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<document id="3176AE31ECCD3A576F28106F4C1C0D4F" ID-DOI="10.5281/zenodo.180435" ID-GBIF-Dataset="91705fcc-498c-41e0-a210-445a05bdfdab" ID-ISSN="1175-5326" ID-Zenodo-Dep="180435" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459807571021" checkinUser="plazi" docAuthor="Maruyama, Munetoshi, Klimaszewski, Jan &amp; Gusarov, Vladimir" docDate="2008" docId="191C6D0BFFC7FFCBFF7F3E24F428F9AA" docLanguage="en" docName="zt01683p050.pdf" docOrigin="Zootaxa 1683" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Osakatheta Maruyama, Klimaszewski &amp; Gusarov" docType="treatment" docVersion="9" lastPageNumber="41" masterDocId="E5251573FFC6FFC9FFE83D0BF630FF80" masterDocTitle="Osakatheta yasukoae, a new intertidal genus and species of athetine rove beetles (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="50" masterPageNumber="39" pageNumber="40" updateTime="1698220674226" updateUser="plazi">
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<mods:title id="15CC3EB03302EA5CD0343B08BFAE44C1">Osakatheta yasukoae, a new intertidal genus and species of athetine rove beetles (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="03D9D02EA22C5599A9C02657DAE797E9">Maruyama, Munetoshi</mods:namePart>
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<treatment id="191C6D0BFFC7FFCBFF7F3E24F428F9AA" ID-DOI="http://doi.org/10.5281/zenodo.6233900" ID-GBIF-Taxon="119353205" ID-Zenodo-Dep="6233900" LSID="urn:lsid:plazi:treatment:191C6D0BFFC7FFCBFF7F3E24F428F9AA" httpUri="http://treatment.plazi.org/id/191C6D0BFFC7FFCBFF7F3E24F428F9AA" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
<subSubSection id="D9AF8F96FFC7FFC8FF7F3E24F716FCEC" pageId="1" pageNumber="40" type="nomenclature">
<paragraph id="910ADC1DFFC7FFC8FF7F3E24F54BFCC9" blockId="1.[151,891,815,876]" box="[151,891,815,841]" pageId="1" pageNumber="40">
<heading id="CA426B71FFC7FFC8FF7F3E24F54BFCC9" bold="true" box="[151,891,815,841]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
<emphasis id="A3C1000FFFC7FFC8FF7F3E24F54BFCC9" bold="true" box="[151,891,815,841]" pageId="1" pageNumber="40">
<taxonomicName id="56B5A79EFFC7FFC8FF7F3E24F4CFFCC9" ID-CoL="8W4VY" authority="Maruyama, Klimaszewski &amp; Gusarov" authorityName="Maruyama, Klimaszewski &amp; Gusarov" box="[151,767,815,841]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="A3C1000FFFC7FFC8FF7F3E24F72FFCC9" bold="true" box="[151,287,815,841]" italics="true" pageId="1" pageNumber="40">Osakatheta</emphasis>
Maruyama, Klimaszewski &amp; Gusarov
</taxonomicName>
, gen. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="910ADC1DFFC7FFC8FF7F3E59F716FCEC" blockId="1.[151,891,815,876]" box="[151,294,850,876]" pageId="1" pageNumber="40">
(
<figureCitation id="098EC098FFC7FFC8FF773E59F72DFCEC" box="[159,285,850,876]" captionStart-0="FIGURES 1 5" captionStart-1="FIGURES 6 10" captionStart-2="FIGURES 11 18" captionStart-3="FIGURES 19 23" captionStart-4="FIGURES 24 27" captionStart-5="FIGURES 28 35" captionStart-6="FIGURE 36" captionStart-7="FIGURE 37" captionStartId-0="3.[151,269,1928,1952]" captionStartId-1="4.[151,269,1666,1690]" captionStartId-2="5.[151,269,1938,1962]" captionStartId-3="6.[151,269,1122,1146]" captionStartId-4="6.[151,269,1975,1999]" captionStartId-5="7.[151,269,1875,1899]" captionStartId-6="8.[151,255,1072,1096]" captionStartId-7="9.[151,255,1564,1588]" captionTargetBox-0="[239,1355,338,1901]" captionTargetBox-1="[155,1429,199,1636]" captionTargetBox-2="[189,1422,588,1913]" captionTargetBox-3="[226,1360,182,1098]" captionTargetBox-4="[241,1364,1224,1957]" captionTargetBox-5="[155,1424,194,1846]" captionTargetBox-6="[151,1435,194,1047]" captionTargetBox-7="[151,1435,394,1540]" captionTargetId-0="figure@3.