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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1128.89751" ID-Pensoft-Pub="1313-2970-1128-129" ID-Pensoft-UUID="9462C7DFB85D5DC780F645BC75150827" ID-ZooBank="7AC1EB83EE914109892AE5714A65EFB9" ModsDocID="1313-2970-1128-129" checkinTime="1668024534632" checkinUser="pensoft" docAuthor="Saxton, Natalie A., Marinov, Milen G. &amp; Bybee, Seth M." docDate="2022" docId="1CE81D420BD9534DAEE8C965351459BE" docLanguage="en" docName="ZooKeys 1128: 129-169" docOrigin="ZooKeys 1128" docPubDate="2022-11-09" docSource="http://dx.doi.org/10.3897/zookeys.1128.89751" docTitle="Vanuatubasis evelynae Saxton &amp; Marinov &amp; Bybee 2022, sp. nov." docType="treatment" docUuid="5974BE63-DD97-411B-998E-6BD01F582838" docUuidSource="ZooBank" docVersion="1" id="9462C7DFB85D5DC780F645BC75150827" lastPageNumber="129" masterDocId="9462C7DFB85D5DC780F645BC75150827" masterDocTitle="Revision of Vanuatubasis Ober &amp; Staniczek, 2009 (Odonata, Coenagrionidae), with description of seven new species" masterLastPageNumber="169" masterPageNumber="129" pageNumber="129" updateTime="1668024534632" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Vanuatubasis Ober &amp; Staniczek, 2009 (Odonata, Coenagrionidae), with description of seven new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saxton, Natalie A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5993-9782</mods:nameIdentifier>
<mods:affiliation>Research and Collections Division, The Cleveland Museum of Natural History, Cleveland, OH, 44106, USA &amp; Department of Biology, Case Western Reserve University, Cleveland, OH, 44106, USA &amp; Department of Biology and Monte L. Bean Museum Brigham Young University, Provo, UT 84604, USA</mods:affiliation>
<mods:nameIdentifier type="email">nsaxton55@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marinov, Milen G.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3284-2555</mods:nameIdentifier>
<mods:affiliation>Biosecurity Surveillance and Incursion Investigation Plant Health Team, Ministry for Primary Industries, 14 Sir William Pickering Drive, Christchurch 8053, New Zealand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bybee, Seth M.</mods:namePart>
<mods:affiliation>Department of Biology and Monte L. Bean Museum Brigham Young University, Provo, UT 84604, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1128</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>169</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1128.89751</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1128.89751</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1128-129</mods:identifier>
<mods:identifier type="ZooBank">7AC1EB83EE914109892AE5714A65EFB9</mods:identifier>
<mods:identifier type="Pensoft-UUID">9462C7DFB85D5DC780F645BC75150827</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:5974BE63-DD97-411B-998E-6BD01F582838" httpUri="http://treatment.plazi.org/id/1CE81D420BD9534DAEE8C965351459BE" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
<taxonomicName LSID="https://zoobank.org/5974BE63-DD97-411B-998E-6BD01F582838" authority="Saxton &amp; Marinov &amp; Bybee, 2022" authorityName="Saxton &amp; Marinov &amp; Bybee" authorityYear="2022" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis evelynae" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="evelynae" status="sp. nov.">Vanuatubasis evelynae</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="129">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Vanuatubasis evelynae Holotype (♂ BYU) A dorsal thorax B lateral terminalia C dorsal terminalia D lateral habitus V. evelynae Allotype (♀ BYU) E dorsal thorax F lateral terminalia. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1128.89751.figure4" httpUri="https://binary.pensoft.net/fig/767329" pageId="0" pageNumber="129">Figs 4</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Male dorso-posterior and dorsal terminalia of A V. bidens B V. evelynae C V. insularivorum D V. kapularum E V. malekulana F V. nunggoli G V. santoensis." figureDoi="10.3897/zookeys.1128.89751.figure20" httpUri="https://binary.pensoft.net/fig/767346" pageId="0" pageNumber="129">, 20B</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female face and dorsal terminalia of A V. bidens B V. discontinua C V. evelynae D V. insularivorum E V. kapularum F V. malekulana G V. nunggoli H V. rhomboides I V. santoensis J V. xanthochroa." figureDoi="10.3897/zookeys.1128.89751.figure21" httpUri="https://binary.pensoft.net/fig/767347" pageId="0" pageNumber="129">, 21C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="type material">
<paragraph pageId="0" pageNumber="129">Type material.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Holotype</emphasis>
(1
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU).</emphasis>
&quot;VANUATU: Santo Is: | Coulons, -15.2957 | 167.1616092, 28 v. 2018, | coll. S. Bybee and G.
