF8023CFF50FA3BFEA7F9F5" bold="true" box="[199,319,1518,1544]" pageId="6" pageNumber="351">Variation.</emphasis>
<materialsCitation id="14E5E57CFFF8023CFEDFFA3AFF74F9AC" ID-GBIF-Occurrence="3045014301" collectionCode="SVL" pageId="6" pageNumber="351" specimenCount="1" typeStatus="holotype">
Body size (
<collectionCode id="C29C77E4FFF8023CFE5DFA3AFD9AF9F4" box="[458,514,1519,1545]" pageId="6" pageNumber="351">SVL</collectionCode>
) was similar among all specimens, ranging between 41.4 and
<quantity id="637542C4FFF8023CFB5AFA3AFAACF9F4" box="[1229,1332,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="4.21" pageId="6" pageNumber="351" unit="mm" value="42.1">42.1 mm</quantity>
in males and between 41.0 and
<quantity id="637542C4FFF8023CFE03F9C6FE61F9D0" box="[404,505,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="4.18" pageId="6" pageNumber="351" unit="mm" value="41.8">41.8 mm</quantity>
in females. The largest male is the
<typeStatus id="7B365183FFF8023CFC1EF9C6FC73F9D0" box="[905,1003,1555,1581]" pageId="6" pageNumber="351" type="holotype">holotype</typeStatus>
(UPTC-Re 1010) measuring
<quantity id="637542C4FFF8023CFAAFF9C6FA05F9D0" box="[1336,1437,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="4.21" pageId="6" pageNumber="351" unit="mm" value="42.1">42.1 mm</quantity>
in
<collectionCode id="C29C77E4FFF8023CFF23F9E2FF74F9AC" box="[180,236,1591,1617]" pageId="6" pageNumber="351">SVL</collectionCode>
</materialsCitation>
. There are typically three or four (UPTC-Re 1012-13), infralabials to the level of the center of eye. Postrostrals 2–3. Loreal scales 9–11. Scales around the midbody are 102–109 of which about 20–24 are ventrals. There are 36–44 scales along the midventral line between the level of anterior margin of forelimbs and the small scales anterior to vent. Lamellae (infraproximals in parentheses) under the fingers are: I: 10–13 (4–5), II: 13–15 (7–6), III: 15–16 (5–6), IV: 16–15 (6–5), and V: 16–15 (6–6). Lamellae (infraproximals in parentheses) under the toes are: I: 9–10 (4–4), II: 13–14 (5–4), III: 17–16 (6–9), IV: 18–20 (8–6), and V: 18–17 (8–6).
</paragraph>
</subSubSection>
<subSubSection id="EC97BCAAFFF8023DFF50F8DBFD40FF28" lastPageId="7" lastPageNumber="352" pageId="6" pageNumber="351" type="distribution">
<paragraph id="A432EF21FFF8023DFF50F8DBFD40FF28" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,150,213]" lastPageId="7" lastPageNumber="352" pageId="6" pageNumber="351">
<emphasis id="96F93333FFF8023CFF50F8DBFDCEF8D5" bold="true" box="[199,598,1806,1833]" pageId="6" pageNumber="351">Distribution and natural history.</emphasis>
Known only from the
<typeStatus id="7B365183FFF8023CFCF3F8DAFC0CF8D4" box="[868,916,1807,1833]" pageId="6" pageNumber="351">type</typeStatus>
locality (municipality of Norosí,
<collectingRegion id="664921C3FFF8023CFA8CF8DAFF65F8B0" country="Colombia" name="Bolivar" pageId="6" pageNumber="351">Bolívar department</collectingRegion>
) in the lowlands (
<emphasis id="96F93333FFF8023CFE5FF8E6FE7AF8B1" box="[456,482,1843,1868]" italics="true" pageId="6" pageNumber="351">ca</emphasis>
.