[239,1355,336,1902]" captionTargetId-1="figure@4.[151,1436,194,1642]" captionTargetId-2="figure@5.[164,1429,578,1916]" captionTargetId-3="figure@6.[226,1360,182,1098]" captionTargetId-4="figure@6.[238,1366,1220,1966]" captionTargetId-5="figure@7.[151,1436,194,1851]" captionTargetId-6="figure@8.[151,1436,194,1048]" captionTargetId-7="figure@9.[151,1436,394,1540]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionText-0="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." captionText-1="FIGURES 6 10. Osakatheta yasukoae, mouthparts. 6 — labrum, dorsal view; 7 — epipharynx, adoral view; 8 — left mandible, dorsal view; 9 — left mandible, ventral view; 10 — right mandible, ventral view." captionText-2="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." captionText-3="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." captionText-4="FIGURES 24 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." captionText-5="FIGURES 28 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." captionText-6="FIGURE 36. The type locality of Osakatheta yasukoae, a tidal flat at Nankô-yachôen, Ôsaka-fu, Japan. The beetles were found under the stones in the bottom of this photograph. Photographed by Rikio Matsumoto." captionText-7="FIGURE 37. Known localities of Osakatheta yasukoae (black stars)." httpUri-0="https://zenodo.org/record/180436/files/figure.png" httpUri-1="https://zenodo.org/record/180437/files/figure.png" httpUri-2="https://zenodo.org/record/180438/files/figure.png" httpUri-3="https://zenodo.org/record/180439/files/figure.png" httpUri-4="https://zenodo.org/record/180440/files/figure.png" httpUri-5="https://zenodo.org/record/180441/files/figure.png" httpUri-6="https://zenodo.org/record/180442/files/figure.png" httpUri-7="https://zenodo.org/record/180443/files/figure.png" pageId="1" pageNumber="40">Figs. 137</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D9AF8F96FFC7FFC8FF7F3EA9F254FC3C" box="[151,1124,930,956]" pageId="1" pageNumber="40" type="materials_examined">
<paragraph id="910ADC1DFFC7FFC8FF7F3EA9F254FC3C" blockId="1.[151,1437,930,2036]" box="[151,1124,930,956]" pageId="1" pageNumber="40">
<emphasis id="A3C1000FFFC7FFC8FF7F3EA9F709FC3C" bold="true" box="[151,313,930,956]" pageId="1" pageNumber="40">
<typeStatus id="4E0E62BFFFC7FFC8FF7F3EA9F6E4FC3C" box="[151,212,930,956]" pageId="1" pageNumber="40">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="56B5A79EFFC7FFC8FEA83EA9F5CEFC3C" authority="Maruyama, Klimaszewski &amp; Gusarov" authorityName="Maruyama, Klimaszewski &amp; Gusarov" box="[320,1022,930,956]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="yasukoae" status="sp. nov.">
<emphasis id="A3C1000FFFC7FFC8FEA83EA9F40AFC3B" box="[320,570,930,955]" italics="true" pageId="1" pageNumber="40">Osakatheta yasukoae</emphasis>
Maruyama, Klimaszewski &amp; Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="B8F2BD74FFC7FFC8FBE23EA9F254FC3C" box="[1034,1124,930,956]" pageId="1" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D9AF8F96FFC7FFC8FF2E3EC1F762FB8C" pageId="1" pageNumber="40" type="diagnosis">
<paragraph id="910ADC1DFFC7FFC8FF2E3EC1F762FB8C" blockId="1.[151,1437,930,2036]" pageId="1" pageNumber="40">
<emphasis id="A3C1000FFFC7FFC8FF2E3EC1F776FC64" bold="true" box="[198,326,970,996]" pageId="1" pageNumber="40">Diagnosis.</emphasis>
<taxonomicName id="56B5A79EFFC7FFC8FEA53EC1F7E2FC63" box="[333,466,970,995]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="A3C1000FFFC7FFC8FEA53EC1F7E2FC63" box="[333,466,970,995]" italics="true" pageId="1" pageNumber="40">Osakatheta</emphasis>
</taxonomicName>
can be readily distinguished from most athetine genera by the erect setae covering the whole body.
</paragraph>
</subSubSection>
<subSubSection id="D9AF8F96FFC7FFCBFF2E3911F73EFA0A" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40" type="description">
<paragraph id="910ADC1DFFC7FFC8FF2E3911F7E0FBDC" blockId="1.[151,1437,930,2036]" pageId="1" pageNumber="40">
<emphasis id="A3C1000FFFC7FFC8FF2E3911F76FFBB4" bold="true" box="[198,351,1050,1076]" pageId="1" pageNumber="40">Description.</emphasis>
Body (
<figureCitation id="098EC098FFC7FFC8FE503911F418FBB4" box="[440,552,1050,1076]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="1" pageNumber="40">Figs. 12</figureCitation>
) slender, broadened posteriorly; dorsal surface almost completely covered with dense and erect setae.
</paragraph>
<paragraph id="910ADC1DFFC7FFC8FF2D3960F550FB54" blockId="1.[151,1437,930,2036]" pageId="1" pageNumber="40">
<emphasis id="A3C1000FFFC7FFC8FF2D3960F73DFB04" box="[197,269,1131,1156]" italics="true" pageId="1" pageNumber="40">Head.</emphasis>
Head capsule (
<figureCitation id="098EC098FFC7FFC8FE2D3961F406FB04" box="[453,566,1130,1156]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="1" pageNumber="40">Figs. 35</figureCitation>
) approximately circular, gently convex dorsally. Eyes (
<figureCitation id="098EC098FFC7FFC8FB203961F308FB04" box="[1224,1336,1130,1156]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="1" pageNumber="40">Figs. 35</figureCitation>
) oval in lateral view, somewhat prominent. Infraorbital carina absent (
<figureCitation id="098EC098FFC7FFC8FC6A3999F5FDFB2C" box="[898,973,1170,1196]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="1" pageNumber="40">Fig. 5</figureCitation>
). Antennae (
<figureCitation id="098EC098FFC7FFC8FB833999F2EDFB2C" box="[1131,1245,1170,1196]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="1" pageNumber="40">Figs. 15</figureCitation>
) 11-segmented, moniliform, almost as long as head and pronotum combined.
</paragraph>
<paragraph id="910ADC1DFFC7FFCBFF2D39E8F32CFDC2" blockId="1.[151,1437,930,2036]" lastBlockId="2.[151,1437,152,1578]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
<emphasis id="A3C1000FFFC7FFC8FF2D39E8F766FB7C" box="[197,342,1251,1276]" italics="true" pageId="1" pageNumber="40">Mouthparts.</emphasis>
Labrum (
<figureCitation id="098EC098FFC7FFC8FE2239E9F43FFB7C" box="[458,527,1250,1276]" captionStart="FIGURES 6 10" captionStartId="4.[151,269,1666,1690]" captionTargetBox="[155,1429,199,1636]" captionTargetId="figure@4.[151,1436,194,1642]" captionTargetPageId="4" captionText="FIGURES 6 10. Osakatheta yasukoae, mouthparts. 6 — labrum, dorsal view; 7 — epipharynx, adoral view; 8 — left mandible, dorsal view; 9 — left mandible, ventral view; 10 — right mandible, ventral view." httpUri="https://zenodo.org/record/180437/files/figure.png" pageId="1" pageNumber="40">Fig. 6</figureCitation>
) strongly transverse (width/length ratio 2.1), rounded anteriorly; surface covered with some 4550 pseudopores, except for posterior and lateral areas; apodeme roundly produced posteromedially, with lateral arm gently curved and its apex hidden under postero-medial projection of apodeme. Epipharynx as in
<figureCitation id="098EC098FFC7FFC8FE8C3851F79BFAF4" box="[356,427,1370,1396]" captionStart="FIGURES 6 10" captionStartId="4.[151,269,1666,1690]" captionTargetBox="[155,1429,199,1636]" captionTargetId="figure@4.[151,1436,194,1642]" captionTargetPageId="4" captionText="FIGURES 6 10. Osakatheta yasukoae, mouthparts. 6 — labrum, dorsal view; 7 — epipharynx, adoral view; 8 — left mandible, dorsal view; 9 — left mandible, ventral view; 10 — right mandible, ventral view." httpUri="https://zenodo.org/record/180437/files/figure.png" pageId="1" pageNumber="40">Fig. 7</figureCitation>
. with long sensilla
<emphasis id="A3C1000FFFC7FFC8FD643850F4ABFAF4" box="[652,667,1371,1396]" italics="true" pageId="1" pageNumber="40">a</emphasis>