<normalizedToken originalValue="Powell">Powell''</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Paratypes</emphasis>
(1
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">1</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BPBM, 7</emphasis>
♂♂
<emphasis bold="true" pageId="0" pageNumber="129">BYU, 3</emphasis>
♂♂
<emphasis bold="true" pageId="0" pageNumber="129">NHM, 3</emphasis>
♂♂
<emphasis bold="true" pageId="0" pageNumber="129">NZAC). (1</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">1</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BPBM, 3</emphasis>
♂♂
<emphasis bold="true" pageId="0" pageNumber="129">BYU, 3</emphasis>
♂♂
<emphasis bold="true" pageId="0" pageNumber="129">NHM, 3</emphasis>
♂♂
<emphasis bold="true" pageId="0" pageNumber="129">NZAC)</emphasis>
Same label data as holotype.
<emphasis bold="true" pageId="0" pageNumber="129">(1</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU)</emphasis>
&quot;VANUATU: Santo Is., | Coulons, May 20, 2019 | -15. 2955, 167.1614 | coll. SM. Bybee, GS. Powell | #BYU-VU-2019
<normalizedToken originalValue="">''</normalizedToken>
<normalizedToken originalValue="“OD1710”">&quot;OD1710&quot;</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="129">(1</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU cryo)</emphasis>
&quot;VANUATU: Santo Is: | Pelmol,
<geoCoordinate degrees="15.6078" direction="south" orientation="latitude" precision="5" value="-15.6078">-15.6078</geoCoordinate>
,
<geoCoordinate degrees="166.7849" direction="east" orientation="longitude" precision="5" value="166.7849">166.7849</geoCoordinate>
, | 4.vi.2018, |coll. S. Bybee and | G. Powell&quot;
<normalizedToken originalValue="“OD1709”">&quot;OD1709&quot;</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="129">(1</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU cryo)</emphasis>
same data as holotype;
<normalizedToken originalValue="“OD1710”">&quot;OD1710&quot;</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="129">(1</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU cryo)</emphasis>
&quot;VANUATU: Santo Is: | Coulons, -15.2957 | 167.1616092, 28 v. 2018, | coll. S. Bybee and G.
<normalizedToken originalValue="Powell">Powell''</normalizedToken>
<normalizedToken originalValue="“OD1711”">&quot;OD1711&quot;</normalizedToken>
.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Allotype</emphasis>
(1
</emphasis>
<emphasis bold="true" pageId="0" pageNumber="129">BYU).</emphasis>
same label data as holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Head</emphasis>
: Labium overall pale beige; labrum blue-green, darkening posteriorly, with dark brown postero-lateral edges and medially, a black spot at posterior edge; anteclypeus dark green; genae, and mandibles (expect for reddish brown tips) green with paler anterolateral edges; postclypeus overall covered by a black bar that begins medially and extends to the anterior edge, not extending to the dark green, lateral corners; frons dark green, with medial black and extending posteriorly; scapes, and pedicels black, flagella dark brown and lightening apically; vertex and rear of head black with line of setae, a bronze shimmer, and white pruinescence; a pair of white post-ocular spots present; three pale ocelli with a yellow patch apical of the median ocellus; line of setae behind vertex; eyes yellowish green.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Thorax</emphasis>
: Prothorax dorsally dark green to black with a bronze shimmer; laterally blue and green; pronotum black, latero-posterior corners rounded to obtuse angles and explanate, mid-line obviously indented across the pronotum, hind lobe with raised ridge that is shorter than the width of the pronotum, is curved outward medially, and extends to a sharp point that protrudes posteriorly; mesostigmal plate dark brown with yellow-green edges, interior edges raised to form protruding lobes. Pterothorax with black carina; laterally, mesepisternum with black stripe reaching the dorsal carina and extending to the mesopleural suture posteriorly across ~0.5 mm, but only reaching ~ 2/3 of the mesepisternum on the anterior and medial portion; green stripe located on latter 1/3 of mesepisternum and extending past the dark brown mesopleural suture; mesepimeron overall blue with green extending down from mesepisternum, metepisternum overall blue with short, brown line located on metapleural suture near the base of the wings, extending ~ 1/6 the
<normalizedToken originalValue="sutures">suture's</normalizedToken>
length; mesinfraepisternum bluish green with dark brown spot located medially; metepimeron pale blue with white pruinescence; coxae, trochanters, and femora dorsally brown to black and ventrally pale beige with black spines; tibiae pale brown with smaller spines than that of the femora; tarsi beige with dark brown edges and small, dense spines; pale brown tarsal claws that darken apically to reddish tips, claws with a small tooth located on the basal 1/4 of their length.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Wings</emphasis>
