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<document ID-DOI="10.11646/zootaxa.5263.3.6" ID-ISSN="1175-5326" ID-ZooBank="2952C656-9F88-4EF1-B435-2EA07F86DF5C" approvalRequired="5" approvalRequired_for_textStreams="4" approvalRequired_for_treatments="1" checkinTime="1680760756093" checkinUser="plazi" docAuthor="Wesener, Thomas, Moritz, Leif &amp; Akkari, Nesrine" docDate="2023" docId="29675F45FFA8FFFDFF5A6A2EFA8FF988" docLanguage="en" docName="zootaxa.5263.3.6.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dawydoffia siphonocryptida Wesener &amp; Moritz &amp; Akkari 2023, new species" docType="treatment" docVersion="1" lastPageNumber="423" masterDocId="D55E273DFFA3FFF1FFCD6D65FFB8FFF9" masterDocTitle="Integrative redescription of the sucking millipede genus Dawydoffia Attems, 1953 with a description of a new species and a transfer to the family Hirudisomatidae (Diplopoda, Polyzoniida)" masterLastPageNumber="429" masterPageNumber="411" pageNumber="422" updateTime="1680760842833" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Integrative redescription of the sucking millipede genus Dawydoffia Attems, 1953 with a description of a new species and a transfer to the family Hirudisomatidae (Diplopoda, Polyzoniida)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:affiliation>Leibniz Institute for the Study of Biodiversity Change, Zoogisches Forschungsmuseum Alexander Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moritz, Leif</mods:namePart>
<mods:affiliation>Leibniz Institute for the Study of Biodiversity Change, Zoogisches Forschungsmuseum Alexander Koenig, Adenauerallee 127, 53113 Bonn, Germany. &amp; University of Bonn, Institute for Evolutionary Biology and Ecology, An der Immenburg 1, 53121 Bonn, Germany.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Akkari, Nesrine</mods:namePart>
<mods:affiliation>Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5263</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>411</mods:start>
<mods:end>429</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.3.6</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5263.3.6</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="ZooBank">2952C656-9F88-4EF1-B435-2EA07F86DF5C</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:29675F45FFA8FFFDFF5A6A2EFA8FF988" httpUri="http://treatment.plazi.org/id/29675F45FFA8FFFDFF5A6A2EFA8FF988" lastPageId="12" lastPageNumber="423" pageId="11" pageNumber="422">
<subSubSection box="[151,643,1867,1894]" pageId="11" pageNumber="422" type="nomenclature">
<paragraph blockId="11.[151,643,1867,1930]" box="[151,643,1867,1894]" pageId="11" pageNumber="422">
<heading bold="true" box="[151,643,1867,1894]" fontSize="11" level="1" pageId="11" pageNumber="422" reason="1">
<emphasis bold="true" box="[151,643,1867,1894]" pageId="11" pageNumber="422">
<taxonomicName ID-CoL="7NGBG" authority="Wesener &amp; Moritz &amp; Akkari, 2023" authorityName="Wesener &amp; Moritz &amp; Akkari" authorityYear="2023" box="[151,492,1867,1894]" class="Diplopoda" family="Siphonotidae" genus="Dawydoffia" kingdom="Animalia" order="Polyzoniida" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="siphonocryptida" status="new species">
<emphasis bold="true" box="[151,492,1867,1894]" italics="true" pageId="11" pageNumber="422">Dawydoffia siphonocryptida</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[499,643,1868,1894]" pageId="11" pageNumber="422" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,284,1904,1930]" pageId="11" pageNumber="422" type="description">
<paragraph blockId="11.[151,643,1867,1930]" box="[151,284,1904,1930]" pageId="11" pageNumber="422">
<figureCitation box="[151,284,1904,1930]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="12.[151,250,980,1004]" captionStartId-1="13.[151,250,1511,1535]" captionStartId-2="14.[151,250,1866,1890]" captionStartId-3="15.[151,250,1791,1815]" captionTargetBox-0="[168,1429,185,950]" captionTargetBox-1="[151,1435,381,1486]" captionTargetBox-2="[151,1436,453,1842]" captionTargetBox-3="[227,1361,181,1767]" captionTargetId-0="figure-18@12.[151,1436,181,956]" captionTargetId-1="figure-111@13.[151,1435,381,1487]" captionTargetId-2="figure-153@14.[151,1436,453,1842]" captionTargetId-3="figure-19@15.[227,1361,181,1767]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="FIGURE 6. Dawydoffia siphonocryptida n. sp., female paratype (SMF), volume rendering based on µCT-data.A. Habitus lateral view. B. Head and anterior body, ventral view. C. Telson and posterior body, ventral view. Abbreviations: At = Antennae; Co = collum; ep = epicranium; fh = forehead; oz = ozopore, Pl = pleurite; St = sternite; T = tergite; Te = telson; Vu = vulva." captionText-1="FIGURE 7. Dawydoffia siphonocryptida n. sp., male holotype (SMF), scanning electron micrographs (SEM). A. Anterior body, lateral view. B. Anterior body, ventral view. C. Frontal view of specimen. D. Detail of labrum and gnathochilarium, apical view. E. Anterior segments showing fused tergites-pleurites. F. Head, lateral view. G. Right antenna, ventral view. H. Apical tip of right antenna. Abbreviations: aG = anterior gonopod; CO = collum (tergite 1); d = antennal disc; gn = gnathochilarium; La = labrum; ms = macrosetae; O = ommatidium; oz = ozopore; pG = posterior gonopod; sb = sensilla basiconica. Arabian numbers refer to number of leg pair, tergite or antennomere." captionText-2="FIGURE 8. Dawydoffia siphonocryptida n. sp., male holotype (SMF), scanning electron micrographs. A. Posterior body end, dorsal view. B. posterior body end with telson, ventral view. C. First tarsal claw. D. Leg 5, tarsal claw. E. Hirudisoma roseum (Victor, 1839), ZFMK MYR11859, midbody leg, tarsal claw. F. Midbody leg, posterior view. Abbreviations: Cx = coxa; cp = coxal pouch; Fe = femur; Go = tip of male gonopore; oz = ozopore; P = pleurite; paro = paronychium; Po = postfemur; Pre = prefemur; St = sternite; T = tergite; Ta = tarsus; Ti = tibia; tr = trochanter." captionText-3="FIGURE 9. Dawydoffia siphonocryptida n. sp., male holotype (SMF), scanning electron micrographs. A. Sternite 1 and 2, coxa of leg pair 2 with gonapophysis or pseudopenis, ventral view.Arrows point to sternal protuberances. B. Left gonapophysis or pseudopenis, ventral view. C. Anterior and posterior gonopod, lateral view. D. Gonopods, anterior view. E. Anterior and posterior gonopod, posterior view. F. Detail of podomere 5, posterior gonopod. Abbreviations: aG = anterior gonopod; aG-P = process on anterior gonopod; cp = coxal pouch; Cx = coxa; H = head; LP = lateral process of podomere 6 of anterior gonopod; MP = mesal process of podomere 6 of anterior gonopod; pG = posterior gonopod; PP = pseudopenis; St = sternite; St-p = sternite process." pageId="11" pageNumber="422">Figures 69</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="422" type="etymology">
<paragraph blockId="11.[151,1436,1976,2039]" pageId="11" pageNumber="422">
<emphasis bold="true" box="[151,280,1976,2002]" pageId="11" pageNumber="422">Etymology</emphasis>
: The first authors first thought was, when observing this species, that it belongs to the Siphonocryptida, whose members resemble this species in their habitus. Noun in apposition.
</paragraph>
</subSubSection>
<caption pageId="12" pageNumber="423" startId="12.[151,250,980,1004]" targetBox="[168,1429,185,950]" targetPageId="12">
<paragraph blockId="12.[151,1436,980,1077]" pageId="12" pageNumber="423">
<emphasis bold="true" box="[151,273,980,1004]" pageId="12" pageNumber="423">FIGURE 6.</emphasis>
<taxonomicName authority="Wesener &amp; Moritz &amp; Akkari, 2023" authorityName="Wesener &amp; Moritz &amp; Akkari" authorityYear="2023" box="[278,566,980,1004]" class="Diplopoda" family="Siphonotidae" genus="Dawydoffia" kingdom="Animalia" order="Polyzoniida" pageId="12" pageNumber="423" phylum="Arthropoda" rank="species" species="siphonocryptida" status="n. sp.">
<emphasis box="[278,566,980,1004]" italics="true" pageId="12" pageNumber="423">Dawydoffia siphonocryptida</emphasis>
</taxonomicName>
<emphasis bold="true" box="[570,625,981,1005]" pageId="12" pageNumber="423">
<taxonomicNameLabel box="[570,625,981,1005]" pageId="12" pageNumber="423" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female paratype (SMF), volume rendering based on µCT-data.
<emphasis bold="true" box="[1260,1285,980,1004]" pageId="12" pageNumber="423">A.</emphasis>
Habitus lateral view.
<emphasis bold="true" box="[211,234,1017,1041]" pageId="12" pageNumber="423">B.</emphasis>
Head and anterior body, ventral view.
<emphasis bold="true" box="[623,648,1016,1040]" pageId="12" pageNumber="423">C.</emphasis>
Telson and posterior body, ventral view.
<emphasis bold="true" box="[1061,1222,1016,1040]" pageId="12" pageNumber="423">Abbreviations:</emphasis>
At = Antennae; Co = collum; ep = epicranium; fh = forehead; oz = ozopore, Pl = pleurite; St = sternite; T = tergite; Te = telson; Vu = vulva.
</paragraph>
</caption>
<subSubSection pageId="12" pageNumber="423" type="materials_examined">
<paragraph blockId="12.[151,1437,1122,1401]" box="[151,385,1122,1149]" pageId="12" pageNumber="423">
<emphasis bold="true" box="[151,377,1122,1148]" pageId="12" pageNumber="423">Material examined</emphasis>
:
</paragraph>
<paragraph blockId="12.[151,1437,1122,1401]" pageId="12" pageNumber="423">
<materialsCitation collectingDate="2008-02-28" collectionCode="SMF, SEM" collectorName="P. Jager" country="Laos" county="Nam Khan" elevation="372" location="Ban Keng Koung" longitude="102.31228" municipality="Xien Ngeun Dist." pageId="12" pageNumber="423" specimenCount="1" stateProvince="Louangphabang" typeStatus="holotype">
<typeStatus box="[199,308,1159,1185]" pageId="12" pageNumber="423">Holotype</typeStatus>
: 1 ♁,
<emphasis bold="true" box="[371,430,1158,1184]" pageId="12" pageNumber="423">
<collectionCode box="[371,430,1158,1184]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="423">SMF</collectionCode>
</emphasis>
, broken and utilized for
<collectionCode box="[701,759,1159,1185]" pageId="12" pageNumber="423">SEM</collectionCode>
and genetic sampling.
<collectingCountry box="[1013,1070,1159,1185]" name="Laos" pageId="12" pageNumber="423">Laos</collectingCountry>
,
<collectingRegion box="[1080,1316,1159,1185]" country="Laos" name="Louangphabang" pageId="12" pageNumber="423">Luang Prabang Prov.</collectingRegion>
, SE
<collectingRegion country="Laos" name="Louangphabang" pageId="12" pageNumber="423">Luang Prabang</collectingRegion>
,
<collectingCounty box="[257,381,1195,1221]" pageId="12" pageNumber="423">Nam Khan</collectingCounty>
,
<collectingMunicipality box="[393,593,1195,1221]" pageId="12" pageNumber="423">Xien Ngeun Dist.</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:29675F45FFA8FFFDFF5A6A2EFA8FF988:A411B888FFAFFFFDFD9169CEFC9AFB3C" box="[604,802,1195,1221]" country="Laos" county="Nam Khan" longitude="102.31228" municipality="Xien Ngeun Dist." name="Ban Keng Koung" pageId="12" pageNumber="423" stateProvince="Louangphabang">Ban Keng Koung</location>
,
<quantity box="[814,886,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="12" pageNumber="423" unit="m" value="372.0">
<elevation box="[814,886,1195,1221]" metricMagnitude="2" metricUnit="m" metricValue="3.72" pageId="12" pageNumber="423" unit="m" value="372.0">372 m</elevation>
</quantity>
, 19°40'96.3'' N,
<geoCoordinate box="[1071,1242,1195,1221]" degrees="102" direction="east" minutes="18" orientation="longitude" pageId="12" pageNumber="423" precision="1" seconds="44.2" value="102.31228">102°18'44.2'' E</geoCoordinate>
, disturbed forest, valley, waterfall, forest soil,
<collectingMethod box="[470,590,1231,1257]" pageId="12" pageNumber="423">pitfall trap</collectingMethod>
, leg.
