164 lines
18 KiB
XML
164 lines
18 KiB
XML
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<mods:title id="3D0D42EF70BB3376B6B4605CEE6A5F94">The family Cretapsychidae (Insecta, Trichoptera) from mid- Cretaceous Burmese amber, with descriptions of two new species</mods:title>
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<mods:namePart id="D8CC4902BDB273B84BA10C48872E07A0">Wichard, Wilfried</mods:namePart>
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<mods:affiliation id="B068E303DBE0C752CFF3A8561C1106A8">Institute of Biology, University of Cologne, Herbert Lewinstrasse 2, 50931 Köln, Germany.</mods:affiliation>
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<mods:nameIdentifier id="DDCC0D12B72A8C5DDD04DBD289BF273F" type="email">Wichard@uni-koeln.de</mods:nameIdentifier>
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<mods:namePart id="243388B15DE1F23515452A58929FF87E">Espeland, Marianne</mods:namePart>
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<mods:affiliation id="75419886457D92809E2653FE6A0EB0EE">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauerallee 127, 53113 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="25ACC4A5CCEFB8350C06B636123CB25E" type="email">m.espeland@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:title id="6B956BA98B981D21E16F647BCA91951B">European Journal of Taxonomy</mods:title>
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<mods:part id="5D0DD79A54D36786A2B52E78F614119D">
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<mods:date id="E01D9150EAC18C0EE35D894F48D6AA27">2022</mods:date>
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<treatment id="1A5A8785FF83156AC9AFFCB3004BED34" ID-DOI="http://doi.org/10.5281/zenodo.6923668" ID-GBIF-Taxon="197864954" ID-Zenodo-Dep="6923668" LSID="urn:lsid:zoobank.org:act:468AEDBE-C3E8-4C1A-9AA7-578660AC0A16" httpUri="http://treatment.plazi.org/id/1A5A8785FF83156AC9AFFCB3004BED34" lastPageNumber="7" pageId="6" pageNumber="7">
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<subSubSection id="DAE96518FF83156AC9AFFCB306DAE613" box="[643,1020,781,809]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="924C3693FF83156AC9AFFCB306DAE613" blockId="6.[393,1270,781,884]" box="[643,1020,781,809]" pageId="6" pageNumber="7">
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<heading id="C90481FFFF83156AC9AFFCB306DAE613" box="[643,1020,781,809]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
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<taxonomicName id="55F34D10FF83156AC9AFFCB306BDE612" authority="Wichard & Espeland, 2022" authorityName="Wichard & Espeland" authorityYear="2022" box="[643,923,781,808]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="myanmari" status="sp. nov.">
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<emphasis id="A087EA81FF83156AC9AFFCB306BDE612" bold="true" box="[643,923,781,808]" italics="true" pageId="6" pageNumber="7">Cretapsyche myanmari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBB457FAFF83156AC88EFCB106DAE613" box="[930,1020,783,809]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="DAE96518FF83156ACAA5FC8A0644E64E" pageId="6" pageNumber="7" type="description">
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<paragraph id="924C3693FF83156ACAA5FC8A01D0E675" blockId="6.[393,1270,781,884]" box="[393,1270,820,847]" pageId="6" pageNumber="7">
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urn:lsid:zoobank.org:act:
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<uuid id="E6550C46FF83156AC984FC8A01D0E675" box="[680,1270,820,847]" pageId="6" pageNumber="7">468AEDBE-C3E8-4C1A-9AA7-578660AC0A16</uuid>
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</uri>
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</paragraph>
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<paragraph id="924C3693FF83156AC831FCE70644E64E" blockId="6.[393,1270,781,884]" box="[797,866,857,884]" pageId="6" pageNumber="7">
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<figureCitation id="0AC82A16FF83156AC831FCE70644E64E" box="[797,866,857,884]" captionStart="Fig" captionStartId="7.[227,270,1962,1988]" captionTargetBox="[258,1405,302,1920]" captionTargetId="figure-58@7.[258,1405,302,1920]" captionTargetPageId="7" captionText="Fig. 4. Cretapsyche myanmari sp. nov., holotype, ♂ (ZFMK-TRI000830). A. Dorsal view. B. Drawing of a forewing. C. Head in dorsal view, with scapus, compound eye and maxillary palp. D. Left maxillary palp. E. Drawing of a maxillary palp. F. Male genitalia in ventral view. G. Drawing of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.6916928" httpUri="https://zenodo.org/record/6916928/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DAE96518FF83156ACBCFFC23048BE160" pageId="6" pageNumber="7" type="diagnosis">
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<paragraph id="924C3693FF83156ACBCFFC23047AE68D" blockId="6.[227,348,925,951]" box="[227,348,925,951]" pageId="6" pageNumber="7">
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<heading id="C90481FFFF83156ACBCFFC23047AE68D" bold="true" box="[227,348,925,951]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
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<emphasis id="A087EA81FF83156ACBCFFC23047AE68D" bold="true" box="[227,348,925,951]" pageId="6" pageNumber="7">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="924C3693FF83156ACBCFFC71048BE160" blockId="6.[226,1444,975,1115]" pageId="6" pageNumber="7">
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<taxonomicName id="55F34D10FF83156ACBCFFC7104D2E6D2" authority="Wichard & Espeland, 2022" authorityName="Wichard & Espeland" authorityYear="2022" box="[227,500,975,1001]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="myanmari" status="sp. nov.">
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<emphasis id="A087EA81FF83156ACBCFFC7104D2E6D2" box="[227,500,975,1001]" italics="true" pageId="6" pageNumber="7">Cretapsyche myanmari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBB457FAFF83156ACAD1FC71077FE6D3" box="[509,601,975,1001]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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can be distinguished from all other species in the genus by having M branching before Rs in the forewing and an open thyridial cell.