: Hyaline; venation dark brown and thickening towards the dorsal edge; pterostigma dark brown and rhombus-shaped, with edges being the darkest; CuP slightly distal than halfway between antenodals in HW; arculus originates just distal of the second antenodal crossvein in HW and slightly proximal to it in HW; discoidal cells unequal with FW dorsal edge being 1/2 as long as HW. Nodal index: 12/2-2/12 in FW and 11/2-2/10 in HW.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Abdomen</emphasis>
: Overall yellowish green with black dorsal stripe extending from S1-S9 and lightening to brown laterally, brown reaching ventrally at posterior edge of S1-S4, pale brown setae; segment 1 blue; S2 blue with patch of beige at anterior end; S3 anteriorly dark blue, turning yellow posteriorly, S4-S8 yellow with patches of blue on terminal end of each segment; S9 overall brown with blue dorsal stripe extending medially lengthwise, and light patch medially in lateral view; S10 with blue patch laterally that extends to ventral surface, terminal edge raised medially to form protrusion; cerci dark brown with gold setae and appearing as a curved crescent shape dorsally, excavated medially on each lobe and sloping posteriorly; paraprocts in dorsal view dark brown and darkening apically, with bumpy texture; in the lateral perspective the lobes are roughly triangular and tapering apically to form small, acutely rounded lobes, with dorsal edge slightly uneven and ventral edge expanded to form small
<normalizedToken originalValue="“hump”">&quot;hump&quot;</normalizedToken>
, terminating in small, acutely rounded lobe.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Measurements</emphasis>
(mm)
</emphasis>
: total length 38 mm, abdomen 33 mm, HW 20 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">Description of allotype.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Head</emphasis>
: Labium overall pale beige; labrum blue-green, with black latero-posterior edges and a medially black spot at posterior edge; anteclypeus, genae, and mandibles (expect for reddish tips) blue-green; postclypeus darker blue with a small bar located on the medial, anterior edge; frons blue-green, abruptly changing to black posteriorly; scapes, and pedicels black, flagella dark brown and lightening apically; vertex and rear of head black with line of setae, a bronze shimmer, and white pruinescence; a pair of white post-ocular spots present; three pale ocelli with a yellow patch apical of the median ocellus; eyes green.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Thorax</emphasis>
: Prothorax dorsally dark green to black with bronze shimmer; laterally blue; pronotum black medially with blue edges, latero-posterior corners rounded to obtuse angles and weakly explanate, mid-line obviously indented across the pronotum, hind lobe with raised ridge that is shorter than the width of the pronotum, is curved outward medially, and extends to a sharp point that protrudes posteriorly; mesostigmal plate black with green lateral edges, roughly triangular and not significantly raised. Pterothorax with black carina; laterally, mesepisternum with black stripe reaching the dorsal carina and reaching the mesopleural suture, but only reaching ~ 2/3 of the mesepisternum; dark green stripe located on latter 1/3 of mesepisternum, not quite reaching mesinfraepisternum, turning yellow as it passes the mesopleural suture; mesepimeron overall blue with yellow extending down from mesepisternum; dark brown line on posterior end of mesopleural suture extending ~1/6 the sutures length; metepisternum overall blue with short, black line located on metapleural suture near the base of the wings, extending ~the
<normalizedToken originalValue="sutures">suture's</normalizedToken>
length; mesinfraepisternum blue; metepimeron blue; coxae, trochanters, and femora dorsally pale brown with blue patches, and ventrally pale beige with black spines; tibiae pale brown with slightly darker, and smaller, spines than that of the femora; tarsi beige with dark brown edges and small, dense spines; pale brown tarsal claws that darken apically to reddish tips, claws with a small tooth located on the basal 1/4 of their length.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Wings</emphasis>
: Hyaline with dark brown venation, thickening towards the dorsal edge. Pterostigma dark brown and rhombus-shaped, with edges being the darkest; CuP slightly distal than halfway between antenodals in HW, and approximately halfway in FW; arculus originates at the second antenodal crossvein in front wings and slightly distal to it in HW; discoidal cells unequal with FW dorsal edge being 1/2 as long as HW. Nodal index: 13/2-2/11 in FW and 13/2-2/11 in HW.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" italics="true" pageId="0" pageNumber="129">Abdomen</emphasis>