<collectorName box="[650,736,1231,1257]" pageId="12" pageNumber="423">P. Jäger</collectorName>
21
<date box="[783,900,1231,1257]" pageId="12" pageNumber="423" value="2008-02-28">
<collectingDate box="[783,900,1231,1257]" pageId="12" pageNumber="423" value="2008-02-28">28.2.2008</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="12.[151,1437,1122,1401]" pageId="12" pageNumber="423">
<materialsCitation box="[199,1371,1266,1293]" collectingDate="2008-02-23" collectionCode="SMF" collectorName="Winkler &amp; P. Jager" country="Laos" county="Nam Khan" elevation="372" location="Ban Keng Koung" longitude="102.31228" municipality="Xien Ngeun Dist." pageId="12" pageNumber="423" specimenCount="1" specimenCount-female="1" stateProvince="Louangphabang" typeStatus="paratype">
<typeStatus box="[199,312,1267,1293]" pageId="12" pageNumber="423">Paratypes</typeStatus>
:
<specimenCount box="[325,370,1267,1293]" pageId="12" pageNumber="423" type="female">1 ♀</specimenCount>
,
<emphasis bold="true" box="[386,445,1266,1292]" pageId="12" pageNumber="423">
<collectionCode box="[386,445,1266,1292]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="423">SMF</collectionCode>
</emphasis>
, same data as holotype, but coll. by
<collectorName box="[873,964,1267,1293]" pageId="12" pageNumber="423">Winkler</collectorName>
extraction, leg.
<collectorName box="[1152,1242,1267,1293]" pageId="12" pageNumber="423">P. Jäger</collectorName>
,
<date box="[1255,1371,1267,1293]" pageId="12" pageNumber="423" value="2008-02-23">
<collectingDate box="[1255,1371,1267,1293]" pageId="12" pageNumber="423" value="2008-02-23">23.2.2008</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2007-03-24" collectionCode="SMF" collectorName="P. Jager" country="Laos" elevation="363" latitude="19.747557" location="Nam Khan" longLatPrecision="1" longitude="102.22095" pageId="12" pageNumber="423" specimenCount="1" specimenCount-female="1" stateProvince="Louangphabang" typeStatus="paratype">
<specimenCount box="[1385,1429,1267,1293]" pageId="12" pageNumber="423" type="female">1 ♀</specimenCount>
,
<emphasis bold="true" box="[151,210,1302,1328]" pageId="12" pageNumber="423">
<collectionCode box="[151,210,1302,1328]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="12" pageNumber="423">SMF</collectionCode>
</emphasis>
,
<collectingCountry box="[227,284,1303,1329]" name="Laos" pageId="12" pageNumber="423">Laos</collectingCountry>
,
<collectingRegion box="[299,543,1303,1329]" country="Laos" name="Louangphabang" pageId="12" pageNumber="423">Luang Prabang Prov.</collectingRegion>
, SE
<collectingRegion box="[599,776,1303,1329]" country="Laos" name="Louangphabang" pageId="12" pageNumber="423">Luang Prabang</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:29675F45FFA8FFFDFF5A6A2EFA8FF988:A411B888FFAFFFFDFCDB6872FC2DFAC8" box="[790,917,1303,1329]" country="Laos" latitude="19.747557" longLatPrecision="1" longitude="102.22095" name="Nam Khan" pageId="12" pageNumber="423" stateProvince="Louangphabang">Nam Khan</location>
,
<locationDeviation firstLocation="Ban Khon Why" pageId="12" pageNumber="423" secondLocation="Xieng Ngeun, Houay Tham">between Ban Khon Why and Xieng Ngeun, Houay Tham</locationDeviation>
,
<quantity box="[309,381,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="3.63" pageId="12" pageNumber="423" unit="m" value="363.0">
<elevation box="[309,381,1339,1365]" metricMagnitude="2" metricUnit="m" metricValue="3.63" pageId="12" pageNumber="423" unit="m" value="363.0">363 m</elevation>
</quantity>
,
<geoCoordinate box="[391,551,1338,1365]" degrees="19" direction="north" minutes="44" orientation="latitude" pageId="12" pageNumber="423" precision="1" seconds="51.2" value="19.747557">19°44'51.2'' N</geoCoordinate>
,
<geoCoordinate box="[561,731,1338,1365]" degrees="102" direction="east" minutes="13" orientation="longitude" pageId="12" pageNumber="423" precision="1" seconds="15.4" value="102.22095">102°13'15.4'' E</geoCoordinate>
, along stream, leaf litter at rocks, daytime, sieving, leg.
<collectorName box="[1346,1432,1339,1365]" pageId="12" pageNumber="423">P. Jäger</collectorName>
,
<date box="[151,267,1375,1401]" pageId="12" pageNumber="423" value="2007-03-24">
<collectingDate box="[151,267,1375,1401]" pageId="12" pageNumber="423" value="2007-03-24">24.3.2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation box="[278,831,1374,1401]" collectingDate="2007-03-24" collectionCode="ZFMK" collectorName="P. Jager" country="Laos" elevation="363" latitude="19.747557" location="Nam Khan" longLatPrecision="1" longitude="102.22095" pageId="12" pageNumber="423" specimenCode="MYR 12266" specimenCount="1" specimenCount-female="1" stateProvince="Louangphabang" typeStatus="paratype">
<specimenCount box="[278,320,1375,1401]" pageId="12" pageNumber="423" type="female">1 ♀</specimenCount>
,
<emphasis bold="true" box="[334,569,1374,1401]" pageId="12" pageNumber="423">
<collectionCode box="[334,418,1375,1401]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="12" pageNumber="423" type="Museum">ZFMK</collectionCode>
<specimenCode box="[425,569,1374,1401]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="12" pageNumber="423" type="Museum">MYR 12266</specimenCode>
</emphasis>
, same data as previous
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="423" type="diagnosis">
<paragraph blockId="12.[151,1437,1442,1649]" box="[151,274,1442,1469]" pageId="12" pageNumber="423">
<emphasis bold="true" box="[151,266,1442,1468]" pageId="12" pageNumber="423">Diagnosis</emphasis>
:
</paragraph>
<paragraph blockId="12.[151,1437,1442,1649]" pageId="12" pageNumber="423">
<taxonomicName authority="Wesener &amp; Moritz &amp; Akkari, 2023" authorityName="Wesener &amp; Moritz &amp; Akkari" authorityYear="2023" box="[199,412,1479,1505]" class="Diplopoda" family="Siphonotidae" genus="Dawydoffia" kingdom="Animalia" order="Polyzoniida" pageId="12" pageNumber="423" phylum="Arthropoda" rank="species" species="siphonocryptida" status="n. sp.">
<emphasis box="[199,412,1479,1505]" italics="true" pageId="12" pageNumber="423">D siphonocryptida</emphasis>
</taxonomicName>
<emphasis bold="true" box="[422,488,1479,1505]" pageId="12" pageNumber="423">
<taxonomicNameLabel box="[422,488,1479,1505]" pageId="12" pageNumber="423" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName authorityName="Attems" authorityYear="1953" box="[647,783,1479,1505]" class="Diplopoda" family="Siphonotidae" genus="Dawydoffia" kingdom="Animalia" order="Polyzoniida" pageId="12" pageNumber="423" phylum="Arthropoda" rank="species" species="kalonota">
<emphasis box="[647,783,1479,1505]" italics="true" pageId="12" pageNumber="423">D. kalonota</emphasis>
</taxonomicName>
in the coloration: body white, with black stripe starting at tergite 3 (body red-brown, with conspicuous black-brown stripe running from collum to anal segment in
<taxonomicName authorityName="Attems" authorityYear="1953" class="Diplopoda" family="Siphonotidae" genus="Dawydoffia" kingdom="Animalia" order="Polyzoniida" pageId="12" pageNumber="423" phylum="Arthropoda" rank="species" species="kalonota">
<emphasis italics="true" pageId="12" pageNumber="423">D. kalonota</emphasis>
</taxonomicName>
). It also differs from
<taxonomicName authorityName="Attems" authorityYear="1953" box="[493,625,1551,1577]" class="Diplopoda" family="Siphonotidae" genus="Dawydoffia" kingdom="Animalia" order="Polyzoniida" pageId="12" pageNumber="423" phylum="Arthropoda" rank="species" species="kalonota">
<emphasis box="[493,625,1551,1577]" italics="true" pageId="12" pageNumber="423">D. kalonota</emphasis>
</taxonomicName>
in the number of conical protuberances on the second sternite (4 versus 2) and in the posterior gonopod, with podomere 2 being the largest, as long as (but wider then) podomeres 3 and 4 combined (in
<taxonomicName authorityName="Attems" authorityYear="1953" box="[307,439,1623,1649]" class="Diplopoda" family="Siphonotidae" genus="Dawydoffia" kingdom="Animalia" order="Polyzoniida" pageId="12" pageNumber="423" phylum="Arthropoda" rank="species" species="kalonota">
<emphasis box="[307,439,1623,1649]" italics="true" pageId="12" pageNumber="423">D. kalonota</emphasis>
</taxonomicName>
with podomere 3 being larger than 2, cylindrical, as long as 4 and 5 combined).
</paragraph>
</subSubSection>
</treatment>
</document>