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<taxonomicName id="55F34D10FF83156AC8EAFC4A01F3E134" authority="Wichard & Espeland, 2022" authorityName="Wichard & Espeland" authorityYear="2022" box="[966,1237,1012,1038]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="myanmari" status="sp. nov.">
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<emphasis id="A087EA81FF83156AC8EAFC4A01F3E134" box="[966,1237,1012,1038]" italics="true" pageId="6" pageNumber="7">Cretapsyche myanmari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBB457FAFF83156ACFF0FC4B0010E135" box="[1244,1334,1013,1039]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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can also be distinguished in the male genitalia by the mesodorsal processes of tergum X, scale-shaped, each with two black spines.
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</paragraph>
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</subSubSection>
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<subSubSection id="DAE96518FF83156ACBCFFB3A04A7E1CF" pageId="6" pageNumber="7" type="etymology">
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<paragraph id="924C3693FF83156ACBCFFB3A044CE1A4" blockId="6.[227,362,1156,1182]" box="[227,362,1156,1182]" pageId="6" pageNumber="7">
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<heading id="C90481FFFF83156ACBCFFB3A044CE1A4" bold="true" box="[227,362,1156,1182]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
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<emphasis id="A087EA81FF83156ACBCFFB3A044CE1A4" bold="true" box="[227,362,1156,1182]" pageId="6" pageNumber="7">Etymology</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="924C3693FF83156ACBCFFB0B04A7E1CF" blockId="6.[227,1436,1205,1269]" pageId="6" pageNumber="7">
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<taxonomicName id="55F34D10FF83156ACBCFFB0B04D1E1F5" authority="Wichard & Espeland, 2022" authorityName="Wichard & Espeland" authorityYear="2022" box="[227,503,1205,1231]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="myanmari" status="sp. nov.">
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<emphasis id="A087EA81FF83156ACBCFFB0B04D1E1F5" box="[227,503,1205,1231]" italics="true" pageId="6" pageNumber="7">Cretapsyche myanmari</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BBB457FAFF83156AC92FFB080744E1EA" box="[515,610,1206,1232]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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is named after
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<collectingCountry id="EAE47603FF83156AC804FB0806BAE1EA" box="[808,924,1206,1232]" name="Myanmar" pageId="6" pageNumber="7">Myanmar</collectingCountry>
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, where the amber inclusion was found in
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<collectingRegion id="5037F871FF83156ACBCFFB65045BE1CF" box="[227,381,1243,1269]" country="Myanmar" name="Kachin" pageId="6" pageNumber="7">Kachin State</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="DAE96518FF83156ACBCFFAA1061CE37F" pageId="6" pageNumber="7" type="materials_examined">
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<paragraph id="924C3693FF83156ACBCFFAA104F6E003" blockId="6.[227,464,1311,1337]" box="[227,464,1311,1337]" pageId="6" pageNumber="7">
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<heading id="C90481FFFF83156ACBCFFAA104F6E003" bold="true" box="[227,464,1311,1337]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
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<emphasis id="A087EA81FF83156ACBCFFAA104F6E003" bold="true" box="[227,464,1311,1337]" pageId="6" pageNumber="7">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="924C3693FF83156ACA3CFAEE04EBE0E1" blockId="6.[227,1437,1360,1499]" pageId="6" pageNumber="7">
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<materialsCitation id="229B3CCEFF83156ACA3CFAEE04EBE0E1" ID-GBIF-Occurrence="3866793302" collectionCode="ZFMK" country="Myanmar" location="Kachin State" pageId="6" pageNumber="7" specimenCode="ZFMK-TRI000830" specimenCount="1" specimenCount-male="1" stateProvince="Kachin State" typeStatus="holotype">
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<emphasis id="A087EA81FF83156ACA3CFAEE04A4E050" bold="true" box="[272,386,1360,1386]" pageId="6" pageNumber="7">
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<typeStatus id="4D488831FF83156ACA3CFAEE04A4E050" box="[272,386,1360,1386]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectingCountry id="EAE47603FF83156ACBCFFAC8045AE0AA" box="[227,380,1398,1424]" name="Myanmar" pageId="6" pageNumber="7">MYANMAR</collectingCountry>
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–
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<emphasis id="A087EA81FF83156ACAB0FACB0776E0AA" bold="true" box="[412,592,1397,1424]" pageId="6" pageNumber="7">
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<collectingRegion id="5037F871FF83156ACAB0FACB0719E0B5" box="[412,575,1397,1423]" country="Myanmar" name="Kachin" pageId="6" pageNumber="7">Kachin State</collectingRegion>