: Overall yellow with black dorsal stripe extending from S1-S9 and lightening laterally, with pale brown setae; S1 and S2 blue laterally; S3 blue anteriorly but turns to pale yellow posteriorly; S4-S8 pale yellow laterally; S9 with dorsal blue patch extending
<normalizedToken originalValue="¾">3/4</normalizedToken>
of its length; S10 blue dorsally and laterally. Ovipositor overall pale yellow and reddish brown ventrally, with serrated ventral edge; stylus with rounded edges, dark brown and lightening apically; gonapophysis dark reddish -brown with slightly serrated dorsal edge. Cerci roughly triangular, dark brown, and narrowing to a rounded apex, dorsal edge slightly rounded.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Measurements</emphasis>
(mm)
</emphasis>
: total length 35 mm, abdomen 28 mm, HW 20 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Male.</emphasis>
<taxonomicName authorityName="Saxton &amp; Marinov &amp; Bybee" authorityYear="2022" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis evelynae" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="evelynae">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis evelynae</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis</emphasis>
</taxonomicName>
species by a black pterostigma, forked (longer than wide) lateral lobes of the genital ligula, cerci curving medially their entire length, and black postclypeus with green lateral margins.
<emphasis bold="true" pageId="0" pageNumber="129">Female.</emphasis>
<taxonomicName authorityName="Saxton &amp; Marinov &amp; Bybee" authorityYear="2022" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis evelynae" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="evelynae">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis evelynae</emphasis>
</taxonomicName>
can be distinguished from other females in this genus, by their dark colored cerci, postclypeus with dark maculation, and uneven dorsal black area on S9 appearing almost sinusoidal.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="variation">
<paragraph pageId="0" pageNumber="129">Variation.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">Male.</emphasis>
Immature specimens are pale blue with less thoracic coloring.
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Measurements</emphasis>
(mm)
</emphasis>
: length 37-38 mm, abdomen 29-33 mm, HW 19-20 mm (
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
= 5).
<emphasis bold="true" pageId="0" pageNumber="129">Female.</emphasis>
The color of the pterostigma is lighter.
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Measurements</emphasis>
(mm)
</emphasis>
: total length 35-36, abdomen 28-29 mm, HW 20-21 (
<emphasis italics="true" pageId="0" pageNumber="129">n</emphasis>
= 2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">Espiritu Santo, Vanuatu.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="etymology">
<paragraph pageId="0" pageNumber="129">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="129">
The specific epithet of this species is a Latinized noun in the genitive case of the name
<normalizedToken originalValue="Evelyn">'Evelyn'</normalizedToken>
in honor of L. Evelyn Cheesman, a prominent female entomologist whose early expeditions in Vanuatu significantly paved the way for future work in this region.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="notes">
<paragraph pageId="0" pageNumber="129">Notes.</paragraph>
<paragraph pageId="0" pageNumber="129">
This species is likely the one referred to as &quot;
<taxonomicName authorityName="Ober &amp; Staniczek" authorityYear="2009" class="Insecta" family="Coenagrionidae" genus="Vanuatubasis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Vanuatubasis" order="Odonata" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Vanuatubasis</emphasis>
</taxonomicName>
sp.&quot; in
<bibRefCitation author="Staniczek, A" editor="Bouchet, PH" journalOrPublisher="Museum national d'Histoire naturelle, Paris" pageId="0" pageNumber="129" pagination="251 - 257" refId="B14" refString="Staniczek, A, 2011. Focus on aquatic insects. In: Bouchet, PH, Guyader, HL, Pascal, O, Eds., The natural history of Santo. Museum national d'Histoire naturelle, Paris: 251 - 257" title="Focus on aquatic insects." volumeTitle="The natural history of Santo." year="2011">Staniczek (2011)</bibRefCitation>
. With the exception of one individual, all specimens were collected at the same site (Coulons, Espiritu Santo). One
<normalizedToken originalValue="individuals">individuals'</normalizedToken>
label data reflects a locality on the other side of the island (Pelmol, Espiritu Santo). While this disjunct range is possible, it does seem suspect and is likely an error.
</paragraph>
</subSubSection>
</treatment>
</document>