272 lines
37 KiB
XML
272 lines
37 KiB
XML
<document id="3C70C02689487AF606809C2A7E85D8F7" ID-DOI="10.5281/zenodo.180435" ID-GBIF-Dataset="91705fcc-498c-41e0-a210-445a05bdfdab" ID-ISSN="1175-5326" ID-Zenodo-Dep="180435" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459807571021" checkinUser="plazi" docAuthor="Maruyama, Munetoshi, Klimaszewski, Jan & Gusarov, Vladimir" docDate="2008" docId="191C6D0BFFC4FFC1FF7F3B8CF2BAF93B" docLanguage="en" docName="zt01683p050.pdf" docOrigin="Zootaxa 1683" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Osakatheta yasukoae Maruyama, Klimaszewski & Gusarov, sp. nov." docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="E5251573FFC6FFC9FFE83D0BF630FF80" masterDocTitle="Osakatheta yasukoae, a new intertidal genus and species of athetine rove beetles (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="50" masterPageNumber="39" pageNumber="41" updateTime="1698220674226" updateUser="plazi">
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<mods:mods id="65FA8D1E784F0D2C91C3D12C1B4E13DA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="34EF7E5189C1DD61830FDE172574571F">
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<mods:title id="AA5D16D7609B77EEA996DD79C350AFC4">Osakatheta yasukoae, a new intertidal genus and species of athetine rove beetles (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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</mods:titleInfo>
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<mods:name id="AF84E04C0CDECDE06F0180A94752248B" type="personal">
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<mods:roleTerm id="A83208EF283F183D30A99064A813A358">Author</mods:roleTerm>
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<mods:namePart id="63E51B0225F28B586B30535CA60C7793">Maruyama, Munetoshi</mods:namePart>
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</mods:name>
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<mods:name id="4D5F5C7FDD58B84F7D40131B6F141356" type="personal">
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<mods:roleTerm id="EB14730033FA7774838782D9E0CB303C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="124A1C5BD7624DBF6728322B8E69F720">Klimaszewski, Jan</mods:namePart>
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</mods:name>
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<mods:name id="6C95844C455023733DFA1AD41C261B93" type="personal">
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<mods:roleTerm id="B14946EF3C4C7FC2F9997F59309DC33A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="95DAC0F62C7A835C9429910F81FBE372">Gusarov, Vladimir</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="31FF7AC3908A522C5E4389CC21FE358E">text</mods:typeOfResource>
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<mods:title id="CC28787EF61B0FE9FC6803C57691550B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="035F5E061120CFEDEDC14B57EEF4969F">
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<mods:date id="9E0F67AA828DCC99DA7B78165A1AEED0">2008</mods:date>
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<mods:detail id="FC653402145F57F9540156831DACCD03" type="volume">
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<mods:number id="A665DC87CD63AC459FA26E10C5BFF5A3">1683</mods:number>
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<mods:classification id="74BDA1E27F941512DA10782D2D6136F0">journal article</mods:classification>
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<mods:identifier id="280E0F55C45D1975162971556BCCB5CB" type="DOI">10.5281/zenodo.180435</mods:identifier>
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<mods:identifier id="5D0976EBCD2C1DF423FF8E3F042240E4" type="GBIF-Dataset">91705fcc-498c-41e0-a210-445a05bdfdab</mods:identifier>
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<mods:identifier id="8945D32FBAAD7ABDE81B02E1DECEB31E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E6B09AEEB701325DD0642441DD2F1266" type="Zenodo-Dep">180435</mods:identifier>
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<treatment id="191C6D0BFFC4FFC1FF7F3B8CF2BAF93B" ID-DOI="http://doi.org/10.5281/zenodo.6233902" ID-GBIF-Taxon="119353207" ID-Zenodo-Dep="6233902" LSID="urn:lsid:plazi:treatment:191C6D0BFFC4FFC1FF7F3B8CF2BAF93B" httpUri="http://treatment.plazi.org/id/191C6D0BFFC4FFC1FF7F3B8CF2BAF93B" lastPageId="8" lastPageNumber="47" pageId="2" pageNumber="41">
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<subSubSection id="D9AF8F96FFC4FFCBFF7F3B8CF716F944" pageId="2" pageNumber="41" type="nomenclature">
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<paragraph id="910ADC1DFFC4FFCBFF7F3B8CF5D2F921" blockId="2.[151,994,1671,1732]" box="[151,994,1671,1697]" pageId="2" pageNumber="41">
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<heading id="CA426B71FFC4FFCBFF7F3B8CF5D2F921" bold="true" box="[151,994,1671,1697]" fontSize="11" level="1" pageId="2" pageNumber="41" reason="1">
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<emphasis id="A3C1000FFFC4FFCBFF7F3B8CF5D2F921" bold="true" box="[151,994,1671,1697]" pageId="2" pageNumber="41">