, its medial sensory field with 11 pores and numerous minute sensory pseudopores; lateral row with 35 pores distributed between membranous wrinkles; 3 marginal setae slender, about 10 times as long as wide. Mandibles (
<figureCitation id="098EC098FFC7FFC8FCF938A1F5A0FA44" box="[785,912,1450,1476]" captionStart="FIGURES 6 10" captionStartId="4.[151,269,1666,1690]" captionTargetBox="[155,1429,199,1636]" captionTargetId="figure@4.[151,1436,194,1642]" captionTargetPageId="4" captionText="FIGURES 6 10. Osakatheta yasukoae, mouthparts. 6 — labrum, dorsal view; 7 — epipharynx, adoral view; 8 — left mandible, dorsal view; 9 — left mandible, ventral view; 10 — right mandible, ventral view." httpUri="https://zenodo.org/record/180437/files/figure.png" pageId="1" pageNumber="40">Figs. 810</figureCitation>
) slightly asymmetric, mesal areas of dorsal surface covered with 3040 pseudopores; prostheca obtuse at apex, its inner margin densely spinulate; left mandible (
<figureCitation id="098EC098FFC7FFC8FEFB38F1F7B3F994" box="[275,387,1530,1556]" captionStart="FIGURES 6 10" captionStartId="4.[151,269,1666,1690]" captionTargetBox="[155,1429,199,1636]" captionTargetId="figure@4.[151,1436,194,1642]" captionTargetPageId="4" captionText="FIGURES 6 10. Osakatheta yasukoae, mouthparts. 6 — labrum, dorsal view; 7 — epipharynx, adoral view; 8 — left mandible, dorsal view; 9 — left mandible, ventral view; 10 — right mandible, ventral view." httpUri="https://zenodo.org/record/180437/files/figure.png" pageId="1" pageNumber="40">Figs. 89</figureCitation>
) generalised, without teeth; 3 small setae present laterally; right mandible (
<figureCitation id="098EC098FFC7FFC8FB1838F1F374F994" box="[1264,1348,1530,1556]" captionStart="FIGURES 6 10" captionStartId="4.[151,269,1666,1690]" captionTargetBox="[155,1429,199,1636]" captionTargetId="figure@4.[151,1436,194,1642]" captionTargetPageId="4" captionText="FIGURES 6 10. Osakatheta yasukoae, mouthparts. 6 — labrum, dorsal view; 7 — epipharynx, adoral view; 8 — left mandible, dorsal view; 9 — left mandible, ventral view; 10 — right mandible, ventral view." httpUri="https://zenodo.org/record/180437/files/figure.png" pageId="1" pageNumber="40">Fig. 10</figureCitation>
) with a small tooth at middle of inner margin, weakly serrate between tooth and apex; 3 small setae present laterally. Labium as in
<figureCitation id="098EC098FFC7FFC8FEA13B41F7ECF9E4" box="[329,476,1610,1636]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Figs. 1112</figureCitation>
. Prementum (
<figureCitation id="098EC098FFC7FFC8FD623B41F4D3F9E4" box="[650,739,1610,1636]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 11</figureCitation>
) with 2 real pores and 1 setal pore latero-mesally, 35 pseudopores between real pores, and 2530 medial pseudopores; internal ridge (
<figureCitation id="098EC098FFC7FFC8FBD73B79F2A4F90C" box="[1087,1172,1650,1676]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 11</figureCitation>
; indicated with arrow) very small, its length less than 1/6 that of prementum; apodeme (
<figureCitation id="098EC098FFC7FFC8FC633B91F5EEF934" box="[907,990,1690,1716]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 11</figureCitation>
) with broom-like projections posteromedially, lateral lobe of apodeme (
<figureCitation id="098EC098FFC7FFC8FDC63BC9F4B2F95C" box="[558,642,1730,1756]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 11</figureCitation>
) gently curved, narrowed apically, pointed at apex. Ligula (
<figureCitation id="098EC098FFC7FFC8FAD73BC9F3A2F95C" box="[1343,1426,1730,1756]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 11</figureCitation>
) bilobed, with each lobe rounded and covered with minute pores apically; apical setae absent; lateral margin weakly sclerotized. Labial palpus (
<figureCitation id="098EC098FFC7FFC8FDDC3A19F4A3F8AC" box="[564,659,1810,1836]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Figs. 11</figureCitation>
) with segment I long and twice as long as II; segment II notched from base to apex forming membranous zone, which bears one pore; segment III longer than II, slightly dilated apically, with 1 pore near apex, covered with spinulae at apex; setae α, β, and γ present. Hypopharynx (
<figureCitation id="098EC098FFC7FFC8FF773A81F6C7F824" box="[159,247,1930,1956]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 12</figureCitation>
) generalised, without apical seta; medial sensory field simplified, with small ridges. Maxilla as in
<figureCitation id="098EC098FFC7FFC8FF7F3AB9F716F84C" box="[151,294,1970,1996]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Figs. 1317</figureCitation>
. Cardo (
<figureCitation id="098EC098FFC7FFC8FE623AB9F7EFF84C" box="[394,479,1970,1996]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 13</figureCitation>
) generalised, almost elliptical in ventral view, with about 10 pseudopores. Stipes (
<figureCitation id="098EC098FFC7FFC8FF773AD1F6C6F874" box="[159,246,2010,2036]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 13</figureCitation>
) small, triangular, with 3 setae around base, and 5 or 6 pseudopores. Palpifer (
<figureCitation id="098EC098FFC7FFC8FB4A3AD1F2C9F874" box="[1186,1273,2010,2036]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="1" pageNumber="40">Fig. 13</figureCitation>
) triangular in lateral view, with one large seta near apex, two small setae at middle and apex, and with some 5 pseudopores. Lacinia (
<figureCitation id="098EC098FFC4FFCBFEE83DCBF7BEFF5A" box="[256,398,192,218]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="2" pageNumber="41">Figs. 1315</figureCitation>
) with some 68 pores around base and 2 setae laterally; apical inner margin with a row of 8 spines forming comb; surface around comb densely pubescent, and with a row of pores. Subgalea (
<figureCitation id="098EC098FFC4FFCBFA853DE3F685FEAA" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="2" pageNumber="41">Fig. 13</figureCitation>
) weakly sclerotized. Galea (
<figureCitation id="098EC098FFC4FFCBFDE93C1BF488FEAA" box="[513,696,272,298]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="2" pageNumber="41">Figs. 13, 1617</figureCitation>
) broad, about 2.5 times as long as wide, slightly curved at middle, gently narrowed apically, with about 10 pores around middle of outer margin, densely pubescent and with pampas grass-like projection apically. Maxillary palpus as in
<figureCitation id="098EC098FFC4FFCBFC953C6BF5E6FEFA" box="[893,982,352,378]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="2" pageNumber="41">Fig. 13</figureCitation>
; segment I very small, without setae; segment II gently curved, and much dilated apically, with some pores along apical margin; segment III largest, 1.5 times longer than 2nd, almost straight, widest at middle, moderately covered with pores except for basal and apical margins; 4th long, narrowed apically, with one pore around middle. Mentum (
<figureCitation id="098EC098FFC4FFCBFB4E3CD3F2CBFE72" box="[1190,1275,472,498]" captionStart="FIGURES 11 18" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[189,1422,588,1913]" captionTargetId="figure@5.[164,1429,578,1916]" captionTargetPageId="5" captionText="FIGURES 11 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="2" pageNumber="41">Fig. 18</figureCitation>
) trapeziform; anterior margin with a pair of projections, which bear seta
<emphasis id="A3C1000FFFC4FFCBFCB53F0BF55CFD99" box="[861,876,512,537]" italics="true" pageId="2" pageNumber="41">u</emphasis>
; posterior margin completely truncate; surface moderately covered with pseudopores except for antero-lateral area; seta
<emphasis id="A3C1000FFFC4FFCBFC193F23F234FDC1" box="[1009,1028,552,577]" italics="true" pageId="2" pageNumber="41">w</emphasis>
located antero-mesally.
</paragraph>
<paragraph id="910ADC1DFFC4FFCBFF2D3F5BF6C2FCDA" blockId="2.[151,1437,152,1578]" pageId="2" pageNumber="41">
<emphasis id="A3C1000FFFC4FFCBFF2D3F5BF729FDE9" box="[197,281,592,617]" italics="true" pageId="2" pageNumber="41">Thorax</emphasis>
. Pronotum (
<figureCitation id="098EC098FFC4FFCBFE443F5BF7CDFDEA" box="[428,509,592,618]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="2" pageNumber="41">Figs. 1</figureCitation>
,
<figureCitation id="098EC098FFC4FFCBFDE33F5BF419FDEA" box="[523,553,592,618]" captionStart="FIGURES 19 23" captionStartId="6.[151,269,1122,1146]" captionTargetBox="[226,1360,182,1098]" captionTargetId="figure@6.[226,1360,182,1098]" captionTargetPageId="6" captionText="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="2" pageNumber="41">19</figureCitation>
) slightly transverse, angles widely rounded; microsetae directed anteriad in midline; in lateral portions of disk microsetae directed laterad (
<typeStatus id="4E0E62BFFFC4FFCBFC7E3F73F5E2FD12" box="[918,978,632,658]" pageId="2" pageNumber="41">Type</typeStatus>
I,
<bibRefCitation id="F524A1ECFFC4FFCBFC103F73F339FD12" author="Benick" box="[1016,1289,632,658]" pageId="2" pageNumber="49" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
). Hypomera (
<figureCitation id="098EC098FFC4FFCBFF773FABF6C4FD3A" box="[159,244,672,698]" captionStart="FIGURES 19 23" captionStartId="6.[151,269,1122,1146]" captionTargetBox="[226,1360,182,1098]" captionTargetId="figure@6.[226,1360,182,1098]" captionTargetPageId="6" captionText="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="2" pageNumber="41">Fig. 19</figureCitation>
) visible laterally. Prosternum (
<figureCitation id="098EC098FFC4FFCBFDB33FABF481FD3A" box="[603,689,672,698]" captionStart="FIGURES 19 23" captionStartId="6.[151,269,1122,1146]" captionTargetBox="[226,1360,182,1098]" captionTargetId="figure@6.[226,1360,182,1098]" captionTargetPageId="6" captionText="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="2" pageNumber="41">Fig. 20</figureCitation>
) transversely sutured; densely covered with setae along anterior margin; posterior margin truncate. Scutellum generalised, its apex acutely pointed (
<figureCitation id="098EC098FFC4FFCBFB833FC3F28DFD62" box="[1131,1213,712,738]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="2" pageNumber="41">Figs. 1</figureCitation>
,
<figureCitation id="098EC098FFC4FFCBFB233FC3F2DCFD62" box="[1227,1260,712,738]" captionStart="FIGURES 19 23" captionStartId="6.[151,269,1122,1146]" captionTargetBox="[226,1360,182,1098]" captionTargetId="figure@6.[226,1360,182,1098]" captionTargetPageId="6" captionText="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="2" pageNumber="41">22</figureCitation>
). Mesoventrite (
<figureCitation id="098EC098FFC4FFCBFF773FFBF6C7FC8A" box="[159,247,752,778]" captionStart="FIGURES 19 23" captionStartId="6.[151,269,1122,1146]" captionTargetBox="[226,1360,182,1098]" captionTargetId="figure@6.[226,1360,182,1098]" captionTargetPageId="6" captionText="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="2" pageNumber="41">Fig. 21</figureCitation>
) with mesoventral process moderate in length, 0.6 times as long as coxal cavities. Mesocoxae are moderately separated. Mesocoxal cavity not margined (
<figureCitation id="098EC098FFC4FFCBFCDD3E13F5BFFCB2" box="[821,911,792,818]" captionStart="FIGURES 19 23" captionStartId="6.[151,269,1122,1146]" captionTargetBox="[226,1360,182,1098]" captionTargetId="figure@6.[226,1360,182,1098]" captionTargetPageId="6" captionText="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="2" pageNumber="41">Fig. 21</figureCitation>
).
<collectingCountry id="E9A29C8DFFC4FFCBFC403E13F5D3FCB2" box="[936,995,792,818]" name="India" pageId="2" pageNumber="41">Hind</collectingCountry>
wing reduced, and about as long as elytron.
</paragraph>
<paragraph id="910ADC1DFFC4FFCBFF2D3E63F478FC2A" blockId="2.[151,1437,152,1578]" pageId="2" pageNumber="41">
<emphasis id="A3C1000FFFC4FFCBFF2D3E63F6CCFC01" box="[197,252,872,897]" italics="true" pageId="2" pageNumber="41">Legs</emphasis>
. Tarsal formula 4-5-5. Medial macroseta of mesotibia thin, slightly longer than tibial width. Empodium (
<figureCitation id="098EC098FFC4FFCBFF0B3E9BF708FC2A" box="[227,312,912,938]" captionStart="FIGURES 19 23" captionStartId="6.[151,269,1122,1146]" captionTargetBox="[226,1360,182,1098]" captionTargetId="figure@6.[226,1360,182,1098]" captionTargetPageId="6" captionText="FIGURES 19 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="2" pageNumber="41">Fig. 23</figureCitation>
) simple, without setae.
</paragraph>
<paragraph id="910ADC1DFFC4FFCBFF2D3EB3F417FBCA" blockId="2.[151,1437,152,1578]" pageId="2" pageNumber="41">
<emphasis id="A3C1000FFFC4FFCBFF2D3EB3F70CFC51" box="[197,316,952,977]" italics="true" pageId="2" pageNumber="41">Abdomen.</emphasis>
Segments IIIVII (
<figureCitation id="098EC098FFC4FFCBFDCC3EB3F459FC52" box="[548,617,952,978]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="2" pageNumber="41">Fig. 1</figureCitation>
) with posterior margin almost straight; surface densely covered with erect setae. Tergites IIIV with moderate basal impressions. Abdominal tergite VIII (
<figureCitation id="098EC098FFC4FFCBFB9B3EEBF2F8FC7A" box="[1139,1224,992,1018]" captionStart="FIGURES 24 27" captionStartId="6.[151,269,1975,1999]" captionTargetBox="[241,1364,1224,1957]" captionTargetId="figure@6.[238,1366,1220,1966]" captionTargetPageId="6" captionText="FIGURES 24 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." httpUri="https://zenodo.org/record/180440/files/figure.png" pageId="2" pageNumber="41">Fig. 24</figureCitation>
) with basal suture curved laterally and continued apicad; sternite (
<figureCitation id="098EC098FFC4FFCBFD273903F515FBA2" box="[719,805,1032,1058]" captionStart="FIGURES 24 27" captionStartId="6.[151,269,1975,1999]" captionTargetBox="[241,1364,1224,1957]" captionTargetId="figure@6.[238,1366,1220,1966]" captionTargetPageId="6" captionText="FIGURES 24 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." httpUri="https://zenodo.org/record/180440/files/figure.png" pageId="2" pageNumber="41">Fig. 25</figureCitation>
) with basal suture reaching lateral sides. Tergites IX and X densely covered with setae.
</paragraph>
<paragraph id="910ADC1DFFC4FFCBFF2D3953F7B1FA92" blockId="2.[151,1437,152,1578]" pageId="2" pageNumber="41">
<emphasis id="A3C1000FFFC4FFCBFF2D3953F7B8FBF1" box="[197,392,1112,1137]" italics="true" pageId="2" pageNumber="41">Male characters</emphasis>
. Sternite VIII (
<figureCitation id="098EC098FFC4FFCBFDAA3953F4E1FBF2" box="[578,721,1112,1138]" captionStart="FIGURES 24 27" captionStartId="6.[151,269,1975,1999]" captionTargetBox="[241,1364,1224,1957]" captionTargetId="figure@6.[238,1366,1220,1966]" captionTargetPageId="6" captionText="FIGURES 24 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." httpUri="https://zenodo.org/record/180440/files/figure.png" pageId="2" pageNumber="41">Figs. 2526</figureCitation>
) with apical margin weakly sclerotized forming transparent zone, lacking minute pubescence and with apical sensory setae which are 1.5 times as long as in female. Aedeagus: median lobe (
<figureCitation id="098EC098FFC4FFCBFE5139A3F476FB42" box="[441,582,1192,1218]" captionStart="FIGURES 28 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="2" pageNumber="41">Figs. 2829</figureCitation>
) with basal capsule bulbous; distal crests fused on their sides, projecting; copulatory piece (
<figureCitation id="098EC098FFC4FFCBFE8639DBF430FB6A" box="[366,512,1232,1258]" captionStart="FIGURES 28 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="2" pageNumber="41">Figs. 30, 31</figureCitation>
) bent in lateral view; paramere (
<figureCitation id="098EC098FFC4FFCBFC6539DBF22DFB6A" box="[909,1053,1232,1258]" captionStart="FIGURES 28 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="2" pageNumber="41">Figs. 3233</figureCitation>
) with condylite almost straight; hinge zone obscure.
</paragraph>
<paragraph id="910ADC1DFFC4FFCBFF2D382BF73EFA0A" blockId="2.[151,1437,152,1578]" pageId="2" pageNumber="41">
<emphasis id="A3C1000FFFC4FFCBFF2D382BF793FAB9" box="[197,419,1312,1337]" italics="true" pageId="2" pageNumber="41">Female characters</emphasis>
. Sternite VIII (
<figureCitation id="098EC098FFC4FFCBFDB5382BF484FABA" box="[605,692,1312,1338]" captionStart="FIGURES 24 27" captionStartId="6.[151,269,1975,1999]" captionTargetBox="[241,1364,1224,1957]" captionTargetId="figure@6.[238,1366,1220,1966]" captionTargetPageId="6" captionText="FIGURES 24 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." httpUri="https://zenodo.org/record/180440/files/figure.png" pageId="2" pageNumber="41">Fig. 27</figureCitation>
) with apical margin fully sclerotized, with minute pubescence and apical sensory setae shorter than in male. Spermatheca (
<figureCitation id="098EC098FFC4FFCBFCBA3843F5EFFAE2" box="[850,991,1352,1378]" captionStart="FIGURES 28 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="2" pageNumber="41">Figs. 3435</figureCitation>
) with capsule much shorter than elongate stem.
</paragraph>
</subSubSection>
<subSubSection id="D9AF8F96FFC4FFCBFF2D389CF53BFA5A" pageId="2" pageNumber="41" type="etymology">
<paragraph id="910ADC1DFFC4FFCBFF2D389CF53BFA5A" blockId="2.[151,1437,152,1578]" pageId="2" pageNumber="41">
<emphasis id="A3C1000FFFC4FFCBFF2D389CF762FA31" bold="true" box="[197,338,1431,1457]" pageId="2" pageNumber="41">Etymology.</emphasis>
<taxonomicName id="56B5A79EFFC4FFCBFEB33893F7D0FA31" box="[347,480,1432,1457]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="A3C1000FFFC4FFCBFEB33893F7D0FA31" box="[347,480,1432,1457]" italics="true" pageId="2" pageNumber="41">Osakatheta</emphasis>
</taxonomicName>
is a combination of “Ôsaka”, the
<typeStatus id="4E0E62BFFFC4FFCBFC9C3893F596FA32" box="[884,934,1432,1458]" pageId="2" pageNumber="41">type</typeStatus>
locality of the
<typeStatus id="4E0E62BFFFC4FFCBFBB43893F2BFFA32" box="[1116,1167,1432,1458]" pageId="2" pageNumber="41">type</typeStatus>
species, and “
<taxonomicName id="56B5A79EFFC4FFCBFAD33893F3B7FA31" box="[1339,1415,1432,1457]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="A3C1000FFFC4FFCBFAD33893F3B7FA31" box="[1339,1415,1432,1457]" italics="true" pageId="2" pageNumber="41">Atheta</emphasis>
</taxonomicName>
”, the
<typeStatus id="4E0E62BFFFC4FFCBFF2A38CBF6C5FA5A" box="[194,245,1472,1498]" pageId="2" pageNumber="41">type</typeStatus>
genus of the tribe Athetini. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="D9AF8F96FFC4FFCBFF2D38ECF428F9AA" pageId="2" pageNumber="41" type="distribution">
<paragraph id="910ADC1DFFC4FFCBFF2D38ECF428F9AA" blockId="2.[151,1437,152,1578]" pageId="2" pageNumber="41">
<emphasis id="A3C1000FFFC4FFCBFF2D38ECF756F981" bold="true" box="[197,358,1511,1537]" pageId="2" pageNumber="41">Distribution.</emphasis>
<collectingCountry id="E9A29C8DFFC4FFCBFE8638E3F785F982" box="[366,437,1512,1538]" name="Japan" pageId="2" pageNumber="41">Japan</collectingCountry>
: Honshû. So far known from Honshû,
<collectingCountry id="E9A29C8DFFC4FFCBFC6838E3F5F7F982" box="[896,967,1512,1538]" name="Japan" pageId="2" pageNumber="41">Japan</collectingCountry>
. It is probably more broadly distributed in the other regions of
<collectingCountry id="E9A29C8DFFC4FFCBFE493B1BF424F9AA" box="[417,532,1552,1578]" name="China" pageId="2" pageNumber="41">East Asia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>