<quantity id="637542C4FFF8023CFE66F8E7FDA1F8B0" box="[497,569,1842,1869]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="351" unit="m" value="150.0">150 m</quantity>
a.s.l.) of the northeastern versant of the Serranía de San Lucas, northwestern
<collectingCountry id="DC9AAFB1FFF8023CFF00F882FE9DF88C" box="[151,261,1879,1905]" name="Colombia" pageId="6" pageNumber="351">Colombia</collectingCountry>
(
<figureCitation id="3CB6F3A4FFF8023CFE8FF882FEC7F88C" box="[280,351,1878,1905]" captionStart="FIGURE 5" captionStartId="8.[151,250,945,969]" captionTargetBox="[154,1435,181,918]" captionTargetId="figure-23@8.[151,1436,181,921]" captionTargetPageId="8" captionText="FIGURE 5. Distribution of Gonatodes castanae sp. nov. (red dot) in the municipality of Norosí, south of the Bolívar department, foothills of the Serranía de San Lucas, Middle Magdalena Valley, Colombia." figureDoi="http://doi.org/10.5281/zenodo.4576581" httpUri="https://zenodo.org/record/4576581/files/figure.png" pageId="6" pageNumber="351">Fig. 5</figureCitation>
). Specimens collected were found in an isolated relictual riparian forest surrounded by an oil palm monoculture at Hacienda IRHOPALMA (
<figureCitation id="3CB6F3A4FFF8023CFD3FF8AEFC9AF86B" box="[680,770,1915,1942]" captionStart="FIGURE 6" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[151,1436,1029,1849]" captionTargetId="figure-59@8.[151,1436,1029,1849]" captionTargetPageId="8" captionText="FIGURE 6. Habitat of Gonatodes castanae sp. nov. at Hacienda IRHOPALMA, municipality of Norisí, department of Bolívar, Colombia. General view of the San Pedro stream (A) and microhabitat where the holotype specimen was collected (B). The arrowhead indicates the collection site. Photos by Willinton Barranco." figureDoi="http://doi.org/10.5281/zenodo.4576583" httpUri="https://zenodo.org/record/4576583/files/figure.png" pageId="6" pageNumber="351">Fig. 6A</figureCitation>
). The collected individuals were first observed basking and foraging on the bark of trees located near to the margin of the
<collectingCountry id="DC9AAFB1FFF8023CFCCFF84AFC52F844" box="[856,970,1951,1977]" name="Saint Pierre and Miquelon" pageId="6" pageNumber="351">San Pedro</collectingCountry>
stream (see
<figureCitation id="3CB6F3A4FFF8023CFBCDF84AFB2AF847" box="[1114,1202,1951,1978]" captionStart="FIGURE 6" captionStartId="8.[151,250,1873,1897]" captionTargetBox="[151,1436,1029,1849]" captionTargetId="figure-59@8.[151,1436,1029,1849]" captionTargetPageId="8" captionText="FIGURE 6. Habitat of Gonatodes castanae sp. nov. at Hacienda IRHOPALMA, municipality of Norisí, department of Bolívar, Colombia. General view of the San Pedro stream (A) and microhabitat where the holotype specimen was collected (B). The arrowhead indicates the collection site. Photos by Willinton Barranco." figureDoi="http://doi.org/10.5281/zenodo.4576583" httpUri="https://zenodo.org/record/4576583/files/figure.png" pageId="6" pageNumber="351">Fig. 6B</figureCitation>
). The sightings were made between 3:00 and 5:00 p.m., in a patch of secondary vegetation with a high degree of intervention, with an approximate area of five hectares. The males occupied the upper parts of the tree stem, at heights from
<quantity id="637542C4FFF9023DFAB2FF43FAF4FF4C" box="[1317,1388,150,177]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="7" pageNumber="352" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">1–5 m</quantity>
and the females the lower parts, between 0.6 and
<quantity id="637542C4FFF9023DFD06FF6EFD4CFF28" box="[657,724,187,213]" metricMagnitude="0" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="352" unit="m" value="1.3">1.3 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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