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<taxonomicName id="56B5A79EFFC4FFCBFF7F3B8CF547F921" ID-CoL="6SYZZ" authority="Maruyama, Klimaszewski & Gusarov" authorityName="Maruyama, Klimaszewski & Gusarov" box="[151,887,1671,1697]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="yasukoae" status="sp. nov.">
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<emphasis id="A3C1000FFFC4FFCBFF7F3B8CF7A6F921" bold="true" box="[151,406,1671,1697]" italics="true" pageId="2" pageNumber="41">Osakatheta yasukoae</emphasis>
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Maruyama, Klimaszewski & Gusarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="B8F2BD74FFC4FFCBFC6C3B8CF5D2F921" box="[900,994,1671,1697]" pageId="2" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="910ADC1DFFC4FFCBFF7F3BA1F716F944" blockId="2.[151,994,1671,1732]" box="[151,294,1706,1732]" pageId="2" pageNumber="41">
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(
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<figureCitation id="098EC098FFC4FFCBFF773BA1F72DF944" box="[159,285,1706,1732]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 10" captionStart-2="FIGURES 11 – 18" captionStart-3="FIGURES 19 – 23" captionStart-4="FIGURES 24 – 27" captionStart-5="FIGURES 28 – 35" captionStart-6="FIGURE 36" captionStart-7="FIGURE 37" captionStartId-0="3.[151,269,1928,1952]" captionStartId-1="4.[151,269,1666,1690]" captionStartId-2="5.[151,269,1938,1962]" captionStartId-3="6.[151,269,1122,1146]" captionStartId-4="6.[151,269,1975,1999]" captionStartId-5="7.[151,269,1875,1899]" captionStartId-6="8.[151,255,1072,1096]" captionStartId-7="9.[151,255,1564,1588]" captionTargetBox-0="[239,1355,338,1901]" captionTargetBox-1="[155,1429,199,1636]" captionTargetBox-2="[189,1422,588,1913]" captionTargetBox-3="[226,1360,182,1098]" captionTargetBox-4="[241,1364,1224,1957]" captionTargetBox-5="[155,1424,194,1846]" captionTargetBox-6="[151,1435,194,1047]" captionTargetBox-7="[151,1435,394,1540]" captionTargetId-0="figure@3.[239,1355,336,1902]" captionTargetId-1="figure@4.[151,1436,194,1642]" captionTargetId-2="figure@5.[164,1429,578,1916]" captionTargetId-3="figure@6.[226,1360,182,1098]" captionTargetId-4="figure@6.[238,1366,1220,1966]" captionTargetId-5="figure@7.[151,1436,194,1851]" captionTargetId-6="figure@8.[151,1436,194,1048]" captionTargetId-7="figure@9.[151,1436,394,1540]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionText-0="FIGURES 1 – 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." captionText-1="FIGURES 6 – 10. Osakatheta yasukoae, mouthparts. 6 — labrum, dorsal view; 7 — epipharynx, adoral view; 8 — left mandible, dorsal view; 9 — left mandible, ventral view; 10 — right mandible, ventral view." captionText-2="FIGURES 11 – 18. Osakatheta yasukoae, mouthparts. 11 — prementum; 12 — hypopharynx, adoral view; 1 — right maxilla, ventral view; 14 — right lacinia, ventral view; 15 — right lacinia, dorsal view; 16 — right galea, ventral view; 17 — right galea, dorsal view; 18 — mentum." captionText-3="FIGURES 19 – 23. Osakatheta yasukoae, SEM photographs. 19 — pronotum, dorso-lateral view; 20 — prosternite, ventral view; 21 — meso- and metathorax, ventral view; 22 — elytra, dorso-lateral view; 23 — claws, right mid leg." captionText-4="FIGURES 24 – 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." captionText-5="FIGURES 28 – 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." captionText-6="FIGURE 36. The type locality of Osakatheta yasukoae, a tidal flat at Nankô-yachôen, Ôsaka-fu, Japan. The beetles were found under the stones in the bottom of this photograph. Photographed by Rikio Matsumoto." captionText-7="FIGURE 37. Known localities of Osakatheta yasukoae (black stars)." httpUri-0="https://zenodo.org/record/180436/files/figure.png" httpUri-1="https://zenodo.org/record/180437/files/figure.png" httpUri-2="https://zenodo.org/record/180438/files/figure.png" httpUri-3="https://zenodo.org/record/180439/files/figure.png" httpUri-4="https://zenodo.org/record/180440/files/figure.png" httpUri-5="https://zenodo.org/record/180441/files/figure.png" httpUri-6="https://zenodo.org/record/180442/files/figure.png" httpUri-7="https://zenodo.org/record/180443/files/figure.png" pageId="2" pageNumber="41">Figs. 1–37</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D9AF8F96FFC4FFCAFF7F3BF1F6C3FF5A" lastPageId="3" lastPageNumber="42" pageId="2" pageNumber="41" type="materials_examined">
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<paragraph id="910ADC1DFFC4FFCAFF7F3BF1F6C3FF5A" blockId="2.[151,1437,1786,2012]" lastBlockId="3.[151,1436,152,298]" lastPageId="3" lastPageNumber="42" pageId="2" pageNumber="41">
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<emphasis id="A3C1000FFFC4FFCBFF7F3BF1F7FDF894" bold="true" box="[151,461,1786,1812]" pageId="2" pageNumber="41">
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<typeStatus id="4E0E62BFFFC4FFCBFF7F3BF1F6E2F894" box="[151,210,1786,1812]" pageId="2" pageNumber="41">Type</typeStatus>
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material.
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<typeStatus id="4E0E62BFFFC4FFCBFEB33BF1F7FDF894" box="[347,461,1786,1812]" pageId="2" pageNumber="41" type="holotype">Holotype</typeStatus>
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</emphasis>
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ɗ:
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<emphasis id="A3C1000FFFC4FFCBFE173BF1F457F894" bold="true" box="[511,615,1786,1812]" pageId="2" pageNumber="41">
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<collectingCountry id="E9A29C8DFFC4FFCBFE173BF1F451F894" box="[511,609,1786,1812]" name="Japan" pageId="2" pageNumber="41">JAPAN</collectingCountry>
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:
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</emphasis>
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Ôsaka-fu, Ôsaka-shi, Suminoe-ku, Nankô-yachôen,
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<date id="E50BFADDFFC4FFCBFB0D3BF1F344F894" box="[1253,1396,1786,1812]" pageId="2" pageNumber="41" value="2002-04-11">11 IV 2002</date>
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, Y. Kawakami (
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<collectionCode id="F7A444D8FFC4FFCBFECD3A29F74DF8BC" box="[293,381,1826,1852]" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="2" pageNumber="41">NSMT</collectionCode>
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).
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<emphasis id="A3C1000FFFC4FFCBFE783A29F43DF8BC" bold="true" box="[400,525,1826,1852]" pageId="2" pageNumber="41">
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<typeStatus id="4E0E62BFFFC4FFCBFE783A29F43DF8BC" box="[400,525,1826,1852]" pageId="2" pageNumber="41" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(all collected by Y. Kawakami):
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<emphasis id="A3C1000FFFC4FFCBFC663A29F5C6F8BC" bold="true" box="[910,1014,1826,1852]" pageId="2" pageNumber="41">
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<collectingCountry id="E9A29C8DFFC4FFCBFC663A29F5C0F8BC" box="[910,1008,1826,1852]" name="Japan" pageId="2" pageNumber="41">JAPAN</collectingCountry>
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:
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</emphasis>
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7 ex., same data as
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<typeStatus id="4E0E62BFFFC4FFCBFB353A29F374F8BC" box="[1245,1348,1826,1852]" pageId="2" pageNumber="41" type="holotype">holotype</typeStatus>
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but,
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<date id="E50BFADDFFC4FFCBFA973A29F6C7F8E4" pageId="2" pageNumber="41" value="2002-03-27">27 III 2002</date>
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(
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<collectionCode id="F7A444D8FFC4FFCBFEE03A41F756F8E4" box="[264,358,1866,1892]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="2" pageNumber="41">OMNH</collectionCode>
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, cMM); 1 ex., ditto,
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<date id="E50BFADDFFC4FFCBFDB63A41F4D5F8E4" box="[606,741,1866,1892]" pageId="2" pageNumber="41" value="2002-03-28">28 III 2002</date>
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(cMM); 2 exs., ditto,
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<date id="E50BFADDFFC4FFCBFC063A41F246F8E4" box="[1006,1142,1866,1892]" pageId="2" pageNumber="41" value="2002-04-12">12 IV 2002</date>
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(cMM); 2 exs. (teneral), ditto,
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<date id="E50BFADDFFC4FFCBFF343A79F753F80C" box="[220,355,1906,1932]" pageId="2" pageNumber="41" value="2003-03-19">19 III 2003</date>
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(cMM); 24 exs., ditto,
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<date id="E50BFADDFFC4FFCBFD943A79F534F80C" box="[636,772,1906,1932]" pageId="2" pageNumber="41" value="2003-04-16">16 IV 2003</date>
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(
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<collectionCode id="F7A444D8FFC4FFCBFCFD3A79F55FF80C" LSID="urn:lsid:biocol.org:col:34795" box="[789,879,1906,1932]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="41">FMNH</collectionCode>
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,
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<collectionCode id="F7A444D8FFC4FFCBFC943A79F585F80C" box="[892,949,1906,1932]" pageId="2" pageNumber="41">LFC</collectionCode>
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,
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<collectionCode id="F7A444D8FFC4FFCBFC293A79F230F80C" box="[961,1024,1906,1932]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="2" pageNumber="41">CNC</collectionCode>
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,
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<collectionCode id="F7A444D8FFC4FFCBFBE63A79F25DF80C" box="[1038,1133,1906,1932]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="2" pageNumber="41">NHMO</collectionCode>
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); 7 exs., ditto,
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<date id="E50BFADDFFC4FFCBFAF63A79F3ADF80C" box="[1310,1437,1906,1932]" pageId="2" pageNumber="41" value="2003-05-21">21 V 2003</date>
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(cMM); 2 exs., ditto,
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<date id="E50BFADDFFC4FFCBFE713A91F413F834" box="[409,547,1946,1972]" pageId="2" pageNumber="41" value="2003-06-25">25 VI 2003</date>
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(cMM); 3 exs., Suminoe-ku, Nankô-kita Sakishima,
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<date id="E50BFADDFFC4FFCBFB773A91F32FF834" box="[1183,1311,1946,1972]" pageId="2" pageNumber="41" value="2003-05-22">22 V 2003</date>
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(cMM); 1 ex., Nishiyodogawa-ku, Hanakawa, Yodo-gawa,
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<date id="E50BFADDFFC4FFCBFD303AC9F56EF85C" box="[728,862,1986,2012]" pageId="2" pageNumber="41" value="2002-03-28">28 III 2002</date>
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(cMM); 3 exs., Nishiyodogawa-ku, Himeshima, Yodo-gawa,
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<date id="E50BFADDFFC5FFCAFED83D93F7F1FF32" box="[304,449,152,178]" pageId="3" pageNumber="42" value="2003-04-12">12 IV 2003</date>
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(cMM); [Mie-ken (Ise-wan)]: 1 ex., Matsuzaka-shi, Kushida-gawa,
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<date id="E50BFADDFFC5FFCAFAE43D93F3ACFF32" box="[1292,1436,152,178]" pageId="3" pageNumber="42" value="2002-04-28">28 IV 2002</date>
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(cMM).
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</paragraph>
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</subSubSection>
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<subSubSection id="D9AF8F96FFC5FFCAFF2D3DECF308FEAA" pageId="3" pageNumber="42" type="diagnosis">
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<paragraph id="910ADC1DFFC5FFCAFF2D3DECF308FEAA" blockId="3.[151,1436,152,298]" pageId="3" pageNumber="42">
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<emphasis id="A3C1000FFFC5FFCAFF2D3DECF775FE81" bold="true" box="[197,325,231,257]" pageId="3" pageNumber="42">Diagnosis.</emphasis>
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This is the only species of the genus currently known. If additional species of
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<taxonomicName id="56B5A79EFFC5FFCAFB033DE3F340FE81" box="[1259,1392,232,257]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
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<emphasis id="A3C1000FFFC5FFCAFB033DE3F340FE81" box="[1259,1392,232,257]" italics="true" pageId="3" pageNumber="42">Osakatheta</emphasis>
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||
</taxonomicName>
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are discovered, they are likely to differ in the shape of the median lobe of the aedeagus and spermatheca.
|
||
</paragraph>
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||
</subSubSection>
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<caption id="C5CA8C95FFC5FFCAFF7F3A83F36BF842" httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="3" pageNumber="42" targetBox="[239,1355,338,1901]" targetPageId="3">
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<paragraph id="910ADC1DFFC5FFCAFF7F3A83F36BF842" blockId="3.[151,1436,1928,1986]" pageId="3" pageNumber="42">
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<emphasis id="A3C1000FFFC5FFCAFF7F3A83F776F820" bold="true" box="[151,326,1928,1952]" pageId="3" pageNumber="42">FIGURES 1–5.</emphasis>
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<taxonomicName id="56B5A79EFFC5FFCAFEB93A82F407F820" box="[337,567,1929,1952]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="yasukoae">
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<emphasis id="A3C1000FFFC5FFCAFEB93A82F407F820" box="[337,567,1929,1952]" italics="true" pageId="3" pageNumber="42">Osakatheta yasukoae</emphasis>
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</taxonomicName>
|
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, SEM photographs. 1—habitus, dorsal view; 1a—pronotum, dorsal view; 2— facies, antero-dorsolateral view; 3—head, antero-dorsal view; 4—head, lateral view; 5—head, ventro-lateral view.
|
||
</paragraph>
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</caption>
|
||
<caption id="C5CA8C95FFC2FFCDFF7F3B89F211F93B" httpUri="https://zenodo.org/record/180437/files/figure.png" pageId="4" pageNumber="43" targetBox="[155,1429,199,1636]" targetPageId="4">
|
||
<paragraph id="910ADC1DFFC2FFCDFF7F3B89F211F93B" blockId="4.[151,1436,1666,1723]" pageId="4" pageNumber="43">
|
||
<emphasis id="A3C1000FFFC2FFCDFF7F3B89F760F91A" bold="true" box="[151,336,1666,1690]" pageId="4" pageNumber="43">FIGURES 6–10.</emphasis>
|
||
<taxonomicName id="56B5A79EFFC2FFCDFEBF3B89F40BF919" box="[343,571,1666,1689]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="yasukoae">
|
||
<emphasis id="A3C1000FFFC2FFCDFEBF3B89F40BF919" box="[343,571,1666,1689]" italics="true" pageId="4" pageNumber="43">Osakatheta yasukoae</emphasis>
|
||
</taxonomicName>
|
||
, mouthparts. 6—labrum, dorsal view; 7—epipharynx, adoral view; 8—left mandible, dorsal view; 9—left mandible, ventral view; 10—right mandible, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D9AF8F96FFC2FFC1FF2D3BE1F5B5FA4B" lastPageId="8" lastPageNumber="47" pageId="4" pageNumber="43" type="description">
|
||
<paragraph id="910ADC1DFFC2FFCCFF2D3BE1F4ECFD9A" blockId="4.[151,1437,1770,2036]" lastBlockId="5.[151,1436,152,538]" lastPageId="5" lastPageNumber="44" pageId="4" pageNumber="43">
|
||
<emphasis id="A3C1000FFFC2FFCDFF2D3BE1F76DF884" bold="true" box="[197,349,1770,1796]" pageId="4" pageNumber="43">Description.</emphasis>
|
||
Reddish brown in ground colour; antennae, mouthparts and legs paler. Head (
|
||
<figureCitation id="098EC098FFC2FFCDFAC83BE1F3A3F884" box="[1312,1427,1770,1796]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="4" pageNumber="43">Figs. 3–5</figureCitation>
|
||
) wider than long (head width/head length ratio ca. 1.09); apical margin of clypeus truncate; surface smooth, moderately covered with setae, which are somewhat shorter than those on pronotum and elytra; length of eyes 0.39–0.40 times as long as head width. Antennae (
|
||
<figureCitation id="098EC098FFC2FFCDFD0E3A69F565F8FC" box="[742,853,1890,1916]" captionStart="FIGURES 1 – 5" captionStartId="3.[151,269,1928,1952]" captionTargetBox="[239,1355,338,1901]" captionTargetId="figure@3.[239,1355,336,1902]" captionTargetPageId="3" captionText="FIGURES 1 – 5. Osakatheta yasukoae, SEM photographs. 1 — habitus, dorsal view; 1 a—pronotum, dorsal view; 2 — facies, antero-dorsolateral view; 3 — head, antero-dorsal view; 4 — head, lateral view; 5 — head, ventro-lateral view." httpUri="https://zenodo.org/record/180436/files/figure.png" pageId="4" pageNumber="43">Figs. 1, 2</figureCitation>
|
||
) almost as long as head and pronotum combined; antennomere I long, slightly longer than II; antennomere II almost as long as III; antennomeres 4–10 moniliform, each almost as long as wide; antennomere XI slightly shorter than 1st; approximate relative lengths of segments from basal to apex: 5.0:4.0:4.0:2.0:2.0:2.0:2.0:2.5:2.5:2.5:4.0. Pronotum slightly transverse, about 1.2 times as wide as long, widest around anterior 2/5, narrowed posteriorly; postero-lateral corner rounded; surface finely punctured, densely covered with setae, with 6 macrosetae along lateral margin. Scutellum with integument smooth and sparsely covered with setae. Elytra slightly wider than long (length measured at humeral level), slightly broadened posteriorly; surface finely punctured, densely covered with setae, with 4 or 5 small macrosetae laterally. Legs long, mesotibia 0.9 times shorter than elytra; approximate relative lengths of tarsal segments from basal to apical: 9.0:9.0:8.0:14.5 (protarsus); 15.0:10.0:8.0:7.0:12.0 (mesotarsus); 14.5:12.5:10.0:8.0:15.0 (metatarsus). Abdomen oblong, wider than elytra, widest around segments V and VI; surface smooth; tergites III–VII densely covered with setae; tergite VIII (
|
||
<figureCitation id="098EC098FFC3FFCCFC143CBBF263FE4A" box="[1020,1107,432,458]" captionStart="FIGURES 24 – 27" captionStartId="6.[151,269,1975,1999]" captionTargetBox="[241,1364,1224,1957]" captionTargetId="figure@6.[238,1366,1220,1966]" captionTargetPageId="6" captionText="FIGURES 24 – 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." httpUri="https://zenodo.org/record/180440/files/figure.png" pageId="5" pageNumber="44">Fig. 24</figureCitation>
|
||
) broadly rounded on posterior margin, densely covered with setae, with 4 macrosetae on each side; sternite VIII (
|
||
<figureCitation id="098EC098FFC3FFCCFB443CD3F333FE72" box="[1196,1283,472,498]" captionStart="FIGURES 24 – 27" captionStartId="6.[151,269,1975,1999]" captionTargetBox="[241,1364,1224,1957]" captionTargetId="figure@6.[238,1366,1220,1966]" captionTargetPageId="6" captionText="FIGURES 24 – 27. Osakatheta yasukoae, abdominal segment VIII. 24 — male tergite VIII; 25 — male sternite VIII; 26 — apex of male sternite VIII; 27 — apex of female sternite VIII." httpUri="https://zenodo.org/record/180440/files/figure.png" pageId="5" pageNumber="44">Fig. 25</figureCitation>
|
||
) rounded on posterior margin, with 6 macrosetae on each side.
|
||
</paragraph>
|
||
<caption id="C5CA8C95FFC3FFCCFF7F3A99F404F86F" httpUri="https://zenodo.org/record/180438/files/figure.png" pageId="5" pageNumber="44" targetBox="[189,1422,588,1913]" targetPageId="5">
|
||
<paragraph id="910ADC1DFFC3FFCCFF7F3A99F404F86F" blockId="5.[151,1436,1938,2031]" pageId="5" pageNumber="44">
|
||
<emphasis id="A3C1000FFFC3FFCCFF7F3A99F76EF82A" bold="true" box="[151,350,1938,1962]" pageId="5" pageNumber="44">FIGURES 11–18.</emphasis>
|
||
<taxonomicName id="56B5A79EFFC3FFCCFE8F3A99F47CF829" box="[359,588,1938,1961]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="44" phylum="Arthropoda" rank="species" species="yasukoae">
|
||
<emphasis id="A3C1000FFFC3FFCCFE8F3A99F47CF829" box="[359,588,1938,1961]" italics="true" pageId="5" pageNumber="44">Osakatheta yasukoae</emphasis>
|
||
</taxonomicName>
|
||
, mouthparts. 11—prementum; 12—hypopharynx, adoral view; 1—right maxilla, ventral view; 14—right lacinia, ventral view; 15—right lacinia, dorsal view; 16—right galea, ventral view; 17— right galea, dorsal view; 18—mentum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C5CA8C95FFC0FFCFFF7F3969F2C9FB1C" httpUri="https://zenodo.org/record/180439/files/figure.png" pageId="6" pageNumber="45" targetBox="[226,1360,182,1098]" targetPageId="6">
|
||
<paragraph id="910ADC1DFFC0FFCFFF7F3969F2C9FB1C" blockId="6.[151,1436,1122,1180]" pageId="6" pageNumber="45">
|
||
<emphasis id="A3C1000FFFC0FFCFFF7F3969F76EFBFA" bold="true" box="[151,350,1122,1146]" pageId="6" pageNumber="45">FIGURES 19–23.</emphasis>
|
||
<taxonomicName id="56B5A79EFFC0FFCFFE8C3968F477FBFA" box="[356,583,1123,1146]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="yasukoae">
|
||
<emphasis id="A3C1000FFFC0FFCFFE8C3968F477FBFA" box="[356,583,1123,1146]" italics="true" pageId="6" pageNumber="45">Osakatheta yasukoae</emphasis>
|
||
</taxonomicName>
|
||
, SEM photographs. 19—pronotum, dorso-lateral view; 20—prosternite, ventral view; 21—meso- and metathorax, ventral view; 22—elytra, dorso-lateral view; 23—claws, right mid leg.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C5CA8C95FFC0FFCFFF7F3ABCF52DF871" httpUri="https://zenodo.org/record/180440/files/figure.png" pageId="6" pageNumber="45" targetBox="[241,1364,1224,1957]" targetPageId="6">
|
||
<paragraph id="910ADC1DFFC0FFCFFF7F3ABCF52DF871" blockId="6.[151,1436,1975,2033]" pageId="6" pageNumber="45">
|
||
<emphasis id="A3C1000FFFC0FFCFFF7F3ABCF76EF84F" bold="true" box="[151,350,1975,1999]" pageId="6" pageNumber="45">FIGURES 24–27.</emphasis>
|
||
<taxonomicName id="56B5A79EFFC0FFCFFE8D3AB3F479F84F" box="[357,585,1976,1999]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="45" phylum="Arthropoda" rank="species" species="yasukoae">
|
||
<emphasis id="A3C1000FFFC0FFCFFE8D3AB3F479F84F" box="[357,585,1976,1999]" italics="true" pageId="6" pageNumber="45">Osakatheta yasukoae</emphasis>
|
||
</taxonomicName>
|
||
, abdominal segment VIII. 24—male tergite VIII; 25—male sternite VIII; 26— apex of male sternite VIII; 27—apex of female sternite VIII.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C5CA8C95FFC1FFCEFF7F3A58F2CAF830" httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="7" pageNumber="46" targetBox="[155,1424,194,1846]" targetPageId="7">
|
||
<paragraph id="910ADC1DFFC1FFCEFF7F3A58F2CAF830" blockId="7.[151,1436,1875,1968]" pageId="7" pageNumber="46">
|
||
<emphasis id="A3C1000FFFC1FFCEFF7F3A58F750F8EB" bold="true" box="[151,352,1875,1899]" pageId="7" pageNumber="46">FIGURES 28–35.</emphasis>
|
||
<taxonomicName id="56B5A79EFFC1FFCEFE803A58F47DF8EA" box="[360,589,1875,1898]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="yasukoae">
|
||
<emphasis id="A3C1000FFFC1FFCEFE803A58F47DF8EA" box="[360,589,1875,1898]" italics="true" pageId="7" pageNumber="46">Osakatheta yasukoae</emphasis>
|
||
</taxonomicName>
|
||
, male genitalia and spermatheca. 28—median lobe of aedeagus, lateral view; 29—median lobe of aedeagus, parameral view; 30—everted internal sac, lateral view; 31—everted internal sac, parameral view; 32—paramere, lateral view; 33—apical lobe of paramerite, lateral view; 34, 35—spermatheca.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C5CA8C95FFCEFFC1FF7F393BF244FBEA" httpUri="https://zenodo.org/record/180442/files/figure.png" pageId="8" pageNumber="47" targetBox="[151,1435,194,1047]" targetPageId="8">
|
||
<paragraph id="910ADC1DFFCEFFC1FF7F393BF244FBEA" blockId="8.[151,1436,1072,1130]" pageId="8" pageNumber="47">
|
||
<emphasis id="A3C1000FFFCEFFC1FF7F393BF716FBC8" bold="true" box="[151,294,1072,1096]" pageId="8" pageNumber="47">FIGURE 36.</emphasis>
|
||
The type locality of
|
||
<taxonomicName id="56B5A79EFFCEFFC1FDEA393AF4D4FBC8" box="[514,740,1073,1096]" class="Insecta" family="Staphylinidae" genus="Osakatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="47" phylum="Arthropoda" rank="species" species="yasukoae">
|
||
<emphasis id="A3C1000FFFCEFFC1FDEA393AF4D4FBC8" box="[514,740,1073,1096]" italics="true" pageId="8" pageNumber="47">Osakatheta yasukoae</emphasis>
|
||
</taxonomicName>
|
||
, a tidal flat at Nankô-yachôen, Ôsaka-fu, Japan. The beetles were found under the stones in the bottom of this photograph. Photographed by Rikio Matsumoto.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="910ADC1DFFCEFFC1FF2D3992F5E7FAAB" blockId="8.[151,1437,1177,1723]" pageId="8" pageNumber="47">
|
||
<emphasis id="A3C1000FFFCEFFC1FF2D3992F731FB32" box="[197,257,1177,1202]" italics="true" pageId="8" pageNumber="47">Male</emphasis>
|
||
. Median lobe of aedeagus as in
|
||
<figureCitation id="098EC098FFCEFFC1FD903992F536FB33" box="[632,774,1177,1203]" captionStart="FIGURES 28 – 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 – 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="8" pageNumber="47">Figs. 28–29</figureCitation>
|
||
; in lateral view (
|
||
<figureCitation id="098EC098FFCEFFC1FC2F3992F22BFB33" box="[967,1051,1177,1203]" captionStart="FIGURES 28 – 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 – 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="8" pageNumber="47">Fig. 28</figureCitation>
|
||
) apical lobe slightly bent paramerally, much narrowed subapically, sinuate abparamerally and rounded at apex; basal ridge weakly convex; copulatory piece of inner sac as in
|
||
<figureCitation id="098EC098FFCEFFC1FDC739E2F48EFA83" box="[559,702,1257,1283]" captionStart="FIGURES 28 – 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 – 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="8" pageNumber="47">Figs. 30, 31</figureCitation>
|
||
. Paramere (
|
||
<figureCitation id="098EC098FFCEFFC1FCA039E2F5E5FA83" box="[840,981,1257,1283]" captionStart="FIGURES 28 – 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 – 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="8" pageNumber="47">Figs. 32–33</figureCitation>
|
||
) with apical lobe of paramerite dilated apically, with basal two setae about 4 times as long as apical two setae.
|
||
</paragraph>
|
||
<paragraph id="910ADC1DFFCEFFC1FF2D3832F339FAFB" blockId="8.[151,1437,1177,1723]" pageId="8" pageNumber="47">
|
||
<emphasis id="A3C1000FFFCEFFC1FF2D3832F72CFAD2" box="[197,284,1337,1362]" italics="true" pageId="8" pageNumber="47">Female</emphasis>
|
||
. Spermatheca (
|
||
<figureCitation id="098EC098FFCEFFC1FE3B3832F450FAD3" box="[467,608,1337,1363]" captionStart="FIGURES 28 – 35" captionStartId="7.[151,269,1875,1899]" captionTargetBox="[155,1424,194,1846]" captionTargetId="figure@7.[151,1436,194,1851]" captionTargetPageId="7" captionText="FIGURES 28 – 35. Osakatheta yasukoae, male genitalia and spermatheca. 28 — median lobe of aedeagus, lateral view; 29 — median lobe of aedeagus, parameral view; 30 — everted internal sac, lateral view; 31 — everted internal sac, parameral view; 32 — paramere, lateral view; 33 — apical lobe of paramerite, lateral view; 34, 35 — spermatheca." httpUri="https://zenodo.org/record/180441/files/figure.png" pageId="8" pageNumber="47">Figs. 34–35</figureCitation>
|
||
) with basal part of stem twice curved around base; capsule dilated apically, about 1/5 as long as stem, densely covered with wrinkles on internal side of apical 2/5–1/2.
|
||
</paragraph>
|
||
<paragraph id="910ADC1DFFCEFFC1FF2D3882F5B5FA4B" blockId="8.[151,1437,1177,1723]" pageId="8" pageNumber="47">
|
||
<emphasis id="A3C1000FFFCEFFC1FF2D3882F75FFA22" box="[197,367,1417,1442]" italics="true" pageId="8" pageNumber="47">Measurements</emphasis>
|
||
. Body length, ca.
|
||
<quantity id="564D71F8FFCEFFC1FDA73882F4D1FA23" box="[591,737,1417,1443]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="8" pageNumber="47" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
|
||
; fore body length, ca.
|
||
<quantity id="564D71F8FFCEFFC1FC193882F2B4FA23" box="[1009,1156,1417,1443]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="8" pageNumber="47" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
|
||
; head width,
|
||
<quantity id="564D71F8FFCEFFC1FACF3882F6F8FA4B" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.6" metricValueMin="4.5" pageId="8" pageNumber="47" unit="mm" value="0.455" valueMax="0.46" valueMin="0.45">0.45–0.46 mm</quantity>
|
||
; pronotal length, 0.43–0.45; pronotal width,
|
||
<quantity id="564D71F8FFCEFFC1FD3B38BAF5B1FA4B" box="[723,897,1457,1483]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="8" pageNumber="47" unit="mm" value="0.51" valueMax="0.52" valueMin="0.5">0.50–0.52 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AF8F96FFCEFFC1FF2D38D3F5BAF9C3" pageId="8" pageNumber="47" type="biology_ecology">
|
||
<paragraph id="910ADC1DFFCEFFC1FF2D38D3F5BAF9C3" blockId="8.[151,1437,1177,1723]" pageId="8" pageNumber="47">
|
||
<emphasis id="A3C1000FFFCEFFC1FF2D38D3F71AFA72" bold="true" box="[197,298,1496,1522]" pageId="8" pageNumber="47">Biology.</emphasis>
|
||
The adult beetles were found under stones on mudflats at river mouths, which are located under the high water line (
|
||
<figureCitation id="098EC098FFCEFFC1FE6C3B0AF7ECF99B" box="[388,476,1537,1563]" captionStart="FIGURE 36" captionStartId="8.[151,255,1072,1096]" captionTargetBox="[151,1435,194,1047]" captionTargetId="figure@8.[151,1436,194,1048]" captionTargetPageId="8" captionText="FIGURE 36. The type locality of Osakatheta yasukoae, a tidal flat at Nankô-yachôen, Ôsaka-fu, Japan. The beetles were found under the stones in the bottom of this photograph. Photographed by Rikio Matsumoto." httpUri="https://zenodo.org/record/180442/files/figure.png" pageId="8" pageNumber="47">Fig. 36</figureCitation>
|
||
). The beetles were observed from the beginning of March to the end of June, and in March were all teneral. Immature stages have not been found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AF8F96FFCEFFC1FF2D3B5BF5AAF9EB" box="[197,922,1616,1643]" pageId="8" pageNumber="47" type="distribution">
|
||
<paragraph id="910ADC1DFFCEFFC1FF2D3B5BF5AAF9EB" blockId="8.[151,1437,1177,1723]" box="[197,922,1616,1643]" pageId="8" pageNumber="47">
|
||
<emphasis id="A3C1000FFFCEFFC1FF2D3B5BF756F9EA" bold="true" box="[197,358,1616,1642]" pageId="8" pageNumber="47">Distribution.</emphasis>
|
||
<collectingCountry id="E9A29C8DFFCEFFC1FE853B5AF784F9EB" box="[365,436,1617,1643]" name="Japan" pageId="8" pageNumber="47">Japan</collectingCountry>
|
||
: Honshû (Ôsaka-wan, Ise-wan) (
|
||
<figureCitation id="098EC098FFCEFFC1FCDC3B5AF5BCF9EB" box="[820,908,1617,1643]" captionStart="FIGURE 37" captionStartId="9.[151,255,1564,1588]" captionTargetBox="[151,1435,394,1540]" captionTargetId="figure@9.[151,1436,394,1540]" captionTargetPageId="9" captionText="FIGURE 37. Known localities of Osakatheta yasukoae (black stars)." httpUri="https://zenodo.org/record/180443/files/figure.png" pageId="8" pageNumber="47">Fig. 37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9AF8F96FFCEFFC1FF2D3B73F2BAF93B" pageId="8" pageNumber="47" type="etymology">
|
||
<paragraph id="910ADC1DFFCEFFC1FF2D3B73F2BAF93B" blockId="8.[151,1437,1177,1723]" pageId="8" pageNumber="47">
|
||
<emphasis id="A3C1000FFFCEFFC1FF2D3B73F762F912" bold="true" box="[197,338,1656,1682]" pageId="8" pageNumber="47">Etymology.</emphasis>
|
||
Dedicated to Mrs. Yasuko Kawakami, Ôsaka-fu, collector of the
|
||
<typeStatus id="4E0E62BFFFCEFFC1FBB83B72F2B2F913" box="[1104,1154,1657,1683]" pageId="8" pageNumber="47">type</typeStatus>
|
||
series, for her contribution to the knowledge of the coastal beetle fauna of the Ôsaka-wan and adjacent areas.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |