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<document id="814840EB6798DA911A1382B732B3DD96" ID-CLB-Dataset="107636" ID-DOI="10.5852/ejt.2022.833.1879" ID-GBIF-Dataset="69674de4-93c1-463c-9ae1-2a9939cfc966" ID-ISSN="2118-9773" ID-Zenodo-Dep="6916920" ID-ZooBank="EADA40BF-86E5-4D5B-8D43-3F6FB8F2C687" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658962515064" checkinUser="felipe" docAuthor="Wichard, Wilfried &amp; Espeland, Marianne" docDate="2022" docId="1A5A8785FF871568C97BFA0206A2E7B9" docLanguage="en" docName="ejt-833_1-11.pdf.imf" docOrigin="European Journal of Taxonomy 833" docSource="http://zoobank.org/eada40bf-86e5-4d5b-8d43-3f6fb8f2c687" docStyle="DocumentStyle:8DBBCD0342F59C7B93E4D60327A07408.3:EJT.2018-.journal_article.type2" docStyleId="8DBBCD0342F59C7B93E4D60327A07408" docStyleName="EJT.2018-.journal_article.type2" docStyleVersion="3" docTitle="Cretapsychidae Wichard 2021" docType="treatment" docVersion="7" lastPageNumber="5" masterDocId="E663FFFDFF85156CCB2CFFBE0526E53A" masterDocTitle="The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1699362387126" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="19CEED04BEB7A0E253D270FE96306F72">The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species</mods:title>
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<mods:namePart id="C81184F3E948A736E12A2386FF6160BC">Wichard, Wilfried</mods:namePart>
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<mods:affiliation id="C801A85175E6F9FC841D14E2A3A11341">Institute of Biology, University of Cologne, Herbert Lewinstrasse 2, 50931 Köln, Germany.</mods:affiliation>
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<mods:namePart id="DD92A8E84DFD387007AA3E0842005763">Espeland, Marianne</mods:namePart>
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<mods:affiliation id="6592FCC72BD005E17C1DA4D6321E6020">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
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<mods:date id="124E6F26CA346104878D114E20559843">2022</mods:date>
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<treatment id="1A5A8785FF871568C97BFA0206A2E7B9" ID-DOI="http://doi.org/10.5281/zenodo.6923660" ID-GBIF-Taxon="197864950" ID-Zenodo-Dep="6923660" LSID="urn:lsid:plazi:treatment:1A5A8785FF871568C97BFA0206A2E7B9" httpUri="http://treatment.plazi.org/id/1A5A8785FF871568C97BFA0206A2E7B9" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
<subSubSection id="DAE96518FF87156EC97BFA02010FE0EC" box="[599,1065,1467,1494]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="924C3693FF87156EC97BFA02010FE0EC" blockId="2.[599,1065,1467,1494]" box="[599,1065,1467,1494]" pageId="2" pageNumber="3">
Family
<taxonomicName id="55F34D10FF87156EC99DFA05010FE0EC" authority="Wichard, 2021" authorityName="Wichard" authorityYear="2021" box="[689,1065,1467,1494]" class="Insecta" family="Cretapsychidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">
<emphasis id="A087EA81FF87156EC99DFA050652E0EF" bold="true" box="[689,884,1467,1493]" pageId="2" pageNumber="3">Cretapsychidae</emphasis>
<bibRefCitation id="F6624B62FF87156EC857FA02010FE0EC" author="Wichard W." box="[891,1065,1468,1494]" pageId="2" pageNumber="3" pagination="104707" refId="ref3509" refString="Wichard W. 2021. Overview of the caddisflies (Insecta, Trichoptera) in mid-Cretaceous Burmese amber. Cretaceous Research 119: e 104707. https: // doi. org / 10.1016 / j. cretres. 2020.104707" type="journal article" year="2021">Wichard, 2021</bibRefCitation>
</taxonomicName>
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<subSubSection id="DAE96518FF87156ECBCFFA40065EE37D" pageId="2" pageNumber="3" type="type_taxon">
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<heading id="C90481FFFF87156ECBCFFA40045BE32D" bold="true" box="[227,381,1533,1560]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="A087EA81FF87156ECBCFFA40045BE32D" bold="true" box="[227,381,1533,1560]" pageId="2" pageNumber="3">
<typeStatus id="4D488831FF87156ECBCFFA400406E322" box="[227,288,1534,1560]" pageId="2" pageNumber="3">Type</typeStatus>
species
</emphasis>
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</paragraph>
<paragraph id="924C3693FF87156ECBCFF993065EE37D" blockId="2.[227,888,1581,1607]" box="[227,888,1581,1607]" pageId="2" pageNumber="3">
<taxonomicName id="55F34D10FF87156ECBCFF9930652E37D" authority="Wichard, Neumann, Muller &amp; Wang, 2018" authorityName="Wichard, Neumann, Muller &amp; Wang" authorityYear="2018" box="[227,884,1581,1607]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A087EA81FF87156ECBCFF9930453E37D" box="[227,373,1581,1607]" italics="true" pageId="2" pageNumber="3">Cretapsyche</emphasis>
<bibRefCitation id="F6624B62FF87156ECA50F9930652E37D" author="Wichard W. &amp; Neumann C. &amp; Muller P. &amp; Wang B." box="[380,884,1581,1607]" pageId="2" pageNumber="3" pagination="138 - 146" refId="ref3595" refString="Wichard W., Neumann C., Muller P. &amp; Wang B. 2018. Family Dysoneuridae (Insecta, Trichoptera) in Cretaceous Burmese amber. Cretaceous Research 82: 138 - 146. https: // doi. org / 10.1016 / j. cretres. 2017.10.008" type="journal article" year="2018">Wichard, Neumann, Müller &amp; Wang, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="DAE96518FF871568CBCFF9D106A2E7B9" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="924C3693FF87156ECBCFF9D104FEE3B3" blockId="2.[227,472,1647,1673]" box="[227,472,1647,1673]" pageId="2" pageNumber="3">
<heading id="C90481FFFF87156ECBCFF9D104FEE3B3" bold="true" box="[227,472,1647,1673]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="A087EA81FF87156ECBCFF9D104FEE3B3" bold="true" box="[227,472,1647,1673]" pageId="2" pageNumber="3">Revised description</emphasis>
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<paragraph id="924C3693FF87156ECBCFF9210140E272" blockId="2.[227,1439,1694,1864]" pageId="2" pageNumber="3">
HEAD. Male adult embedded in amber, light brown coloration. Large complex eyes on sides of head, slightly longer in diameter than head length. Ocelli absent. Antennae as long as, or slightly longer than forewings, each with scapus about as long as head, often bearing brush of dark setae; flagellomeres uniformly bar-shaped. Maxillary palps five-segmented; second segment longest, following three segments successively smaller and shorter; fifth segment shortest (
<figureCitation id="0AC82A16FF87156EC8C2F890017CE272" box="[1006,1114,1838,1864]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[227,270,1965,1991]" captionStartId-1="5.[227,270,1964,1990]" captionStartId-2="7.[227,270,1962,1988]" captionTargetBox-0="[258,1405,304,1917]" captionTargetBox-1="[263,1401,302,1912]" captionTargetBox-2="[258,1405,302,1920]" captionTargetId-0="figure-45@3.[258,1405,304,1917]" captionTargetId-1="figure-52@5.[263,1401,302,1912]" captionTargetId-2="figure-58@7.[258,1405,302,1920]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="7" captionText-0="Fig. 2. The 5-segmented maxillary palps of species of Cretapsyche Wichard, Neumann, Müller &amp; Wang, 2018. A–B. C. elegans Wichard, Neumann, Müller &amp; Wang, 2018. C–D. C. circula Wichard, Neumann, Müller &amp; Wang, 2018. E–F. C. palpinova Wichard &amp; Neumann, 2019." captionText-1="Fig. 3. Cretapsyche kachini sp. nov., holotype, ♂ (ZFMK-TRI000829). A. Dorsal view. B. Drawing of a forewing. C. Maxillary and labial palps. D. Drawing of the maxillary palp. E. Male genitalia in ventrolateral view, distinctly flattened and partially distorted. F. Drawing of male genitalia, left half." captionText-2="Fig. 4. Cretapsyche myanmari sp. nov., holotype, ♂ (ZFMK-TRI000830). A. Dorsal view. B. Drawing of a forewing. C. Head in dorsal view, with scapus, compound eye and maxillary palp. D. Left maxillary palp. E. Drawing of a maxillary palp. F. Male genitalia in ventral view. G. Drawing of male genitalia." figureDoi-0="http://doi.org/10.5281/zenodo.6916924" figureDoi-1="http://doi.org/10.5281/zenodo.6916926" figureDoi-2="http://doi.org/10.5281/zenodo.6916928" httpUri-0="https://zenodo.org/record/6916924/files/figure.png" httpUri-1="https://zenodo.org/record/6916926/files/figure.png" httpUri-2="https://zenodo.org/record/6916928/files/figure.png" pageId="2" pageNumber="3">Figs 2–4</figureCitation>
).
</paragraph>
<paragraph id="924C3693FF87156ECBCFF8CE00A0E2B0" blockId="2.[227,1414,1904,1930]" box="[227,1414,1904,1930]" pageId="2" pageNumber="3">THORAX. Pronotum with one pair of oval setal warts. Mesonotum without setal warts on mesoscutum.</paragraph>
<paragraph id="924C3693FF87156ECBCFF80D01D4E2F6" blockId="2.[227,1266,1970,1997]" box="[227,1266,1970,1997]" pageId="2" pageNumber="3">MESOSCUTELLUM. With one domed setal wart or pair of domed setal warts fused medially.</paragraph>
<paragraph id="924C3693FF87156ECBCFF84B04E2ED34" blockId="2.[227,452,2036,2063]" box="[227,452,2036,2063]" pageId="2" pageNumber="3">TIBIAL SPURS. 2/2/4.</paragraph>
<caption id="C68C661BFF86156FCBCFF8130107ED34" ID-DOI="http://doi.org/10.5281/zenodo.6916924" ID-Zenodo-Dep="6916924" httpUri="https://zenodo.org/record/6916924/files/figure.png" pageId="3" pageNumber="4" startId="3.[227,270,1965,1991]" targetBox="[258,1405,304,1917]" targetPageId="3">
<paragraph id="924C3693FF86156FCBCFF8130107ED34" blockId="3.[226,1436,1965,2063]" pageId="3" pageNumber="4">
<emphasis id="A087EA81FF86156FCBCFF8130409E2FD" bold="true" box="[227,303,1965,1991]" pageId="3" pageNumber="4">Fig. 2.</emphasis>
The 5-segmented maxillary palps of species of
<taxonomicName id="55F34D10FF86156FC87BF8100406E2D1" authority="Wichard, Neumann, Muller &amp; Wang, 2018" authorityName="Wichard, Neumann, Muller &amp; Wang" authorityYear="2018" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="A087EA81FF86156FC87BF81006CFE2F2" box="[855,1001,1966,1992]" italics="true" pageId="3" pageNumber="4">Cretapsyche</emphasis>
<bibRefCitation id="F6624B62FF86156FC8C2F8100406E2D1" author="Wichard W. &amp; Neumann C. &amp; Muller P. &amp; Wang B." pageId="3" pageNumber="4" pagination="138 - 146" refId="ref3595" refString="Wichard W., Neumann C., Muller P. &amp; Wang B. 2018. Family Dysoneuridae (Insecta, Trichoptera) in Cretaceous Burmese amber. Cretaceous Research 82: 138 - 146. https: // doi. org / 10.1016 / j. cretres. 2017.10.008" type="journal article" year="2018">Wichard, Neumann, Müller &amp; Wang, 2018</bibRefCitation>
</taxonomicName>
.
<emphasis id="A087EA81FF86156FCA07F86F0445E2D1" bold="true" box="[299,355,2001,2027]" pageId="3" pageNumber="4">A–B</emphasis>
.
<taxonomicName id="55F34D10FF86156FCA5CF86F06C3E2D1" authority="Wichard, Neumann, Muller &amp; Wang, 2018" authorityName="Wichard, Neumann, Muller &amp; Wang" authorityYear="2018" box="[368,997,2001,2027]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="A087EA81FF86156FCA5CF86F04CAE2D1" box="[368,492,2001,2027]" italics="true" pageId="3" pageNumber="4">C. elegans</emphasis>
<bibRefCitation id="F6624B62FF86156FCADDF86F06C3E2D1" author="Wichard W. &amp; Neumann C. &amp; Muller P. &amp; Wang B." box="[497,997,2001,2027]" pageId="3" pageNumber="4" pagination="138 - 146" refId="ref3595" refString="Wichard W., Neumann C., Muller P. &amp; Wang B. 2018. Family Dysoneuridae (Insecta, Trichoptera) in Cretaceous Burmese amber. Cretaceous Research 82: 138 - 146. https: // doi. org / 10.1016 / j. cretres. 2017.10.008" type="journal article" year="2018">Wichard, Neumann, Müller &amp; Wang, 2018</bibRefCitation>
</taxonomicName>
.
<emphasis id="A087EA81FF86156FC8DCF86F010EE2D1" bold="true" box="[1008,1064,2001,2027]" pageId="3" pageNumber="4">C–D</emphasis>
.
<taxonomicName id="55F34D10FF86156FCF1BF86F04C1ED34" authority="Wichard, Neumann, Muller &amp; Wang, 2018" authorityName="Wichard, Neumann, Muller &amp; Wang" authorityYear="2018" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="circula">
<emphasis id="A087EA81FF86156FCF1BF86F018DE2D1" box="[1079,1195,2001,2027]" italics="true" pageId="3" pageNumber="4">C. circula</emphasis>
<bibRefCitation id="F6624B62FF86156FCF9CF86F04C1ED34" author="Wichard W. &amp; Neumann C. &amp; Muller P. &amp; Wang B." pageId="3" pageNumber="4" pagination="138 - 146" refId="ref3595" refString="Wichard W., Neumann C., Muller P. &amp; Wang B. 2018. Family Dysoneuridae (Insecta, Trichoptera) in Cretaceous Burmese amber. Cretaceous Research 82: 138 - 146. https: // doi. org / 10.1016 / j. cretres. 2017.10.008" type="journal article" year="2018">Wichard, Neumann, Müller &amp; Wang, 2018</bibRefCitation>
</taxonomicName>
.
<emphasis id="A087EA81FF86156FCADFF84A0701ED34" bold="true" box="[499,551,2036,2062]" pageId="3" pageNumber="4">E–F</emphasis>
.
<taxonomicName id="55F34D10FF86156FC91AF84A013CED34" authority="Wichard &amp; Neumann, 2019" authorityName="Wichard &amp; Neumann" authorityYear="2019" box="[566,1050,2036,2063]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="palpinova">
<emphasis id="A087EA81FF86156FC91AF84A07EBED34" box="[566,717,2036,2062]" italics="true" pageId="3" pageNumber="4">C. palpinova</emphasis>
<bibRefCitation id="F6624B62FF86156FC9F9F84A013CED34" author="Wichard W. &amp; Neumann C." box="[725,1050,2036,2063]" pageId="3" pageNumber="4" pagination="51 - 56" refId="ref3549" refString="Wichard W. &amp; Neumann C. 2019. A new bizarre dysoneurid species (Insecta, Trichoptera) in Burmese amber. Fossil Record 22: 51 - 56. https: // doi. org / 10.5194 / fr- 22 - 51 - 2019" type="journal article" year="2019">Wichard &amp; Neumann, 2019</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="924C3693FF811568CBCFFE910196E48E" blockId="4.[227,1437,303,436]" pageId="4" pageNumber="5">
WINGS (
<figureCitation id="0AC82A16FF811568CA6CFE9104A2E473" box="[320,388,303,329]" captionStart="Fig" captionStartId="1.[227,270,1966,1992]" captionTargetBox="[233,1432,1133,1918]" captionTargetId="figure-302@1.[227,1436,1131,1921]" captionTargetPageId="1" captionText="Fig. 1. Cretapsyche Wichard, Neumann, Müller &amp; Wang, 2018, wing venation in forewing with forks II, III, and V present, discoidal and thyridial cells open or closed, nygmata absent, in hindwing with forks I, III, and V usually present." figureDoi="http://doi.org/10.5281/zenodo.6916922" httpUri="https://zenodo.org/record/6916922/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
). Forewings broader than hindwings, each with rounded apex. In forewings, forks II, III, and V present; cubitus vein Cu2 ending in Cu1b or connected to Cu1b by crossvein. Discoidal, median, and thyridial cells present or absent. Nygmata absent. Anal vein reaching wing margin before fork V or before crossvein reaching wing margin. Hindwings with forks I, III, and V present.
</paragraph>
<paragraph id="924C3693FF811568CBCFFE6206A2E7B9" blockId="4.[227,1436,475,644]" pageId="4" pageNumber="5">MALE GENITALIA. Inferior appendages two-segmented, consisting of basal coxopodite and harpago simply attached to apex of coxopodite. Coxopodite arcuate, sub-triangular or scaly, broad, long and curved; harpago finger- or needle-shaped, often straight at base and curved mesad at apex. Two parallel appendages located centrally in genitalia interpreted as probable mesodorsal pair of lobes of tergum X. Preanal appendages small, elongate or finger-like curved.</paragraph>
</subSubSection>
</treatment>
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