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•
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</emphasis>
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<specimenCount id="84F5FD1AFF83156AC974FAC80757E0AA" box="[600,625,1398,1424]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
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; exact locality unknown;
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<geologicalTimeScale id="E0D3B638FF83156AC88EFAC800BEE0AA" box="[930,1432,1398,1424]" pageId="6" pageNumber="7">Mid-Cretaceous Burmese amber inclusion</geologicalTimeScale>
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; deposited in the amber collection of the
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<collectionCode id="F4E2AE56FF83156AC9D8FA22066FE08C" box="[756,841,1436,1462]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="7" type="Museum">ZFMK</collectionCode>
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; former Patrick Müller Collection, BUB 3597;
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<specimenCode id="C2559EE8FF83156ACBCFFA7F04EFE0E1" box="[227,457,1473,1499]" collectionCode="ZFMK-TRI" pageId="6" pageNumber="7">ZFMK-TRI000830</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="924C3693FF83156ACBCFF9BB061CE37F" blockId="6.[227,1436,1541,1606]" pageId="6" pageNumber="7">Body well preserved; forewing in dorsal view, hindwing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view.</paragraph>
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</subSubSection>
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<subSubSection id="DAE96518FF83156ACBCFF9D1004BED34" pageId="6" pageNumber="7" type="description">
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<paragraph id="924C3693FF83156ACBCFF9D10452E3B3" blockId="6.[227,372,1647,1673]" box="[227,372,1647,1673]" pageId="6" pageNumber="7">
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<heading id="C90481FFFF83156ACBCFF9D10452E3B3" bold="true" box="[227,372,1647,1673]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
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||
<emphasis id="A087EA81FF83156ACBCFF9D10452E3B3" bold="true" box="[227,372,1647,1673]" pageId="6" pageNumber="7">Description</emphasis>
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||
</heading>
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</paragraph>
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<paragraph id="924C3693FF83156ACA3CF91E0476E380" blockId="6.[226,1438,1696,2063]" box="[272,336,1696,1722]" pageId="6" pageNumber="7">
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||
<emphasis id="A087EA81FF83156ACA3CF91E0476E380" bold="true" box="[272,336,1696,1722]" pageId="6" pageNumber="7">Male</emphasis>
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</paragraph>
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<paragraph id="924C3693FF83156ACBCFF978004BED34" blockId="6.[226,1438,1696,2063]" pageId="6" pageNumber="7">
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Ocelli absent. Antennae little longer than forewings. Scapus about as long as head, smaller pedicellus, followed by more than 30 cylindrical flagellomeres gradually shortening in length apically. Maxillary palps 5-segmented, 2
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<superScript id="65869BDBFF83156ACACFF8AF04D2E225" attach="left" box="[483,500,1809,1823]" fontSize="6" pageId="6" pageNumber="7">nd</superScript>
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segment longest, following three segments successively smaller and shorter. Body and wings light brown. Forewing with length about
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<quantity id="550B9B76FF83156AC8A1F88906CCE268" box="[909,1002,1847,1874]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="7" unit="mm" value="2.8">2.8 mm</quantity>
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. In forewings, apical forks II, III and V present, discoidal and thyridial cells absent, Cu2 running continuously to Cu1b. In hindwings, apical forks I, III and V present (
|
||
<typeStatus id="4D488831FF83156AC93DF83D075CE2A7" box="[529,634,1923,1949]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
hindwings partially spread laterally and visible). Tibial spur formula 2/2/4. In male genitalia, two-segmented inferior appendages characterized by finger-shaped, curved coxopodite and by short and cone-shaped harpago. Two appendages, each with two dark spines, located centrally in genitalia interpreted as probable mesodorsal pair of lobes of tergum X (
|
||
<bibRefCitation id="F6624B62FF83156ACF9FF84A0046ED34" author="Wichard W." box="[1203,1376,2036,2062]" pageId="6" pageNumber="7" pagination="104707" refId="ref3509" refString="Wichard W. 2021. Overview of the caddisflies (Insecta, Trichoptera) in mid-Cretaceous Burmese amber. Cretaceous Research 119: e 104707. https: // doi. org / 10.1016 / j. cretres. 2020.104707" type="journal article" year="2021">Wichard 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |