treatments-xml/data/07/34/87/073487A0FFC1FE34FF24C974FCA5FF06.xml

<document id="E0AB438FDE9F38AD505416E9553F26B0" ID-CLB-Dataset="45829" ID-DOI="10.5281/zenodo.204780" ID-GBIF-Dataset="0fb466d4-dcfd-484f-9c9c-c78b9a3f18d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="204780" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460417569566" checkinUser="plazi" docAuthor="Breitman, Maria Florencia, Pérez, Cristian Hernán Fulvio, Parra, Micaela, Morando, Mariana, Sites, Jack Walter &amp; Avila, Luciano Javier" docDate="2011" docId="073487A0FFC1FE34FF24C974FCA5FF06" docLanguage="en" docName="zt03123p048.pdf" docOrigin="Zootaxa 3123" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Liolaemus caparensis Breitman, Pérez, Parra, Morando, Sites &amp; Avila, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FB0DFFD8FFC5FE38FFB3CF26FFA1FFF1" masterDocTitle="New species of lizard from the magellanicus clade of the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemidae) from southern Patagonia" masterLastPageNumber="48" masterPageNumber="32" pageNumber="36" updateTime="1698329089384" updateUser="ExternalLinkService">
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<mods:title id="EA6A2820ECAE4667B7617A07EFED01D6">New species of lizard from the magellanicus clade of the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemidae) from southern Patagonia</mods:title>
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<mods:namePart id="E41A3C411E6975D852B94214F587A9DC">Breitman, Maria Florencia</mods:namePart>
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<mods:namePart id="868C740296610E4596FFA465DFA1C108">Pérez, Cristian Hernán Fulvio</mods:namePart>
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<mods:namePart id="0E246E858A72F5F92574CC0F5842D55F">Parra, Micaela</mods:namePart>
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<mods:namePart id="6B2D56B78B71BDA6212C45781F37DD08">Morando, Mariana</mods:namePart>
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<mods:namePart id="DB72A5DB89542246884D1801E6DFB452">Sites, Jack Walter</mods:namePart>
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<mods:namePart id="58DD9723345F27FCB243A0BE5DF3C4B4">Avila, Luciano Javier</mods:namePart>
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<mods:date id="29BC5A6B7DD8FAEEEFE7032953A4402D">2011</mods:date>
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<treatment id="073487A0FFC1FE34FF24C974FCA5FF06" ID-DOI="http://doi.org/10.5281/zenodo.6191226" ID-GBIF-Taxon="119425581" ID-Zenodo-Dep="6191226" LSID="urn:lsid:plazi:treatment:073487A0FFC1FE34FF24C974FCA5FF06" httpUri="http://treatment.plazi.org/id/073487A0FFC1FE34FF24C974FCA5FF06" lastPageId="12" lastPageNumber="43" pageId="4" pageNumber="36">
<subSubSection id="C787653DFFC1FE3CFF24C974FEA6F97C" pageId="4" pageNumber="36" type="nomenclature">
<paragraph id="8F2236B6FFC1FE3CFF24C974FDA3F99D" blockId="4.[151,514,1618,1677]" box="[151,514,1618,1644]" pageId="4" pageNumber="36">
<heading id="D46A81DAFFC1FE3CFF24C974FDA3F99D" bold="true" box="[151,514,1618,1644]" fontSize="11" level="1" pageId="4" pageNumber="36" reason="1">
<emphasis id="BDE9EAA4FFC1FE3CFF24C974FDA3F99D" bold="true" box="[151,514,1618,1644]" pageId="4" pageNumber="36">
<taxonomicName id="489D4D35FFC1FE3CFF24C974FE3FF99D" box="[151,414,1618,1644]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC1FE3CFF24C974FE3FF99D" bold="true" box="[151,414,1618,1644]" italics="true" pageId="4" pageNumber="36">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC1FE3CFE16C974FDA3F99D" box="[421,514,1618,1644]" pageId="4" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8F2236B6FFC1FE3CFF24C953FEA6F97C" blockId="4.[151,514,1618,1677]" box="[151,263,1653,1677]" pageId="4" pageNumber="36">
(
<figureCitation id="17A62A33FFC1FE3CFF2DC953FF5EF97C" box="[158,255,1653,1677]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1758]" captionTargetBox="[254,1327,265,1713]" captionTargetId="figure@7.[254,1327,265,1714]" captionTargetPageId="7" captionText="FIGURE 2. Upper, dorsal and ventral views of male holotype in life of Liolaemus caparensis sp. nov. MLP. S 2628; below, dorsal and ventral views of female (LJAMM-CNP 9387)." httpUri="https://zenodo.org/record/204782/files/figure.png" pageId="4" pageNumber="36">Figure 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C787653DFFC1FE3CFF24C99EFDB1F878" pageId="4" pageNumber="36" type="materials_examined">
<paragraph id="8F2236B6FFC1FE3CFF24C99EFBC8F8EB" blockId="4.[151,1436,1719,2038]" pageId="4" pageNumber="36">
<emphasis id="BDE9EAA4FFC1FE3CFF24C99EFEAAF921" bold="true" box="[151,267,1720,1744]" pageId="4" pageNumber="36">
<typeStatus id="50268814FFC1FE3CFF24C99EFEA7F921" box="[151,262,1720,1744]" pageId="4" pageNumber="36" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="E98CAE73FFC1FE3CFEA1C991FEEFF93E" box="[274,334,1719,1743]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="4" pageNumber="36">MLP</collectionCode>
.S 2628 (
<figureCitation id="17A62A33FFC1FE3CFE1DC991FDB0F93E" box="[430,529,1719,1743]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1758]" captionTargetBox="[254,1327,265,1713]" captionTargetId="figure@7.[254,1327,265,1714]" captionTargetPageId="7" captionText="FIGURE 2. Upper, dorsal and ventral views of male holotype in life of Liolaemus caparensis sp. nov. MLP. S 2628; below, dorsal and ventral views of female (LJAMM-CNP 9387)." httpUri="https://zenodo.org/record/204782/files/figure.png" pageId="4" pageNumber="36">Figure 2</figureCitation>
), an adult male from Campo Las Piedras plateau, 
<quantity id="48659B53FFC1FE3CFBF4C99EFB20F93E" box="[1095,1153,1719,1743]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="36" unit="km" value="7.0">7 km</quantity>
N Punta del Lago ranch, Lago 
<collectingCountry id="F78A7626FFC1FE3CFF6BC9FAFEE8F905" box="[216,329,1756,1780]" name="Argentina" pageId="4" pageNumber="36">Argentino</collectingCountry>
department, Santa Cruz province, 
<collectingCountry id="F78A7626FFC1FE3CFD61C9FAFCE3F905" box="[722,834,1756,1780]" name="Argentina" pageId="4" pageNumber="36">Argentina</collectingCountry>
(
<geoCoordinate id="EAA95071FFC1FE3CFCE2C9FAFBA3F905" box="[849,1026,1756,1780]" direction="south" orientation="latitude" pageId="4" pageNumber="36" precision="1" value="-49.56972">49º 34’ 11.0” S</geoCoordinate>
, 
<geoCoordinate id="EAA95071FFC1FE3CFBBCC9FAFB69F905" box="[1039,1224,1756,1780]" direction="west" orientation="longitude" pageId="4" pageNumber="36" precision="1" value="-72.04775">72º 02’ 51.9” W</geoCoordinate>
, 
<quantity id="48659B53FFC1FE3CFB67C9FAFABEF905" box="[1236,1311,1756,1780]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="4" pageNumber="36" unit="m" value="868.0">868 m</quantity>
; 
<figureCitation id="17A62A33FFC1FE3CFA98C9FAFA37F905" box="[1323,1430,1756,1780]" captionStart="FIGURE 3" captionStartId="9.[151,250,1967,1990]" captionTargetBox="[210,1374,193,1942]" captionTargetId="figure@9.[210,1375,193,1942]" captionTargetPageId="9" captionText="FIGURE 3. Type locality of Liolaemus caparensis sp. nov., Subantartic Phytogeographic province, Estepa de Festuca pallescens District (49 º 34 ’ 11.0 ” S, 72 º 02 ’ 51.9 ” W)." httpUri="https://zenodo.org/record/204783/files/figure.png" pageId="4" pageNumber="36">Figures 3</figureCitation>
, 
<figureCitation id="17A62A33FFC1FE3CFF24C824FF06F8EB" box="[151,167,1794,1818]" captionStart="FIGURE 4" captionStartId="10.[151,250,1104,1127]" captionTargetBox="[164,1415,205,1066]" captionTargetId="figure@10.[151,1436,193,1083]" captionTargetPageId="10" captionText="FIGURE 4. Distribution map for species of the magellanicus group, with localities sampled superimposed on regional elevation (shading). White square: Liolaemus caparensis sp. nov.; black squares: distribution of L. magellanicus; black and gray squares: L. magellanicus localities used in this study from LJAMM-CNP collection. Arrow indicates type locality of L. magellanicus (in Chile). SC: Santa Cruz province. Notice L. caparensis sp. nov. distribution does not overlap with the distribution of L. magellanicus (south of the Santa Cruz river)." httpUri="https://zenodo.org/record/204784/files/figure.png" pageId="4" pageNumber="36">4</figureCitation>
), L.J. Avila, C.H.F. Pérez, M.F. Breitman and N. Feltrin collectors, 
<date id="FB231076FFC1FE3CFC2FC824FBC4F8EB" box="[924,1125,1791,1819]" pageId="4" pageNumber="36" value="2008-01-16">16th January 2008</date>
.
</paragraph>
<paragraph id="8F2236B6FFC1FE3CFF76C801FDB1F878" blockId="4.[151,1436,1719,2038]" pageId="4" pageNumber="36">
<emphasis id="BDE9EAA4FFC1FE3CFF76C801FEE5F8CE" bold="true" box="[197,324,1831,1855]" pageId="4" pageNumber="36">
<typeStatus id="50268814FFC1FE3CFF76C801FE9EF8CE" box="[197,319,1831,1855]" pageId="4" pageNumber="36" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="E98CAE73FFC1FE3CFEFCC801FE10F8CE" box="[335,433,1831,1855]" pageId="4" pageNumber="36">LJAMM</collectionCode>
–CNP 9379–9380 adult males, 
<collectionCode id="E98CAE73FFC1FE3CFC90C801FC24F8CE" box="[803,901,1831,1855]" pageId="4" pageNumber="36">LJAMM</collectionCode>
–CNP 9383–9387 females and 
<collectionCode id="E98CAE73FFC1FE3CFB46C801FAF6F8CE" box="[1269,1367,1831,1855]" pageId="4" pageNumber="36">LJAMM</collectionCode>
–CNP 9382, 9388, 9389 juveniles; from the same locality of 
<typeStatus id="50268814FFC1FE3CFD42C86CFCF7F893" box="[753,854,1866,1890]" pageId="4" pageNumber="36" type="holotype">holotype</typeStatus>
, L.J. Avila, C.H.F. Pérez, M.F. Breitman and N. Feltrin collectors, 
<date id="FB231076FFC1FE3CFE8CC857FDADF878" box="[319,524,1902,1930]" pageId="4" pageNumber="36" value="2008-01-16">16 th January 2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C787653DFFC1FE3DFF76C8B3FBC4FE3E" lastPageId="5" lastPageNumber="37" pageId="4" pageNumber="36" type="diagnosis">
<paragraph id="8F2236B6FFC1FE3DFF76C8B3FBC4FE3E" blockId="4.[151,1436,1719,2038]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="37" pageId="4" pageNumber="36">
<emphasis id="BDE9EAA4FFC1FE3CFF76C8B3FE9EF85F" bold="true" box="[197,319,1941,1966]" pageId="4" pageNumber="36">Diagnosis.</emphasis>
<taxonomicName id="489D4D35FFC1FE3CFEF6C8B0FD99F85F" box="[325,568,1942,1966]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC1FE3CFEF6C8B0FD99F85F" box="[325,568,1942,1966]" italics="true" pageId="4" pageNumber="36">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC1FE3CFD8CC8B0FD39F85F" box="[575,664,1942,1966]" pageId="4" pageNumber="36" rank="species">
<emphasis id="BDE9EAA4FFC1FE3CFD8CC8B0FD39F85F" bold="true" box="[575,664,1942,1966]" pageId="4" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
is a member of the 
<taxonomicName id="489D4D35FFC1FE3CFCC1C8B0FBA6F85F" box="[882,1031,1942,1966]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFC1FE3CFCC1C8B0FBA6F85F" box="[882,1031,1942,1966]" italics="true" pageId="4" pageNumber="36">magellanicus</emphasis>
</taxonomicName>
group, in the 
<taxonomicName id="489D4D35FFC1FE3CFB12C8B1FAC9F85F" box="[1185,1384,1942,1966]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFC1FE3CFB12C8B1FAC9F85F" box="[1185,1384,1942,1966]" italics="true" pageId="4" pageNumber="36">L. lineomaculatus</emphasis>
</taxonomicName>
section, based on molecular evidence (
<figureCitation id="17A62A33FFC1FE3CFD9EC89FFD2FF820" box="[557,654,1977,2001]" captionStart="FIGURE 1" captionStartId="5.[151,250,1174,1197]" captionTargetBox="[159,1397,515,1143]" captionTargetId="figure@5.[140,1425,515,1153]" captionTargetPageId="5" captionText="FIGURE 1. Relationships between Liolaemus caparensis sp. nov. and species of the L. lineomaculatus section and selected species of the subgenus Eulaemus and Liolaemus sensu stricto. On the left, phylogenetic relationships recovered using seven nuclear genes (Cmos, ACM 4 tg, PRLR, LDA 8 F, LDA 1 D, LDA 9 C and LDA 9 E, 4180 bp total alignment length) with Bayesian species-tree inference; bold branches represent well supported nodes (Pp&gt; 0.95). On the right, the Bayesian tree (modified from Breitman et al. 2011), represents a concatenated analyses (5865 bp) including the seven nuclear genes plus two mitochondrial genes (12 S and cyt- b), and summarizes information from MP and ML methods. Nodes with high support from three methods (MP, jackknife and ML bootstrap&gt; 0.70; Bayesian Pp&gt; 0.95) are identified by bold branches; solid circles show nodes with weak MP support, and open circles nodes with weak MP and ML support." httpUri="https://zenodo.org/record/204781/files/figure.png" pageId="4" pageNumber="36">Figure 1</figureCitation>
). 
<taxonomicName id="489D4D35FFC1FE3CFD1BC89FFC3FF820" box="[680,926,1977,2001]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC1FE3CFD1BC89FFC3FF820" box="[680,926,1977,2001]" italics="true" pageId="4" pageNumber="36">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC1FE3CFC14C89CFBA2F823" box="[935,1027,1978,2002]" pageId="4" pageNumber="36" rank="species">
<emphasis id="BDE9EAA4FFC1FE3CFC14C89CFBA2F823" bold="true" box="[935,1027,1978,2002]" pageId="4" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="489D4D35FFC1FE3CFB2EC89CFAF0F820" box="[1181,1361,1977,2001]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFC1FE3CFB2EC89CFAF0F820" box="[1181,1361,1977,2001]" italics="true" pageId="4" pageNumber="36">L. magellanicus</emphasis>
</taxonomicName>
in the following traits: shorter tibia-fibula length (6.19-8.62, X = 7.24 vs. 6.91-10.76, X = 9.09; 
<emphasis id="BDE9EAA4FFC1FE3CFB07C8F9FB63F807" box="[1204,1218,2015,2038]" italics="true" pageId="4" pageNumber="36">p</emphasis>
&lt;0.0001), shorter radius-ulna length (5.24-5.66, X = 5.24 vs. 5.22-6.88, X = 5.98; 
<emphasis id="BDE9EAA4FFC0FE3DFCD4CFBFFCD4FF41" box="[871,885,153,176]" italics="true" pageId="5" pageNumber="37">p</emphasis>
= 0.0017), larger number of midbody scales (39- 55, X = 43.82 vs. 34-48, X = 40.44; 
<emphasis id="BDE9EAA4FFC0FE3DFD85CF9BFDE5FF25" box="[566,580,189,212]" italics="true" pageId="5" pageNumber="37">p</emphasis>
= 0.0282), larger number of dorsal scales (39-47, X = 41.45 vs. 34-45, X = 37.5; 
<emphasis id="BDE9EAA4FFC0FE3DFF65CFC7FF45FF09" box="[214,228,225,248]" italics="true" pageId="5" pageNumber="37">p</emphasis>
= 0.0003), more ventral scales (64-73, X = 68.36 vs. 49-77, X = 59.17; 
<emphasis id="BDE9EAA4FFC0FE3DFBB0CFC7FBB0FF09" box="[1027,1041,225,248]" italics="true" pageId="5" pageNumber="37">p</emphasis>
&lt;0.0001), more infradigital lamellae of the third finger (14-18, X = 15 vs. 11-15, X = 13.33; 
<emphasis id="BDE9EAA4FFC0FE3DFC99CE23FC99FEED" box="[810,824,261,284]" italics="true" pageId="5" pageNumber="37">p</emphasis>
= 0.0006), and more infradigital lamellae of the fourth toe (18-23, X = 20.3 vs. 15-20, X = 18.24; 
<emphasis id="BDE9EAA4FFC0FE3DFDC0CE0FFD20FEB1" box="[627,641,297,320]" italics="true" pageId="5" pageNumber="37">p</emphasis>
= 0.0011) (for details see 
<tableCitation id="C21F030DFFC0FE3DFC17CE01FBA3FECE" box="[932,1026,295,319]" captionStart="TABLE 1" captionStartId="3.[151,235,1045,1066]" captionTargetBox="[151,1436,1130,1897]" captionTargetPageId="3" captionText="TABLE 1. Values (in millimeters) of morphometric characters from species of the magellanicus group; sample sizes are shown in parentheses, as well as mean ± SD (min – max); abbreviations for characters are defined in Appendix II." httpUri="http://table.plazi.org/id/DBE2663EFFC6FE3BFF24CB33FB64FBBF" pageId="5" pageNumber="37" tableUuid="DBE2663EFFC6FE3BFF24CB33FB64FBBF">Tables 1</tableCitation>
to 4). 
<taxonomicName id="489D4D35FFC0FE3DFBFACE0EFA9DFEB1" box="[1097,1340,296,320]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC0FE3DFBFACE0EFA9DFEB1" box="[1097,1340,296,320]" italics="true" pageId="5" pageNumber="37">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC0FE3DFAF0CE0EFA3DFEB1" box="[1347,1436,296,320]" pageId="5" pageNumber="37" rank="species">
<emphasis id="BDE9EAA4FFC0FE3DFAF0CE0EFA3DFEB1" bold="true" box="[1347,1436,296,320]" pageId="5" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
has smaller dorsal blotches on the hindlimbs and a more defined vertebral line in comparison with 
<taxonomicName id="489D4D35FFC0FE3DFB5ACE6BFA3DFE95" box="[1257,1436,332,356]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFC0FE3DFB5ACE6BFA3DFE95" box="[1257,1436,332,356]" italics="true" pageId="5" pageNumber="37">L. magellanicus</emphasis>
</taxonomicName>
(
<figureCitation id="17A62A33FFC0FE3DFF2DCE49FF5EFE76" box="[158,255,367,391]" captionStart="FIGURE 5" captionStartId="11.[151,250,1276,1299]" captionTargetBox="[155,1429,411,1252]" captionTargetId="figure@11.[151,1436,409,1255]" captionTargetPageId="11" captionText="FIGURE 5. Morphological comparison between Liolaemus magellanicus (left) and L. caparensis sp. nov. (right). Differences in dorsal scales are shown in blue / light blue, in L. magellanicus scales are more mucronated than in L. caparensis sp. nov.; differences in dorsal scales of hindlimbs are shown in green / light green, in L. magellanicus scales are more mucronated and less carinated than in L. caparensis sp. nov. Notice differences in color pattern on the tails, size of blotches on the limbs, and the differences in vertebral lines." httpUri="https://zenodo.org/record/204785/files/figure.png" pageId="5" pageNumber="37">Figure 5</figureCitation>
), as well as fewer precloacal pores than 
<taxonomicName id="489D4D35FFC0FE3DFD74CE57FCDBFE79" box="[711,890,368,392]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFC0FE3DFD74CE57FCDBFE79" box="[711,890,368,392]" italics="true" pageId="5" pageNumber="37">L. magellanicus</emphasis>
</taxonomicName>
(3-4 vs. 3-5). Dorsal scales of 
<taxonomicName id="489D4D35FFC0FE3DFB65CE56FF6EFE5D" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC0FE3DFB65CE56FF6EFE5D" italics="true" pageId="5" pageNumber="37">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC0FE3DFF65CEB2FE91FE5D" box="[214,304,404,428]" pageId="5" pageNumber="37" rank="species">
<emphasis id="BDE9EAA4FFC0FE3DFF65CEB2FE91FE5D" bold="true" box="[214,304,404,428]" pageId="5" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
have a shorter mucron than those of 
<emphasis id="BDE9EAA4FFC0FE3DFD61CEB3FC2CFE5D" box="[722,909,404,428]" italics="true" pageId="5" pageNumber="37">
<taxonomicName id="489D4D35FFC0FE3DFD61CEB3FC28FE5D" box="[722,905,404,428]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="magellanicus">L. magellanicus</taxonomicName>
.
</emphasis>
The dorsal surface of limb scales is more carinated and less mucronated in 
<taxonomicName id="489D4D35FFC0FE3DFE50CE9FFDD9FE21" box="[483,632,441,464]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC0FE3DFE50CE9FFDD9FE21" box="[483,632,441,464]" italics="true" pageId="5" pageNumber="37">L. caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC0FE3DFD33CE9EFD78FE21" box="[640,729,440,464]" pageId="5" pageNumber="37" rank="species">
<emphasis id="BDE9EAA4FFC0FE3DFD33CE9EFD78FE21" bold="true" box="[640,729,440,464]" pageId="5" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
than in 
<taxonomicName id="489D4D35FFC0FE3DFC87CE9FFC46FE21" box="[820,999,440,464]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFC0FE3DFC87CE9FFC46FE21" box="[820,999,440,464]" italics="true" pageId="5" pageNumber="37">L. magellanicus</emphasis>
</taxonomicName>
(
<figureCitation id="17A62A33FFC0FE3DFC46CE91FBF6FE3E" box="[1013,1111,439,463]" captionStart="FIGURE 5" captionStartId="11.[151,250,1276,1299]" captionTargetBox="[155,1429,411,1252]" captionTargetId="figure@11.[151,1436,409,1255]" captionTargetPageId="11" captionText="FIGURE 5. Morphological comparison between Liolaemus magellanicus (left) and L. caparensis sp. nov. (right). Differences in dorsal scales are shown in blue / light blue, in L. magellanicus scales are more mucronated than in L. caparensis sp. nov.; differences in dorsal scales of hindlimbs are shown in green / light green, in L. magellanicus scales are more mucronated and less carinated than in L. caparensis sp. nov. Notice differences in color pattern on the tails, size of blotches on the limbs, and the differences in vertebral lines." httpUri="https://zenodo.org/record/204785/files/figure.png" pageId="5" pageNumber="37">Figure 5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DBE2663EFFC0FE3DFF24CBB0FC2CFA72" httpUri="https://zenodo.org/record/204781/files/figure.png" pageId="5" pageNumber="37" targetBox="[159,1397,515,1143]" targetPageId="5">
<paragraph id="8F2236B6FFC0FE3DFF24CBB0FC2CFA72" blockId="5.[151,1436,1174,1411]" pageId="5" pageNumber="37">
<emphasis id="BDE9EAA4FFC0FE3DFF24CBB0FEB4FB5D" bold="true" box="[151,277,1174,1197]" pageId="5" pageNumber="37">FIGURE 1.</emphasis>
Relationships between 
<taxonomicName id="489D4D35FFC0FE3DFDB8CBB1FD48FB5C" box="[523,745,1175,1197]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC0FE3DFDB8CBB1FD48FB5C" box="[523,745,1175,1197]" italics="true" pageId="5" pageNumber="37">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC0FE3DFD42CBB1FCE5FB5D" box="[753,836,1175,1196]" pageId="5" pageNumber="37" rank="species">
<emphasis id="BDE9EAA4FFC0FE3DFD42CBB1FCE5FB5D" bold="true" box="[753,836,1175,1196]" pageId="5" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
and species of the 
<taxonomicName id="489D4D35FFC0FE3DFBBDCBBEFB67FB5C" box="[1038,1222,1175,1197]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFC0FE3DFBBDCBBEFB67FB5C" box="[1038,1222,1175,1197]" italics="true" pageId="5" pageNumber="37">L. lineomaculatus</emphasis>
</taxonomicName>
section and selected species of the subgenus 
<taxonomicName id="489D4D35FFC0FE3DFE22CB90FE54FB3D" box="[401,501,1206,1228]" class="Squamata" family="Liolaemidae" genus="Eulaemus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="BDE9EAA4FFC0FE3DFE22CB90FE54FB3D" box="[401,501,1206,1228]" italics="true" pageId="5" pageNumber="37">Eulaemus</emphasis>
</taxonomicName>
and 
<taxonomicName id="489D4D35FFC0FE3DFD9ACB90FD33FB3D" box="[553,658,1206,1228]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="genus" sensu="stricto">
<emphasis id="BDE9EAA4FFC0FE3DFD9ACB90FD33FB3D" box="[553,658,1206,1228]" italics="true" pageId="5" pageNumber="37">Liolaemus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC0FE3DFD29CB90FCB8FB3A" box="[666,793,1205,1227]" pageId="5" pageNumber="37" sensu="stricto">sensu stricto</taxonomicNameLabel>
. On the left, phylogenetic relationships recovered using seven nuclear genes (Cmos, ACM4tg, PRLR, LDA8F, LDA1D, LDA9C and LDA9E, 4180 bp total alignment length) with Bayesian species-tree inference; bold branches represent well supported nodes (
<emphasis id="BDE9EAA4FFC0FE3DFCDCCBD2FC2DFAF8" box="[879,908,1268,1289]" italics="true" pageId="5" pageNumber="37">Pp</emphasis>
&gt; 0.95). On the right, the Bayesian tree (modified from Breitman 
<emphasis id="BDE9EAA4FFC0FE3DFE81CA35FEC4FAD9" box="[306,357,1298,1320]" italics="true" pageId="5" pageNumber="37">et al.</emphasis>
2011), represents a concatenated analyses (5865 bp) including the seven nuclear genes plus two mitochondrial genes (12S and cyt- 
<emphasis id="BDE9EAA4FFC0FE3DFE21CA16FE3FFAB7" box="[402,414,1328,1350]" italics="true" pageId="5" pageNumber="37">b</emphasis>
), and summarizes information from MP and ML methods. Nodes with high support from three methods (MP, jackknife and ML bootstrap&gt;0.70; Bayesian 
<emphasis id="BDE9EAA4FFC0FE3DFD0DCA76FD7BFA94" box="[702,730,1360,1381]" italics="true" pageId="5" pageNumber="37">Pp</emphasis>
&gt; 0.95) are identified by bold branches; solid circles show nodes with weak MP support, and open circles nodes with weak MP and ML support.
</paragraph>
</caption>
<subSubSection id="C787653DFFC0FE30FF76CA8DFBABFE94" lastPageId="8" lastPageNumber="40" pageId="5" pageNumber="37" type="description">
<paragraph id="8F2236B6FFC0FE3DFF76CA8DFADDF9A2" blockId="5.[151,1437,1451,2015]" pageId="5" pageNumber="37">
<emphasis id="BDE9EAA4FFC0FE3DFF76CA8DFE43FA35" bold="true" box="[197,482,1451,1476]" pageId="5" pageNumber="37">
Description of 
<typeStatus id="50268814FFC0FE3DFEC7CA8AFE7CFA35" box="[372,477,1452,1476]" pageId="5" pageNumber="37" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male. Snout-vent length 59.0 mm. Tail length (complete, not regenerated) 84.0 mm. Radius-ulna length 
<quantity id="48659B53FFC0FE3DFE1DCAE9FDB7FA16" box="[430,534,1487,1511]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="5" pageNumber="37" unit="mm" value="25.9">25.9 mm</quantity>
. Head length 
<quantity id="48659B53FFC0FE3DFD01CAE9FCB6FA16" box="[690,791,1487,1511]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="5" pageNumber="37" unit="mm" value="13.4">13.4 mm</quantity>
(from anterior border of tympanum to tip of snout), 11.0 mm wide (at anterior border of tympanum), 
<quantity id="48659B53FFC0FE3DFD3CCAD2FD47F9FA" box="[655,742,1524,1547]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="5" pageNumber="37" unit="mm" value="7.7">7.7 mm</quantity>
long (at anterior border of tympanum). Snout length 
<quantity id="48659B53FFC0FE3DFAF6CAD5FA3DF9FA" box="[1349,1436,1523,1547]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="37" unit="mm" value="4.9">4.9 mm</quantity>
(orbit-tip of snout distance). Auditory meatus-eye distance 
<quantity id="48659B53FFC0FE3DFCADC931FCD4F9DE" box="[798,885,1559,1583]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="37" unit="mm" value="4.7">4.7 mm</quantity>
. Interorbital distance 
<quantity id="48659B53FFC0FE3DFBD2C931FB18F9DE" box="[1121,1209,1559,1583]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="37" unit="mm" value="4.2">4.2 mm</quantity>
. Eye-nostril distance 
<quantity id="48659B53FFC0FE3DFF24C91DFF4FF9A2" box="[151,238,1595,1619]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="37" unit="mm" value="3.1">3.1 mm</quantity>
. Forelimb length 
<quantity id="48659B53FFC0FE3DFE1DC91DFDB5F9A2" box="[430,532,1595,1619]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="5" pageNumber="37" unit="mm" value="17.5">17.5 mm</quantity>
. Tibial length 
<quantity id="48659B53FFC0FE3DFD1CC91DFCA6F9A2" box="[687,775,1595,1619]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="5" pageNumber="37" unit="mm" value="8.6">8.6 mm</quantity>
. Foot length 
<quantity id="48659B53FFC0FE3DFC27C91DFC57F9A2" box="[916,1014,1595,1619]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="5" pageNumber="37" unit="mm" value="14.6">14.6 mm</quantity>
(ankle to tip of claw on fourth toe).
</paragraph>
<paragraph id="8F2236B6FFC0FE3EFF76C979FD65FD1E" blockId="5.[151,1437,1451,2015]" lastBlockId="6.[151,1437,151,2012]" lastPageId="6" lastPageNumber="38" pageId="5" pageNumber="37">
Dorsal head scales slightly bulged, smooth, 14 between occiput, at level of anterior border of tympanum, to rostral, pitted with numerous scale organs in the anterior region, and reducing to no organ in the posterior half of the head. Rostral scale wider (
<quantity id="48659B53FFC0FE3DFE5CC981FDE9F94E" box="[495,584,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="37" unit="mm" value="2.8">2.8 mm</quantity>
) than long (
<quantity id="48659B53FFC0FE3DFD60C981FC91F94E" box="[723,816,1703,1727]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="37" unit="mm" value="0.9">0.9 mm</quantity>
). Two postrostrals that together with anterior lorilabial separate nasal scales from rostral, surrounded by six scales. Nasal scales longer than wide, irregularly hexagonal; nostril one-half length of nasal, posterior in position. Scales surrounding nasals 7 on each side. Four internasals. Frontonasals and prefrontals 14, irregular in shape, size and position. Two frontal scales. Frontoparietals in two rows, two anterior and one posterior scale. Interparietal pentagonal (
<quantity id="48659B53FFC0FE3DFC07C811FBB2F8BE" box="[948,1043,1847,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="37" unit="mm" value="1.3">1.3 mm</quantity>
), surrounded by five scales; three smaller and irregular in front and sides, two larger in back. Parietal eye evident. Parietals slightly bulged, irregularly shaped, equal in size to interparietal. Circumorbitals: 9 on the left side, incomplete; 10 on the right side, incomplete. Transversally expanded supraoculars 4–3. Smaller lateral supraoculars: 16–19. Three canthals on the left side, one posterior large, higher than wide, in contact with postnasal, two inferior, small; four canthal on the right side, one posterior with similar characteristics that the left side, and three inferior, small. Loreal scales slightly bulged, 4–5 on each side. Lorilabials longer than wide (7–8), slightly smaller than labials. Superciliaries 7 on each side, flattened and elongated, anterior four broadly overlapping dorsally. Orbit with 13–12 upper and 11–10 lower ciliaries on each side. Orbit diameter 4.7 x 
<quantity id="48659B53FFC3FE3EFD3BCE22FD43FEED" box="[648,738,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="38" unit="mm" value="2.3">2.3 mm</quantity>
. Preocular small, unfragmented, longer than wide. Subocular scale elongated, approximately nine times longer than wide (4.0 x 
<quantity id="48659B53FFC3FE3EFCCACE01FC72FECE" box="[889,979,295,319]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="38" unit="mm" value="0.5">0.5 mm</quantity>
). A well marked longitudinal ridge along upper margin of preocular and subocular scales. Postocular small, slightly bulged, quarter superimposed to subocular, with a marked longitudinal ridge. Palpebral scales, small granular and bulged. Supralabials 6 on each side, convex. Temporals smooth, convex, juxtaposed with one scale organ in the tip. Anterior auriculars smaller than adjacent posterior temporals, slightly projecting outward (1 on each side). Posterior auriculars small and granular. External auditory meatus conspicuous, rounded (1.8 x 
<quantity id="48659B53FFC3FE3EFD4CCEFAFCF7FE05" box="[767,854,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="38" unit="mm" value="1.5">1.5 mm</quantity>
). Lateral scales of neck granular with inflated skin. Mental scale wider (3.0 mm) than high (
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), in contact with four scales. Mental followed posteriorly by two postmentals, and two rows of 2–1 chinshields. Five infralabials on each side, first on each side quadrangular two times wider than supralabials, all others elongated, slightly smaller than supralabials. Gular scales smooth, flat, imbricate, with rounded posterior margins, with melanophores. Scales of throat between chinshields slightly imbricate. Twenty-six gular scales between tympanum openings. Infralabials separated from chinshields by one to two rows of scales. Antehumeral and longitudinal neck folds well developed; gular fold incomplete; postauricular, rictal, dorsolateral and oblique folds unconspicuous.
</paragraph>
<paragraph id="8F2236B6FFC3FE3EFF76CDDAFCCAFC1D" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="38">Scales of dorsal neck region rhomboidal, imbricate, strongly keeled, mucronate. Thirty-nine scales between occiput and anterior surface of thighs. Dorsal body scales rhomboidal, imbricate, strongly keeled and mucronate. Dorsal scales grade laterally into slightly smaller, slightly keeled scales at midbody. Scales immediately anterior and posterior to forelimb and hindlimb insertion small, smooth, granular, and non-overlapping. Body lateral scales grading smaller to larger at midbody. Ventral body scales rhomboidal, smooth, flat, imbricate, and bigger than dorsal scales. Forty-four midbody scales; scales between mental and precloacal pores 71. Scales of cloacal region about equal in size to ventral body scales. Four precloacal pores.</paragraph>
<paragraph id="8F2236B6FFC3FE3EFF76CCD1FBA7FB35" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="38">Anterior suprabrachials rhomboidal, imbricate, smooth, slightly larger in size to dorsal body scales. Postabrachials smaller, smooth, becoming granular near axilla. Supra-antebrachials rhomboidal, imbricate, smooth, some of the external lateral slightly keeled. Infra-antebrachials rhomboidals, imbricate, smooth, toward the hand slightly keeled. Supracarpals imbricated, rhomboidal, smooth. Infracarpals strongly imbricate, rhomboidal, slightly keeled and mucronate. Infradigital lamellae with 3-keels, each terminating in a short mucron, 3-mucronate, numbering: I: 8, II: 12, III: 16, IV: 19, V: 11. Claws robust, curved and sharp, opaque brown.</paragraph>
<paragraph id="8F2236B6FFC3FE3EFF76CBF6FD1EFA15" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="38">Suprafemorals as large as dorsal body scales, rhomboidal, imbricated, strongly keeled and mucronate. Postfemorals small, granular shape. Supratibial rhomboidal, imbricated, strongly keeled and mucronate. Infrafemoral scales small, granular and smooth. Supratarsals rhomboidals, imbricated and smooth, some lateral scales slightly keeled. Infratarsals small, rhomboidal, imbricate, smooth, mucronate, some slightly keeled and mucronate near the digit. Subdigital scales 3-keeled, 3-mucronate, numbering: I: 9, II: 13, III: 19, IV: 21, V: 14. Claws robust, curved and sharp, opaque brown. Tail complete, non-regenerated. Dorsal and lateral caudal scales, rhomboidal in the first half of the tail, becoming quadrangular toward the tip, strongly keeled and mucronate. Ventrals subtriangular and smooth, toward posterior half moderately keeled.</paragraph>
<paragraph id="8F2236B6FFC3FE3EFF76CAC9FD21F8F5" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="38">
<emphasis id="BDE9EAA4FFC3FE3EFF76CAC9FE59F9F9" bold="true" box="[197,504,1519,1544]" pageId="6" pageNumber="38">
Color of 
<typeStatus id="50268814FFC3FE3EFE84CAD6FE3FF9F9" box="[311,414,1520,1544]" pageId="6" pageNumber="38" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Grey dorsal background (
<figureCitation id="17A62A33FFC3FE3EFC9ECAD6FC35F9F9" box="[813,916,1520,1544]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1758]" captionTargetBox="[254,1327,265,1713]" captionTargetId="figure@7.[254,1327,265,1714]" captionTargetPageId="7" captionText="FIGURE 2. Upper, dorsal and ventral views of male holotype in life of Liolaemus caparensis sp. nov. MLP. S 2628; below, dorsal and ventral views of female (LJAMM-CNP 9387)." httpUri="https://zenodo.org/record/204782/files/figure.png" pageId="6" pageNumber="38">Figure 2</figureCitation>
). The posterior borders of the dorsal scales, between neck and pelvic region, present one or two contrasting light colors (white or light blue). Dorsal pattern with eight series of paravertebral and quadrangular black (with a white end) blotches, appearing from the nuchal region to about the first autotomy line, those series fuse into a dark line that is present to the tip of the tail. Dorsolateral region, between the occipital and the pelvic regions, has eight irregular black and white-bordered blotches. Between the series of blotches there are three longitudinal lines, a white vertebral and two light brownish paravertebral lines, the last ones are wider than the vertebral (one and half scale vs. a quarter of scale). The coloration in the vertebral and lateral area is light brown.
</paragraph>
<paragraph id="8F2236B6FFC3FE3EFF76C836FCF1F865" blockId="6.[151,1437,151,2012]" pageId="6" pageNumber="38">Anterior region of the head shows a dark gray background with a central black blotch from the internasals to the frontal scales. Orbitals, postfrontals and parietals are light brown, the black circumorbital scales continue in a line of the same color until the nuchal region. Gray temporal and lateral head regions. Limbs dorsal region are gray with a scarce black reticulation with some lighter color scales.</paragraph>
<paragraph id="8F2236B6FFC3FE3FFF76C886FE3AFF25" blockId="6.[151,1437,151,2012]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="39" pageId="6" pageNumber="38">
Ventral scales with melanophores. Black color is present throughout all the ventral body until the postcloacal region. Dark reticulate area is present in the throat and extends to the adjacent malar and maxilla region. Cloacal and postcloacal regions are black. Ventral area of lower belly and femoral region present a bright yellow color. Tail is light gray (
<figureCitation id="17A62A33FFC2FE3FFE98CF9AFE2FFF25" box="[299,398,188,212]" captionStart="FIGURE 2" captionStartId="7.[151,250,1735,1758]" captionTargetBox="[254,1327,265,1713]" captionTargetId="figure@7.[254,1327,265,1714]" captionTargetPageId="7" captionText="FIGURE 2. Upper, dorsal and ventral views of male holotype in life of Liolaemus caparensis sp. nov. MLP. S 2628; below, dorsal and ventral views of female (LJAMM-CNP 9387)." httpUri="https://zenodo.org/record/204782/files/figure.png" pageId="7" pageNumber="39">Figure 2</figureCitation>
).
</paragraph>
<caption id="DBE2663EFFC2FE3FFF24C9E1FD7FF90C" httpUri="https://zenodo.org/record/204782/files/figure.png" pageId="7" pageNumber="39" targetBox="[254,1327,265,1713]" targetPageId="7">
<paragraph id="8F2236B6FFC2FE3FFF24C9E1FD7FF90C" blockId="7.[151,1436,1735,1789]" pageId="7" pageNumber="39">
<emphasis id="BDE9EAA4FFC2FE3FFF24C9E1FEB5F92C" bold="true" box="[151,276,1735,1758]" pageId="7" pageNumber="39">FIGURE 2.</emphasis>
Upper, dorsal and ventral views of male holotype in life of 
<taxonomicName id="489D4D35FFC2FE3FFC3BC9E1FBC6F92F" box="[904,1127,1735,1758]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC2FE3FFC3BC9E1FBC6F92F" box="[904,1127,1735,1758]" italics="true" pageId="7" pageNumber="39">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC2FE3FFBDCC9EEFB60F92C" box="[1135,1217,1736,1757]" pageId="7" pageNumber="39" rank="species">
<emphasis id="BDE9EAA4FFC2FE3FFBDCC9EEFB60F92C" bold="true" box="[1135,1217,1736,1757]" pageId="7" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
MLP.S 2628; below, dorsal and ventral views of female (LJAMM-CNP 9387).
</paragraph>
</caption>
<paragraph id="8F2236B6FFC2FE3FFF76C80EFB49F85C" blockId="7.[151,1437,1832,2037]" pageId="7" pageNumber="39">
<emphasis id="BDE9EAA4FFC2FE3FFF76C80EFDF2F8B0" bold="true" box="[197,595,1832,1857]" pageId="7" pageNumber="39">
Color of 
<typeStatus id="50268814FFC2FE3FFE9EC80FFE35F8B0" box="[301,404,1833,1857]" pageId="7" pageNumber="39" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
After three years in preservative, the dorsal coloration of the head, dorsum, body flanks and tail becomes darker while maintaining the contrast, but the two light brownish paravertebral lines turned gray. Ventral scales of throat, neck, chest, belly and forelimbs maintain the same dark coloration as in life, and the distinctive yellow ventral coloration of the femoral and belly regions turns gray (
<figureCitation id="17A62A33FFC2FE3FFBCBC8B3FB7BF85C" box="[1144,1242,1941,1965]" captionStart="FIGURE 5" captionStartId="11.[151,250,1276,1299]" captionTargetBox="[155,1429,411,1252]" captionTargetId="figure@11.[151,1436,409,1255]" captionTargetPageId="11" captionText="FIGURE 5. Morphological comparison between Liolaemus magellanicus (left) and L. caparensis sp. nov. (right). Differences in dorsal scales are shown in blue / light blue, in L. magellanicus scales are more mucronated than in L. caparensis sp. nov.; differences in dorsal scales of hindlimbs are shown in green / light green, in L. magellanicus scales are more mucronated and less carinated than in L. caparensis sp. nov. Notice differences in color pattern on the tails, size of blotches on the limbs, and the differences in vertebral lines." httpUri="https://zenodo.org/record/204785/files/figure.png" pageId="7" pageNumber="39">Figure 5</figureCitation>
).
</paragraph>
<paragraph id="8F2236B6FFC2FE30FF76C89EFBABFE94" blockId="7.[151,1437,1832,2037]" lastBlockId="8.[151,1436,151,357]" lastPageId="8" lastPageNumber="40" pageId="7" pageNumber="39">
<emphasis id="BDE9EAA4FFC2FE3FFF76C89EFE98F820" bold="true" box="[197,313,1976,2001]" pageId="7" pageNumber="39">Variation.</emphasis>
Morphological and meristic characters variation, between males and females of 
<taxonomicName id="489D4D35FFC2FE3FFB0EC89FFF14F804" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFC2FE3FFB0EC89FFF14F804" italics="true" pageId="7" pageNumber="39">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFC2FE3FFF0FC8FBFEB4F804" box="[188,277,2013,2037]" pageId="7" pageNumber="39" rank="species">
<emphasis id="BDE9EAA4FFC2FE3FFF0FC8FBFEB4F804" bold="true" box="[188,277,2013,2037]" pageId="7" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
are shown in 
<tableCitation id="C21F030DFFC2FE3FFE07C8FBFDABF804" box="[436,522,2013,2037]" captionStart="TABLE 5" captionStartId="8.[151,235,404,425]" captionTargetBox="[151,1435,483,1664]" captionTargetPageId="8" captionText="TABLE 5. Variation between females and males of Liolaemus caparensis sp. nov. summarized as mean ± SD (min – max) values for all meristic and morphometric (in millimeters) variables." httpUri="http://table.plazi.org/id/DBE2663EFFCDFE30FF24CEB2FC83FE39" pageId="7" pageNumber="39" tableUuid="DBE2663EFFCDFE30FF24CEB2FC83FE39">Table 5</tableCitation>
. Females in life present basically the same dorsal and lateral pattern as present in males, but the coloration has a light brown background; vertebral and paravertebral lines are wider and much more notorious than the ones in males, the lines are white; although, one of the lizards presents yellowish dorsolateral lines. Only the females present a white lateral line from the axilla to the groin. Ventral coloration shows a dark reticulate pattern in throat, melanic chest and belly, but two specimens present a yellowish coloration in the ventrolateral, femoral, cloacal and postcloacal regions. All distinctive dorsal lines, and ventral coloration, femoral and lower belly areas turns in preservative from yellowish or bright yellow to gray.
</paragraph>
</subSubSection>
<caption id="DBE2663EFFCDFE30FF24CEB2FC83FE39" ID-Table-UUID="DBE2663EFFCDFE30FF24CEB2FC83FE39" httpUri="http://table.plazi.org/id/DBE2663EFFCDFE30FF24CEB2FC83FE39" pageId="8" pageNumber="40" targetBox="[151,1435,483,1664]" targetIsTable="true" targetPageId="8">
<paragraph id="8F2236B6FFCDFE30FF24CEB2FC83FE39" blockId="8.[151,1435,403,456]" pageId="8" pageNumber="40">
<emphasis id="BDE9EAA4FFCDFE30FF24CEB2FEA5FE58" bold="true" box="[151,260,404,425]" pageId="8" pageNumber="40">TABLE 5.</emphasis>
Variation between females and males of 
<taxonomicName id="489D4D35FFCDFE30FD1ACEB2FC27FE5B" box="[681,902,404,426]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCDFE30FD1ACEB2FC27FE5B" box="[681,902,404,426]" italics="true" pageId="8" pageNumber="40">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCDFE30FC3ECEB2FC7FFE58" box="[909,990,404,425]" pageId="8" pageNumber="40" rank="species">
<emphasis id="BDE9EAA4FFCDFE30FC3ECEB2FC7FFE58" bold="true" box="[909,990,404,425]" pageId="8" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
summarized as mean ± SD (min–max) values for all meristic and morphometric (in millimeters) variables.
</paragraph>
</caption>
<paragraph id="8F2236B6FFCDFE30FF2ECEC3FBCAF98E" pageId="8" pageNumber="40">
<table id="FD9DC416FFCD01C7FF24CEC5FA3AF971" box="[151,1435,483,1664]" gridcols="3" gridrows="29" pageId="8" pageNumber="40">
<tr id="31AD34F4FFCD01C7FF24CEC5FA3AFE0B" box="[151,1435,483,506]" gridrow="0" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CEC5FF74FE0B" box="[151,213,483,506]" gridcol="0" gridrow="0" pageId="8" pageNumber="40">Var</th>
<th id="727C5D88FFCD01C7FED8CEC5FDD2FE0B" box="[363,627,483,506]" gridcol="1" gridrow="0" pageId="8" pageNumber="40">Females (n=5)</th>
<th id="727C5D88FFCD01C7FC3BCEC5FA3AFE0B" box="[904,1435,483,506]" gridcol="2" gridrow="0" pageId="8" pageNumber="40">Males (n=3)</th>
</tr>
<tr id="31AD34F4FFCD01C7FF24CD2BFA3AFDD2" box="[151,1435,525,547]" gridrow="1" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CD2BFF74FDD2" box="[151,213,525,547]" gridcol="0" gridrow="1" pageId="8" pageNumber="40">SCI</th>
<td id="727C5D88FFCD01C7FED8CD2BFDD2FDD2" box="[363,627,525,547]" gridcol="1" gridrow="1" pageId="8" pageNumber="40">6.75±1.04 (5–8)</td>
<td id="727C5D88FFCD01C7FC3BCD2BFA3AFDD2" box="[904,1435,525,547]" gridcol="2" gridrow="1" pageId="8" pageNumber="40">6±1 (5–7)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CD10FA3AFDBD" box="[151,1435,566,588]" gridrow="2" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CD10FF74FDBD" box="[151,213,566,588]" gridcol="0" gridrow="2" pageId="8" pageNumber="40">LS</th>
<td id="727C5D88FFCD01C7FED8CD10FDD2FDBD" box="[363,627,566,588]" gridcol="1" gridrow="2" pageId="8" pageNumber="40">5±0.53 (4–6)</td>
<td id="727C5D88FFCD01C7FC3BCD10FA3AFDBD" box="[904,1435,566,588]" gridcol="2" gridrow="2" pageId="8" pageNumber="40">4.33±1.15 (3–5)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CD46FA3AFD87" box="[151,1435,608,630]" gridrow="3" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CD46FF74FD87" box="[151,213,608,630]" gridcol="0" gridrow="3" pageId="8" pageNumber="40">SS</th>
<td id="727C5D88FFCD01C7FED8CD46FDD2FD87" box="[363,627,608,630]" gridcol="1" gridrow="3" pageId="8" pageNumber="40">6.88±0.83 (6–8)</td>
<td id="727C5D88FFCD01C7FC3BCD46FA3AFD87" box="[904,1435,608,630]" gridcol="2" gridrow="3" pageId="8" pageNumber="40">6.33±0.58 (6–7)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CDAFFA3AFD6E" box="[151,1435,649,671]" gridrow="4" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CDAFFF74FD6E" box="[151,213,649,671]" gridcol="0" gridrow="4" pageId="8" pageNumber="40">IS</th>
<td id="727C5D88FFCD01C7FED8CDAFFDD2FD6E" box="[363,627,649,671]" gridcol="1" gridrow="4" pageId="8" pageNumber="40">4.75±0.71 (3–5)</td>
<td id="727C5D88FFCD01C7FC3BCDAFFA3AFD6E" box="[904,1435,649,671]" gridcol="2" gridrow="4" pageId="8" pageNumber="40">4.67±0.58 (4–5)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CD94FA3AFD38" box="[151,1435,690,713]" gridrow="5" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CD94FF74FD38" box="[151,213,690,713]" gridcol="0" gridrow="5" pageId="8" pageNumber="40">MS</th>
<td id="727C5D88FFCD01C7FED8CD94FDD2FD38" box="[363,627,690,713]" gridcol="1" gridrow="5" pageId="8" pageNumber="40">44.25±4.95 (39–55)</td>
<td id="727C5D88FFCD01C7FC3BCD94FA3AFD38" box="[904,1435,690,713]" gridcol="2" gridrow="5" pageId="8" pageNumber="40">42.67±2.31 (40–44)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CDFDFA3AFD03" box="[151,1435,731,754]" gridrow="6" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CDFDFF74FD03" box="[151,213,731,754]" gridcol="0" gridrow="6" pageId="8" pageNumber="40">DS</th>
<td id="727C5D88FFCD01C7FED8CDFDFDD2FD03" box="[363,627,731,754]" gridcol="1" gridrow="6" pageId="8" pageNumber="40">42.13±2.59 (39–47)</td>
<td id="727C5D88FFCD01C7FC3BCDFDFA3AFD03" box="[904,1435,731,754]" gridcol="2" gridrow="6" pageId="8" pageNumber="40">39.67±1.15 (39–41)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CC23FA3AFCEA" box="[151,1435,773,795]" gridrow="7" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CC23FF74FCEA" box="[151,213,773,795]" gridcol="0" gridrow="7" pageId="8" pageNumber="40">VS</th>
<td id="727C5D88FFCD01C7FED8CC23FDD2FCEA" box="[363,627,773,795]" gridcol="1" gridrow="7" pageId="8" pageNumber="40">68.75±3.01 (64–73)</td>
<td id="727C5D88FFCD01C7FC3BCC23FA3AFCEA" box="[904,1435,773,795]" gridcol="2" gridrow="7" pageId="8" pageNumber="40">67.33±3.21 (65–71)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CC08FA3AFCB4" box="[151,1435,814,837]" gridrow="8" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CC08FF74FCB4" box="[151,213,814,837]" gridcol="0" gridrow="8" pageId="8" pageNumber="40">IL3</th>
<td id="727C5D88FFCD01C7FED8CC08FDD2FCB4" box="[363,627,814,837]" gridcol="1" gridrow="8" pageId="8" pageNumber="40">14.88±1.36 (14–18)</td>
<td id="727C5D88FFCD01C7FC3BCC08FA3AFCB4" box="[904,1435,814,837]" gridcol="2" gridrow="8" pageId="8" pageNumber="40">15.33±0.58 (15–16)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CC7EFA3AFC9F" box="[151,1435,856,878]" gridrow="9" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CC7EFF74FC9F" box="[151,213,856,878]" gridcol="0" gridrow="9" pageId="8" pageNumber="40">IL4</th>
<td id="727C5D88FFCD01C7FED8CC7EFDD2FC9F" box="[363,627,856,878]" gridcol="1" gridrow="9" pageId="8" pageNumber="40">20.25±1.67 (18–23)</td>
<td id="727C5D88FFCD01C7FC3BCC7EFA3AFC9F" box="[904,1435,856,878]" gridcol="2" gridrow="9" pageId="8" pageNumber="40">20.5±0.71 (20–21)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CCA7FA3AFC69" box="[151,1435,897,920]" gridrow="10" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CCA7FF74FC69" box="[151,213,897,920]" gridcol="0" gridrow="10" pageId="8" pageNumber="40">Pores</th>
<td id="727C5D88FFCD01C7FED8CCA7FDD2FC69" box="[363,627,897,920]" gridcol="1" gridrow="10" pageId="8" pageNumber="40">0</td>
<td id="727C5D88FFCD01C7FC3BCCA7FA3AFC69" box="[904,1435,897,920]" gridcol="2" gridrow="10" pageId="8" pageNumber="40">3.67±0.58 (3–4)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CC8CFA3AFC31" box="[151,1435,938,960]" gridrow="11" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CC8CFF74FC31" box="[151,213,938,960]" gridcol="0" gridrow="11" pageId="8" pageNumber="40">SVL</th>
<td id="727C5D88FFCD01C7FED8CC8CFDD2FC31" box="[363,627,938,960]" gridcol="1" gridrow="11" pageId="8" pageNumber="40">49.8±2.77 (47–54)</td>
<td id="727C5D88FFCD01C7FC3BCC8CFA3AFC31" box="[904,1435,938,960]" gridcol="2" gridrow="11" pageId="8" pageNumber="40">55±3.61 (52–59)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CCF5FA3AFC1B" box="[151,1435,979,1002]" gridrow="12" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CCF5FF74FC1B" box="[151,213,979,1002]" gridcol="0" gridrow="12" pageId="8" pageNumber="40">TL</th>
<td id="727C5D88FFCD01C7FED8CCF5FDD2FC1B" box="[363,627,979,1002]" gridcol="1" gridrow="12" pageId="8" pageNumber="40">63±2.94 (60–67)</td>
<td id="727C5D88FFCD01C7FC3BCCF5FA3AFC1B" box="[904,1435,979,1002]" gridcol="2" gridrow="12" pageId="8" pageNumber="40">74±8.72 (68–84)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CCDBFA3AFBE2" box="[151,1435,1021,1043]" gridrow="13" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CCDBFF74FBE2" box="[151,213,1021,1043]" gridcol="0" gridrow="13" pageId="8" pageNumber="40">DFH</th>
<td id="727C5D88FFCD01C7FED8CCDBFDD2FBE2" box="[363,627,1021,1043]" gridcol="1" gridrow="13" pageId="8" pageNumber="40">23.84±1.62 (21.2–25.2)</td>
<td id="727C5D88FFCD01C7FC3BCCDBFA3AFBE2" box="[904,1435,1021,1043]" gridcol="2" gridrow="13" pageId="8" pageNumber="40">24.87±0.93 (24.1–25.9)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CB00FA3AFBCC" box="[151,1435,1062,1085]" gridrow="14" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CB00FF74FBCC" box="[151,213,1062,1085]" gridcol="0" gridrow="14" pageId="8" pageNumber="40">FOL</th>
<td id="727C5D88FFCD01C7FED8CB00FDD2FBCC" box="[363,627,1062,1085]" gridcol="1" gridrow="14" pageId="8" pageNumber="40">12.52±0.71 (11.37–13.19)</td>
<td id="727C5D88FFCD01C7FC3BCB00FA3AFBCC" box="[904,1435,1062,1085]" gridcol="2" gridrow="14" pageId="8" pageNumber="40">14.99±0.44 (14.68–15.3)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CB76FA3AFB97" box="[151,1435,1104,1126]" gridrow="15" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CB76FF74FB97" box="[151,213,1104,1126]" gridcol="0" gridrow="15" pageId="8" pageNumber="40">TFL</th>
<td id="727C5D88FFCD01C7FED8CB76FDD2FB97" box="[363,627,1104,1126]" gridcol="1" gridrow="15" pageId="8" pageNumber="40">6.86±0.55 (6.19–7.55)</td>
<td id="727C5D88FFCD01C7FC3BCB76FA3AFB97" box="[904,1435,1104,1126]" gridcol="2" gridrow="15" pageId="8" pageNumber="40">7.86±0.72 (7.19–8.62)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CB5FFA3AFB7E" box="[151,1435,1145,1167]" gridrow="16" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CB5FFF74FB7E" box="[151,213,1145,1167]" gridcol="0" gridrow="16" pageId="8" pageNumber="40">RUL</th>
<td id="727C5D88FFCD01C7FED8CB5FFDD2FB7E" box="[363,627,1145,1167]" gridcol="1" gridrow="16" pageId="8" pageNumber="40">5.08±0.44 (4.58–5.66)</td>
<td id="727C5D88FFCD01C7FC3BCB5FFA3AFB7E" box="[904,1435,1145,1167]" gridcol="2" gridrow="16" pageId="8" pageNumber="40">5.51±0.12 (5.38–5.61)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CB84FA3AFB49" box="[151,1435,1186,1208]" gridrow="17" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CB84FF74FB49" box="[151,213,1186,1208]" gridcol="0" gridrow="17" pageId="8" pageNumber="40">HAL</th>
<td id="727C5D88FFCD01C7FED8CB84FDD2FB49" box="[363,627,1186,1208]" gridcol="1" gridrow="17" pageId="8" pageNumber="40">7.55±0.53 (7.08–8.36)</td>
<td id="727C5D88FFCD01C7FC3BCB84FA3AFB49" box="[904,1435,1186,1208]" gridcol="2" gridrow="17" pageId="8" pageNumber="40">8.53±0.59 (7.85–8.91)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CBEDFA3AFB13" box="[151,1435,1227,1250]" gridrow="18" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CBEDFF74FB13" box="[151,213,1227,1250]" gridcol="0" gridrow="18" pageId="8" pageNumber="40">HH</th>
<td id="727C5D88FFCD01C7FED8CBEDFDD2FB13" box="[363,627,1227,1250]" gridcol="1" gridrow="18" pageId="8" pageNumber="40">6.16±0.41 (5.65–6.7)</td>
<td id="727C5D88FFCD01C7FC3BCBEDFA3AFB13" box="[904,1435,1227,1250]" gridcol="2" gridrow="18" pageId="8" pageNumber="40">7.05±0.56 (6.72–7.7)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CBD0FA3AFAFD" box="[151,1435,1270,1292]" gridrow="19" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CBD0FF74FAFD" box="[151,213,1270,1292]" gridcol="0" gridrow="19" pageId="8" pageNumber="40">HW</th>
<td id="727C5D88FFCD01C7FED8CBD0FDD2FAFD" box="[363,627,1270,1292]" gridcol="1" gridrow="19" pageId="8" pageNumber="40">8.8±0.43 (8.4–9.33)</td>
<td id="727C5D88FFCD01C7FC3BCBD0FA3AFAFD" box="[904,1435,1270,1292]" gridcol="2" gridrow="19" pageId="8" pageNumber="40">10.41±0.91 (9.37–11.07)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CA38FA3AFAC4" box="[151,1435,1310,1333]" gridrow="20" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CA38FF74FAC4" box="[151,213,1310,1333]" gridcol="0" gridrow="20" pageId="8" pageNumber="40">HL</th>
<td id="727C5D88FFCD01C7FED8CA38FDD2FAC4" box="[363,627,1310,1333]" gridcol="1" gridrow="20" pageId="8" pageNumber="40">10.95±0.37 (10.43–11.4)</td>
<td id="727C5D88FFCD01C7FC3BCA38FA3AFAC4" box="[904,1435,1310,1333]" gridcol="2" gridrow="20" pageId="8" pageNumber="40">12.38±0.91 (11.69–13.41)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CA6EFA3AFAAF" box="[151,1435,1352,1374]" gridrow="21" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CA6EFF74FAAF" box="[151,213,1352,1374]" gridcol="0" gridrow="21" pageId="8" pageNumber="40">EH</th>
<td id="727C5D88FFCD01C7FED8CA6EFDD2FAAF" box="[363,627,1352,1374]" gridcol="1" gridrow="21" pageId="8" pageNumber="40">1.79±0.14 (1.58–1.89)</td>
<td id="727C5D88FFCD01C7FC3BCA6EFA3AFAAF" box="[904,1435,1352,1374]" gridcol="2" gridrow="21" pageId="8" pageNumber="40">1.98±0.27 (1.77–2.29)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CA57FA3AFA79" box="[151,1435,1393,1416]" gridrow="22" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CA57FF74FA79" box="[151,213,1393,1416]" gridcol="0" gridrow="22" pageId="8" pageNumber="40">EL</th>
<td id="727C5D88FFCD01C7FED8CA57FDD2FA79" box="[363,627,1393,1416]" gridcol="1" gridrow="22" pageId="8" pageNumber="40">2.99±0.1 (2.85–3.07)</td>
<td id="727C5D88FFCD01C7FC3BCA57FA3AFA79" box="[904,1435,1393,1416]" gridcol="2" gridrow="22" pageId="8" pageNumber="40">3.16±0.24 (2.91–3.38)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CABCFA3AFA41" box="[151,1435,1434,1456]" gridrow="23" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CABCFF74FA41" box="[151,213,1434,1456]" gridcol="0" gridrow="23" pageId="8" pageNumber="40">RND</th>
<td id="727C5D88FFCD01C7FED8CABCFDD2FA41" box="[363,627,1434,1456]" gridcol="1" gridrow="23" pageId="8" pageNumber="40">1.83±0.22 (1.58–2.15)</td>
<td id="727C5D88FFCD01C7FC3BCABCFA3AFA41" box="[904,1435,1434,1456]" gridcol="2" gridrow="23" pageId="8" pageNumber="40">2.16±0.13 (2.01–2.26)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CAE5FA3AFA2B" box="[151,1435,1475,1498]" gridrow="24" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CAE5FF74FA2B" box="[151,213,1475,1498]" gridcol="0" gridrow="24" pageId="8" pageNumber="40">RH</th>
<td id="727C5D88FFCD01C7FED8CAE5FDD2FA2B" box="[363,627,1475,1498]" gridcol="1" gridrow="24" pageId="8" pageNumber="40">0.97±0.05 (0.9–1.01)</td>
<td id="727C5D88FFCD01C7FC3BCAE5FA3AFA2B" box="[904,1435,1475,1498]" gridcol="2" gridrow="24" pageId="8" pageNumber="40">1±0.1 (0.94–1.12)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24CACBFA3AF9F5" box="[151,1435,1517,1540]" gridrow="25" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24CACBFF74F9F5" box="[151,213,1517,1540]" gridcol="0" gridrow="25" pageId="8" pageNumber="40">RL</th>
<td id="727C5D88FFCD01C7FED8CACBFDD2F9F5" box="[363,627,1517,1540]" gridcol="1" gridrow="25" pageId="8" pageNumber="40">2.41±0.16 (2.26–2.66)</td>
<td id="727C5D88FFCD01C7FC3BCACBFA3AF9F5" box="[904,1435,1517,1540]" gridcol="2" gridrow="25" pageId="8" pageNumber="40">2.83±0.11 (2.71–2.93)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24C930FA3AF9DC" box="[151,1435,1558,1581]" gridrow="26" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24C930FF74F9DC" box="[151,213,1558,1581]" gridcol="0" gridrow="26" pageId="8" pageNumber="40">DRE</th>
<td id="727C5D88FFCD01C7FED8C930FDD2F9DC" box="[363,627,1558,1581]" gridcol="1" gridrow="26" pageId="8" pageNumber="40">4.65±0.25 (4.38–4.99)</td>
<td id="727C5D88FFCD01C7FC3BC930FA3AF9DC" box="[904,1435,1558,1581]" gridcol="2" gridrow="26" pageId="8" pageNumber="40">5.23±0.22 (5.09–5.49)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24C966FA3AF9A7" box="[151,1435,1600,1622]" gridrow="27" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24C966FF74F9A7" box="[151,213,1600,1622]" gridcol="0" gridrow="27" pageId="8" pageNumber="40">AH</th>
<td id="727C5D88FFCD01C7FED8C966FDD2F9A7" box="[363,627,1600,1622]" gridcol="1" gridrow="27" pageId="8" pageNumber="40">1.82±0.18 (1.65–2.09)</td>
<td id="727C5D88FFCD01C7FC3BC966FA3AF9A7" box="[904,1435,1600,1622]" gridcol="2" gridrow="27" pageId="8" pageNumber="40">2.13±0.17 (1.94–2.24)</td>
</tr>
<tr id="31AD34F4FFCD01C7FF24C94FFA3AF971" box="[151,1435,1641,1664]" gridrow="28" pageId="8" pageNumber="40">
<th id="727C5D88FFCD01C7FF24C94FFF74F971" box="[151,213,1641,1664]" gridcol="0" gridrow="28" pageId="8" pageNumber="40">AL</th>
<td id="727C5D88FFCD01C7FED8C94FFDD2F971" box="[363,627,1641,1664]" gridcol="1" gridrow="28" pageId="8" pageNumber="40">1.58±0.25 (1.33–1.88)</td>
<td id="727C5D88FFCD01C7FC3BC94FFA3AF971" box="[904,1435,1641,1664]" gridcol="2" gridrow="28" pageId="8" pageNumber="40">1.62±0.03 (1.59–1.64)</td>
</tr>
</table>
</paragraph>
<subSubSection id="C787653DFFCDFE30FF76C9E2FD89F8D5" pageId="8" pageNumber="40" type="etymology">
<paragraph id="8F2236B6FFCDFE30FF76C9E2FD89F8D5" blockId="8.[151,1436,1732,1936]" pageId="8" pageNumber="40">
<emphasis id="BDE9EAA4FFCDFE30FF76C9E2FEEAF92D" bold="true" box="[197,331,1732,1756]" pageId="8" pageNumber="40">Etymology.</emphasis>
The specific epithet of this species “ 
<taxonomicName id="489D4D35FFCDFE30FD4BC9E2FCD0F92D" box="[760,881,1732,1756]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="caparensis">caparensis</taxonomicName>
” refers to the name “Capar” given to the actual Viedma lake by the Aónikenk aborigines that inhabited this area. The Campo Las Piedras plateau is located on the northwestern edge of Viedma Lake.
</paragraph>
</subSubSection>
<subSubSection id="C787653DFFCDFE34FF76C809FCA5FF06" lastPageId="12" lastPageNumber="44" pageId="8" pageNumber="40" type="distribution">
<paragraph id="8F2236B6FFCDFE30FF76C809FE8EF87E" blockId="8.[151,1436,1732,1936]" pageId="8" pageNumber="40">
<emphasis id="BDE9EAA4FFCDFE30FF76C809FEFFF8B9" bold="true" box="[197,350,1839,1864]" pageId="8" pageNumber="40">Distribution.</emphasis>
<taxonomicName id="489D4D35FFCDFE30FED4C816FDFCF8B9" box="[359,605,1840,1864]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="40" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCDFE30FED4C816FDFCF8B9" box="[359,605,1840,1864]" italics="true" pageId="8" pageNumber="40">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCDFE30FDD5C816FD63F8B9" box="[614,706,1840,1864]" pageId="8" pageNumber="40" rank="species">
<emphasis id="BDE9EAA4FFCDFE30FDD5C816FD63F8B9" bold="true" box="[614,706,1840,1864]" pageId="8" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the 
<typeStatus id="50268814FFCDFE30FC50C817FBB2F8B8" box="[995,1043,1841,1865]" pageId="8" pageNumber="40">type</typeStatus>
locality, from Campo Las Piedras plateau, 
<quantity id="48659B53FFCDFE30FF44C873FE90F89D" box="[247,305,1876,1900]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="40" unit="km" value="7.0">7 km</quantity>
N Punta del Lago ranch, Lago 
<collectingCountry id="F78A7626FFCDFE30FD2AC872FCABF89D" box="[665,778,1876,1900]" name="Argentina" pageId="8" pageNumber="40">Argentino</collectingCountry>
department, Santa Cruz province, 
<collectingCountry id="F78A7626FFCDFE30FB26C872FAA9F89D" box="[1173,1288,1876,1900]" name="Argentina" pageId="8" pageNumber="40">Argentina</collectingCountry>
. 
<quantity id="48659B53FFCDFE30FAA7C872FAFDF89D" box="[1300,1372,1876,1900]" metricMagnitude="2" metricUnit="m" metricValue="8.68" pageId="8" pageNumber="40" unit="m" value="868.0">868 m</quantity>
(Figures 3 and 4).
</paragraph>
<caption id="DBE2663EFFCCFE31FF24C889FD2BF815" httpUri="https://zenodo.org/record/204783/files/figure.png" pageId="9" pageNumber="41" targetBox="[210,1374,193,1942]" targetPageId="9">
<paragraph id="8F2236B6FFCCFE31FF24C889FD2BF815" blockId="9.[151,1436,1967,2021]" pageId="9" pageNumber="41">
<emphasis id="BDE9EAA4FFCCFE31FF24C889FEAFF834" bold="true" box="[151,270,1967,1990]" pageId="9" pageNumber="41">FIGURE 3</emphasis>
. Type locality of 
<taxonomicName id="489D4D35FFCCFE31FE76C896FD02F837" box="[453,675,1968,1990]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="41" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCCFE31FE76C896FD02F837" box="[453,675,1968,1990]" italics="true" pageId="9" pageNumber="41">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCCFE31FD19C896FD5AF834" box="[682,763,1968,1989]" pageId="9" pageNumber="41" rank="species">
<emphasis id="BDE9EAA4FFCCFE31FD19C896FD5AF834" bold="true" box="[682,763,1968,1989]" pageId="9" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
, Subantartic Phytogeographic province, Estepa de 
<taxonomicName id="489D4D35FFCCFE31FABAC897FF6FF814" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="9" pageNumber="41" phylum="Tracheophyta" rank="species" species="pallescens">
<emphasis id="BDE9EAA4FFCCFE31FABAC897FF6FF814" italics="true" pageId="9" pageNumber="41">Festuca pallescens</emphasis>
</taxonomicName>
District (49º 34’ 11.0” S, 72º 02’ 51.9” W).
</paragraph>
</caption>
<caption id="DBE2663EFFCFFE32FF24CB76FDDAFB11" httpUri="https://zenodo.org/record/204784/files/figure.png" pageId="10" pageNumber="42" targetBox="[164,1415,205,1066]" targetPageId="10">
<paragraph id="8F2236B6FFCFFE32FF24CB76FDDAFB11" blockId="10.[151,1436,1104,1249]" pageId="10" pageNumber="42">
<emphasis id="BDE9EAA4FFCFFE32FF24CB76FEB2FB94" bold="true" box="[151,275,1104,1127]" pageId="10" pageNumber="42">FIGURE 4.</emphasis>
Distribution map for species of the 
<taxonomicName id="489D4D35FFCFFE32FD35CB76FCACFB97" box="[646,781,1104,1126]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCFFE32FD35CB76FCACFB97" box="[646,781,1104,1126]" italics="true" pageId="10" pageNumber="42">magellanicus</emphasis>
</taxonomicName>
group, with localities sampled superimposed on regional elevation (shading). White square: 
<taxonomicName id="489D4D35FFCFFE32FE7DCB49FD0CFB74" box="[462,685,1135,1157]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCFFE32FE7DCB49FD0CFB74" box="[462,685,1135,1157]" italics="true" pageId="10" pageNumber="42">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCFFE32FD06CB49FCA9FB75" box="[693,776,1135,1156]" pageId="10" pageNumber="42" rank="species">
<emphasis id="BDE9EAA4FFCFFE32FD06CB49FCA9FB75" bold="true" box="[693,776,1135,1156]" pageId="10" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
; black squares: distribution of 
<taxonomicName id="489D4D35FFCFFE32FBFDCB56FB52FB74" box="[1102,1267,1135,1157]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCFFE32FBFDCB56FB52FB74" box="[1102,1267,1135,1157]" italics="true" pageId="10" pageNumber="42">L. magellanicus</emphasis>
</taxonomicName>
; black and gray squares: 
<taxonomicName id="489D4D35FFCFFE32FF43CBA8FE33FB52" box="[240,402,1165,1187]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCFFE32FF43CBA8FE33FB52" box="[240,402,1165,1187]" italics="true" pageId="10" pageNumber="42">L. magellanicus</emphasis>
</taxonomicName>
localities used in this study from LJAMM-CNP collection. Arrow indicates type locality of 
<taxonomicName id="489D4D35FFCFFE32FA88CBA8FF41FB33" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCFFE32FA88CBA8FF41FB33" italics="true" pageId="10" pageNumber="42">L. magellanicus</emphasis>
</taxonomicName>
(in Chile). SC: Santa Cruz province. Notice 
<taxonomicName id="489D4D35FFCFFE32FD15CB8BFC8CFB33" box="[678,813,1197,1218]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCFFE32FD15CB8BFC8CFB33" box="[678,813,1197,1218]" italics="true" pageId="10" pageNumber="42">L. caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCFFE32FC87CB8AFC25FB30" box="[820,900,1196,1217]" pageId="10" pageNumber="42" rank="species">
<emphasis id="BDE9EAA4FFCFFE32FC87CB8AFC25FB30" bold="true" box="[820,900,1196,1217]" pageId="10" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
distribution does not overlap with the distribution of 
<taxonomicName id="489D4D35FFCFFE32FF24CBEAFE98FB10" box="[151,313,1227,1249]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCFFE32FF24CBEAFE98FB10" box="[151,313,1227,1249]" italics="true" pageId="10" pageNumber="42">L. magellanicus</emphasis>
</taxonomicName>
(south of the Santa Cruz river).
</paragraph>
</caption>
<paragraph id="8F2236B6FFCFFE32FF76CA28FEE3F9D0" blockId="10.[151,1437,1293,2038]" pageId="10" pageNumber="42">
<emphasis id="BDE9EAA4FFCFFE32FF76CA28FE24FAD7" bold="true" box="[197,389,1293,1318]" pageId="10" pageNumber="42">Natural history.</emphasis>
The new species is viviparous and probably insectivorous (inferred from the biology of its sister species 
<taxonomicName id="489D4D35FFCFFE32FE82CA17FDE2FAB8" box="[305,579,1329,1353]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCFFE32FE82CA17FDE2FAB8" box="[305,579,1329,1353]" italics="true" pageId="10" pageNumber="42">Liolaemus magellanicus</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EB0C4B47FFCFFE32FDE1CA17FCDEFAB8" author="Ibarguengoytia" box="[594,895,1329,1353]" pageId="10" pageNumber="42" refString="Ibarguengoytia, N. R., Marlin Medina, S., Fernandez, J. B., Gutierrez, J. A., Tappari, F. &amp; Scolaro, A. (2010) Thermal biology of the southernmost lizards in the world: Liolaemus sarmientoi and Liolaemus magellanicus from Patagonia, Argentina. Journal of Thermal Biology, 35, 21 - 27." type="journal article" year="2010">
Ibargüengoytía 
<emphasis id="BDE9EAA4FFCFFE32FCB1CA14FC9AFAB8" box="[770,827,1329,1353]" italics="true" pageId="10" pageNumber="42">et al.</emphasis>
2010
</bibRefCitation>
). Out of five females, only two presented a ventral yellowish coloration and one of those was gravid, thus we inferred that the yellowish coloration in females is not directly related with the reproductive state. The specimens were found in the Subantarctic Phytogeographic province, Estepa de 
<taxonomicName id="489D4D35FFCFFE32FE35CAB9FDFEFA47" box="[390,607,1438,1462]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="10" pageNumber="42" phylum="Tracheophyta" rank="species" species="pallescens">
<emphasis id="BDE9EAA4FFCFFE32FE35CAB9FDFEFA47" box="[390,607,1438,1462]" italics="true" pageId="10" pageNumber="42">Festuca pallescens</emphasis>
</taxonomicName>
district, in an environment mainly characterized as 
<taxonomicName id="489D4D35FFCFFE32FB77CAB9FABCFA47" box="[1220,1309,1439,1462]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="10" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="BDE9EAA4FFCFFE32FB77CAB9FABCFA47" box="[1220,1309,1439,1462]" italics="true" pageId="10" pageNumber="42">Festuca</emphasis>
</taxonomicName>
grasslands (
<bibRefCitation id="EB0C4B47FFCFFE32FF2DCAE7FEBDFA28" author="Roig" box="[158,284,1473,1497]" pageId="10" pageNumber="42" refString="Roig, A. F. (1998) La vegetacion de la Patagonia. In: Correa M. N. (Ed.), Flora Patagonica. Coleccion cientifica del INTA, Tomo VIII, parte I, Buenos Aires, Argentina, pp. 48 - 174." type="book chapter" year="1998">Roig 1998</bibRefCitation>
). Other dominant vegetation included short bushes (
<emphasis id="BDE9EAA4FFCFFE32FCC9CAE7FAC1FA28" box="[890,1376,1473,1497]" italics="true" pageId="10" pageNumber="42">Anartrophyllum desideratum, Senecio spp.</emphasis>
) and grasses (
<taxonomicName id="489D4D35FFCFFE32FF4ACAC0FE93FA0C" box="[249,306,1510,1533]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="10" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="BDE9EAA4FFCFFE32FF4ACAC0FE93FA0C" box="[249,306,1510,1533]" italics="true" pageId="10" pageNumber="42">Stipa</emphasis>
</taxonomicName>
spp.) (
<figureCitation id="17A62A33FFCFFE32FE32CAC3FE45FA0C" box="[385,484,1509,1533]" captionStart="FIGURE 3" captionStartId="9.[151,250,1967,1990]" captionTargetBox="[210,1374,193,1942]" captionTargetId="figure@9.[210,1375,193,1942]" captionTargetPageId="9" captionText="FIGURE 3. Type locality of Liolaemus caparensis sp. nov., Subantartic Phytogeographic province, Estepa de Festuca pallescens District (49 º 34 ’ 11.0 ” S, 72 º 02 ’ 51.9 ” W)." httpUri="https://zenodo.org/record/204783/files/figure.png" pageId="10" pageNumber="42">Figure 3</figureCitation>
). This species was found in open substrates between rocks, sharing the habitat with 
<emphasis id="BDE9EAA4FFCFFE32FF24C92FFEE3F9D0" box="[151,322,1545,1569]" italics="true" pageId="10" pageNumber="42">
<taxonomicName id="489D4D35FFCFFE32FF24C92FFE9FF9D0" box="[151,318,1545,1569]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="tari">Liolaemus tari</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8F2236B6FFCFFE32FF76C90BFC9AF943" blockId="10.[151,1437,1293,2038]" pageId="10" pageNumber="42">
<emphasis id="BDE9EAA4FFCFFE32FF76C90BFE60F9B7" bold="true" box="[197,449,1581,1606]" pageId="10" pageNumber="42">Concluding remarks.</emphasis>
Lizards from the 
<taxonomicName id="489D4D35FFCFFE32FD34C908FCBCF9B7" box="[647,797,1582,1606]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCFFE32FD34C908FCBCF9B7" box="[647,797,1582,1606]" italics="true" pageId="10" pageNumber="42">magellanicus</emphasis>
</taxonomicName>
group have a general phenotype (general body form, coloration, and color pattern) similar to lizards of the 
<taxonomicName id="489D4D35FFCFFE32FD7FC977FCD6F998" box="[716,887,1617,1641]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCFFE32FD7FC977FCD6F998" box="[716,887,1617,1641]" italics="true" pageId="10" pageNumber="42">lineomaculatus</emphasis>
</taxonomicName>
group, but they have precloacal pores, a synapomorphy shared with all other species of the 
<taxonomicName id="489D4D35FFCFFE32FD30C950FC0BF97F" box="[643,938,1654,1678]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCFFE32FD30C950FC0BF97F" box="[643,938,1654,1678]" italics="true" pageId="10" pageNumber="42">Liolaemus lineomaculatus</emphasis>
</taxonomicName>
section (
<emphasis id="BDE9EAA4FFCFFE32FBA2C950FB7DF97F" box="[1041,1244,1654,1678]" italics="true" pageId="10" pageNumber="42">
<taxonomicName id="489D4D35FFCFFE32FBA2C950FBF1F97F" box="[1041,1104,1654,1678]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="kingii">kingii</taxonomicName>
+ 
<taxonomicName id="489D4D35FFCFFE32FBEDC950FB7DF97F" box="[1118,1244,1654,1678]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="archeforus">archeforus</taxonomicName>
</emphasis>
and 
<taxonomicName id="489D4D35FFCFFE32FAA0C951FA3DF97F" box="[1299,1436,1655,1678]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="somuncurae">
<emphasis id="BDE9EAA4FFCFFE32FAA0C951FA3DF97F" box="[1299,1436,1655,1678]" italics="true" pageId="10" pageNumber="42">somuncurae</emphasis>
</taxonomicName>
groups) but not with species from the 
<taxonomicName id="489D4D35FFCFFE32FDF3C9BFFD4AF940" box="[576,747,1689,1713]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCFFE32FDF3C9BFFD4AF940" box="[576,747,1689,1713]" italics="true" pageId="10" pageNumber="42">lineomaculatus</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8F2236B6FFCFFE32FF76C998FF71F894" blockId="10.[151,1437,1293,2038]" pageId="10" pageNumber="42">
<bibRefCitation id="EB0C4B47FFCFFE32FF76C998FE15F924" box="[197,436,1725,1750]" pageId="10" pageNumber="42" refString="Fouquet, A., Gilles, A., Vences, M., Marty, C., Blanc, M. &amp; Gemmell, N. J. (2007) Underestimation of species richness in neotropical frogs revealed by mtDNA analyses. PLoS ONE, 2, e 1109." type="journal article">
Fouquet 
<emphasis id="BDE9EAA4FFCFFE32FE9BC999FEC1F927" box="[296,352,1726,1750]" italics="true" pageId="10" pageNumber="42">et al.</emphasis>
(2007)
</bibRefCitation>
, proposed for Neotropical frogs a mtDNA approach to species delimitation based on isolation-by-distance population structure. The method tests for correlation of geographic distance with genetic distance, which characterized most samples up to uncorrelated values of 3%. At this value the isolation-by-distance correlation was not significant and 
<bibRefCitation id="EB0C4B47FFCFFE32FD98C80CFCBFF8B0" box="[555,798,1833,1857]" pageId="10" pageNumber="42" refString="Fouquet, A., Gilles, A., Vences, M., Marty, C., Blanc, M. &amp; Gemmell, N. J. (2007) Underestimation of species richness in neotropical frogs revealed by mtDNA analyses. PLoS ONE, 2, e 1109." type="journal article">
Fouquet 
<emphasis id="BDE9EAA4FFCFFE32FD23C80CFD6AF8B0" box="[656,715,1833,1857]" italics="true" pageId="10" pageNumber="42">et al.</emphasis>
(2007)
</bibRefCitation>
interpreted this break as the limit to intra-specific gene flow.
</paragraph>
<paragraph id="8F2236B6FFCFFE33FF76C857FC24FE95" blockId="10.[151,1437,1293,2038]" lastBlockId="11.[151,1436,151,356]" lastPageId="11" lastPageNumber="43" pageId="10" pageNumber="42">
Martinez (
<emphasis id="BDE9EAA4FFCFFE32FE8DC857FDF6F878" box="[318,599,1905,1929]" italics="true" pageId="10" pageNumber="42">personal communication</emphasis>
) found that within 
<taxonomicName id="489D4D35FFCFFE32FCF2C857FC14F878" box="[833,949,1905,1929]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="BDE9EAA4FFCFFE32FCF2C857FC14F878" box="[833,949,1905,1929]" italics="true" pageId="10" pageNumber="42">Liolaemus</emphasis>
</taxonomicName>
the mean genetic distance value of cytochrome 
<emphasis id="BDE9EAA4FFCFFE32FF47C8B0FEA3F85F" box="[244,258,1942,1966]" italics="true" pageId="10" pageNumber="42">b</emphasis>
between sister species was ~ 4%; moreover, she found that different groups of 
<taxonomicName id="489D4D35FFCFFE32FB17C8B0FAB9F85F" box="[1188,1304,1942,1966]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="BDE9EAA4FFCFFE32FB17C8B0FAB9F85F" box="[1188,1304,1942,1966]" italics="true" pageId="10" pageNumber="42">Liolaemus</emphasis>
</taxonomicName>
had values ranging from 1 to 6%. In the 
<taxonomicName id="489D4D35FFCFFE32FE6AC89DFD00F823" box="[473,673,1978,2002]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCFFE32FE6AC89DFD00F823" box="[473,673,1978,2002]" italics="true" pageId="10" pageNumber="42">L. lineomaculatus</emphasis>
</taxonomicName>
section, average genetic distance between described species is 1.6% (based on the sister species 
<taxonomicName id="489D4D35FFCFFE32FE7DC8F9FDE4F807" box="[462,581,2014,2038]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="BDE9EAA4FFCFFE32FE7DC8F9FDE4F807" box="[462,581,2014,2038]" italics="true" pageId="10" pageNumber="42">L. kolengh</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="489D4D35FFCFFE32FDC0C8F9FD4CF807" box="[627,749,2014,2038]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="hatcheri">
<emphasis id="BDE9EAA4FFCFFE32FDC0C8F9FD4CF807" box="[627,749,2014,2038]" italics="true" pageId="10" pageNumber="42">L. hatcheri</emphasis>
</taxonomicName>
), and 2.25% between species of the 
<emphasis id="BDE9EAA4FFCFFE32FB3BC8F8FAF5F807" box="[1160,1364,2014,2038]" italics="true" pageId="10" pageNumber="42">
<taxonomicName id="489D4D35FFCFFE32FB3BC8F8FB66F807" box="[1160,1223,2014,2038]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="kingii">kingii</taxonomicName>
+ 
<taxonomicName id="489D4D35FFCFFE32FB65C8F8FAF5F807" box="[1238,1364,2014,2038]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="42" phylum="Chordata" rank="species" species="archeforus">archeforus</taxonomicName>
</emphasis>
group (based on sister species 
<taxonomicName id="489D4D35FFCEFE33FE00CFBFFDEDFF41" box="[435,588,152,176]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="archeforus">
<emphasis id="BDE9EAA4FFCEFE33FE00CFBFFDEDFF41" box="[435,588,152,176]" italics="true" pageId="11" pageNumber="43">L. archeforus</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="489D4D35FFCEFE33FD30CFBFFCEEFF41" box="[643,847,152,176]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="chacabucoense">
<emphasis id="BDE9EAA4FFCEFE33FD30CFBFFCEEFF41" box="[643,847,152,176]" italics="true" pageId="11" pageNumber="43">L. chacabucoense</emphasis>
</taxonomicName>
and 
<taxonomicName id="489D4D35FFCEFE33FC3ECFBFFBE3FF41" box="[909,1090,152,176]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="escarchadosi">
<emphasis id="BDE9EAA4FFCEFE33FC3ECFBFFBE3FF41" box="[909,1090,152,176]" italics="true" pageId="11" pageNumber="43">L. escarchadosi</emphasis>
</taxonomicName>
vs. 
<taxonomicName id="489D4D35FFCEFE33FBC4CFBFFB60FF41" box="[1143,1217,153,176]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="tari">
<emphasis id="BDE9EAA4FFCEFE33FBC4CFBFFB60FF41" box="[1143,1217,153,176]" italics="true" pageId="11" pageNumber="43">L. tari</emphasis>
</taxonomicName>
). Even though the genetic distance values calculated for 
<taxonomicName id="489D4D35FFCEFE33FD8ECF9BFD72FF25" box="[573,723,189,212]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCEFE33FD8ECF9BFD72FF25" box="[573,723,189,212]" italics="true" pageId="11" pageNumber="43">L. caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCEFE33FD6ACF9AFC93FF25" box="[729,818,188,212]" pageId="11" pageNumber="43" rank="species">
<emphasis id="BDE9EAA4FFCEFE33FD6ACF9AFC93FF25" bold="true" box="[729,818,188,212]" pageId="11" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
relative to 
<taxonomicName id="489D4D35FFCEFE33FC1CCF9BFBC0FF25" box="[943,1121,188,212]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FC1CCF9BFBC0FF25" box="[943,1121,188,212]" italics="true" pageId="11" pageNumber="43">L. magellanicus</emphasis>
</taxonomicName>
are slightly higher than values of other species from the 
<taxonomicName id="489D4D35FFCEFE33FE51CFC7FD0AFF09" box="[482,683,224,248]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCEFE33FE51CFC7FD0AFF09" box="[482,683,224,248]" italics="true" pageId="11" pageNumber="43">L. lineomaculatus</emphasis>
</taxonomicName>
section, we want to highlight that the discovery of the new species started with the genetics analyses presented by 
<bibRefCitation id="EB0C4B47FFCEFE33FD1ECE22FC08FEED" box="[685,937,260,284]" pageId="11" pageNumber="43" refString="Breitman, M. F., Avila, L. J., Sites, J. W., Jr. &amp; Morando, M. (2011) Lizards from the end of the world: Phylogenetic relationships of the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemini). Molecular Phylogenetics and Evolution, 59, 364 - 376." type="journal article">
Breitman 
<emphasis id="BDE9EAA4FFCEFE33FCAECE23FCF7FEED" box="[797,854,260,284]" italics="true" pageId="11" pageNumber="43">et al.</emphasis>
(2011)
</bibRefCitation>
, in which several clades were recognized as candidate species, continued with the nuclear data here presented and concluded with the morphological differentiation and characterization of 
<taxonomicName id="489D4D35FFCEFE33FE6ECE6BFDD2FE95" box="[477,627,333,356]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCEFE33FE6ECE6BFDD2FE95" box="[477,627,333,356]" italics="true" pageId="11" pageNumber="43">L. caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCEFE33FDC9CE6AFD75FE95" box="[634,724,332,356]" pageId="11" pageNumber="43" rank="species">
<emphasis id="BDE9EAA4FFCEFE33FDC9CE6AFD75FE95" bold="true" box="[634,724,332,356]" pageId="11" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
presented here.
</paragraph>
<caption id="DBE2663EFFCEFE33FF24CBDAFE02FA7C" httpUri="https://zenodo.org/record/204785/files/figure.png" pageId="11" pageNumber="43" targetBox="[155,1429,411,1252]" targetPageId="11">
<paragraph id="8F2236B6FFCEFE33FF24CBDAFE02FA7C" blockId="11.[151,1436,1276,1421]" pageId="11" pageNumber="43">
<emphasis id="BDE9EAA4FFCEFE33FF24CBDAFEB2FAE3" bold="true" box="[151,275,1276,1299]" pageId="11" pageNumber="43">FIGURE 5.</emphasis>
Morphological comparison between 
<taxonomicName id="489D4D35FFCEFE33FD3CCBDBFC27FAE2" box="[655,902,1277,1299]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FD3CCBDBFC27FAE2" box="[655,902,1277,1299]" italics="true" pageId="11" pageNumber="43">Liolaemus magellanicus</emphasis>
</taxonomicName>
(left) and 
<taxonomicName id="489D4D35FFCEFE33FC41CBD8FBDBFAE2" box="[1010,1146,1278,1299]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCEFE33FC41CBD8FBDBFAE2" box="[1010,1146,1278,1299]" italics="true" pageId="11" pageNumber="43">L. caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCEFE33FB33CBDBFB70FAE3" box="[1152,1233,1277,1298]" pageId="11" pageNumber="43" rank="species">
<emphasis id="BDE9EAA4FFCEFE33FB33CBDBFB70FAE3" bold="true" box="[1152,1233,1277,1298]" pageId="11" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
(right). Differences in dorsal scales are shown in blue/light blue, in 
<taxonomicName id="489D4D35FFCEFE33FDCBCA3AFCBBFAC0" box="[632,794,1307,1329]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FDCBCA3AFCBBFAC0" box="[632,794,1307,1329]" italics="true" pageId="11" pageNumber="43">L. magellanicus</emphasis>
</taxonomicName>
scales are more mucronated than in 
<taxonomicName id="489D4D35FFCEFE33FB3FCA3AFAB2FAC0" box="[1164,1299,1308,1329]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCEFE33FB3FCA3AFAB2FAC0" box="[1164,1299,1308,1329]" italics="true" pageId="11" pageNumber="43">L. caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCEFE33FAAACA3AFAC8FAC0" box="[1305,1385,1308,1329]" pageId="11" pageNumber="43" rank="species">
<emphasis id="BDE9EAA4FFCEFE33FAAACA3AFAC8FAC0" bold="true" box="[1305,1385,1308,1329]" pageId="11" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
; differences in dorsal scales of hindlimbs are shown in green/light green, in 
<taxonomicName id="489D4D35FFCEFE33FCCECA1DFB81FAA1" box="[893,1056,1338,1360]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FCCECA1DFB81FAA1" box="[893,1056,1338,1360]" italics="true" pageId="11" pageNumber="43">L. magellanicus</emphasis>
</taxonomicName>
scales are more mucronated and less carinated than in 
<taxonomicName id="489D4D35FFCEFE33FEF7CA7CFE6AFA9E" box="[324,459,1370,1391]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCEFE33FEF7CA7CFE6AFA9E" box="[324,459,1370,1391]" italics="true" pageId="11" pageNumber="43">L. caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCEFE33FE63CA7FFD80FA9F" box="[464,545,1369,1390]" pageId="11" pageNumber="43" rank="species">
<emphasis id="BDE9EAA4FFCEFE33FE63CA7FFD80FA9F" bold="true" box="[464,545,1369,1390]" pageId="11" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
Notice differences in color pattern on the tails, size of blotches on the limbs, and the differences in vertebral lines.
</paragraph>
</caption>
<paragraph id="8F2236B6FFCEFE33FF76CA9DFCB6F977" blockId="11.[151,1437,1466,2031]" pageId="11" pageNumber="43">
Even though there is not a formal description of the morphological characteristics that define the 
<taxonomicName id="489D4D35FFCEFE33FAB5CA9CFA3AFA23" box="[1286,1435,1466,1490]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FAB5CA9CFA3AFA23" box="[1286,1435,1466,1490]" italics="true" pageId="11" pageNumber="43">magellanicus</emphasis>
</taxonomicName>
group, the features that defined the species 
<taxonomicName id="489D4D35FFCEFE33FDCDCAF8FC2EFA07" box="[638,911,1502,1526]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FDCDCAF8FC2EFA07" box="[638,911,1502,1526]" italics="true" pageId="11" pageNumber="43">Liolaemus magellanicus</emphasis>
</taxonomicName>
do differentiate it from the other groups of the 
<taxonomicName id="489D4D35FFCEFE33FF24C925FEC5F9EB" box="[151,356,1538,1562]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCEFE33FF24C925FEC5F9EB" box="[151,356,1538,1562]" italics="true" pageId="11" pageNumber="43">L. lineomaculatus</emphasis>
</taxonomicName>
section (scale counts and shape, coloration patterns, presence of precloacal pores and size, among others: 
<bibRefCitation id="EB0C4B47FFCEFE33FEF2C900FE7BF9CF" author="Scolaro" box="[321,474,1574,1598]" pageId="11" pageNumber="43" refString="Scolaro, J. A. (1992) Morphological differences between Vilcunia and L. magellanicus and L. lineomaculatus: Discriminant analysis. Acta Zoologica Lilloana, 41, 287 - 293." type="journal article" year="1992">Scolaro 1992</bibRefCitation>
; 
<bibRefCitation id="EB0C4B47FFCEFE33FE5BC901FD23F9CF" author="Laurent" box="[488,642,1574,1598]" pageId="11" pageNumber="43" refString="Laurent, R. F. (1995) A tentative arrangement of subgenera of the genus Liolaemus Wiegmann (Reptilia: Squamata: Tropiduridae). Bulletin of the Maryland Herpetological Society, 31, 10 - 14." type="journal article" year="1995">Laurent 1995</bibRefCitation>
; 
<bibRefCitation id="EB0C4B47FFCEFE33FD23C900FC6FF9CF" author="Etheridge" box="[656,974,1574,1598]" pageId="11" pageNumber="43" refString="Etheridge, R. &amp; Espinoza, R. E. (2000) Taxonomy of the Liolaeminae (Squamata: Iguania: Tropiduridae) and a semi-annotated bibliography. Smithsonian Herpetological Information Service, 126, 1 - 64." type="journal article" year="2000">Etheridge &amp; Espinoza 2000</bibRefCitation>
; 
<bibRefCitation id="EB0C4B47FFCEFE33FC6FC900FAFCF9CF" author="Pincheira-Donoso" box="[988,1373,1574,1598]" pageId="11" pageNumber="43" refString="Pincheira-Donoso, D. &amp; Nunez, H. (2005) Las especies chilenas del genero Liolaemus (Iguania, Tropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion. Publicacion Ocasional del Museo Nacional de Historia Natural de Chile, Santiago, 486 pp." type="book" year="2005">Pincheira-Donoso &amp; Núñez 2005</bibRefCitation>
). The divergence of the 
<taxonomicName id="489D4D35FFCEFE33FED5C96CFE5AF993" box="[358,507,1610,1634]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FED5C96CFE5AF993" box="[358,507,1610,1634]" italics="true" pageId="11" pageNumber="43">magellanicus</emphasis>
</taxonomicName>
group from the rest of the section was inferred to have occurred during the Late Miocene (~8.46 million years ago, 
<bibRefCitation id="EB0C4B47FFCEFE33FD92C948FCAAF977" author="Breitman" box="[545,779,1646,1670]" pageId="11" pageNumber="43" refString="Breitman, M. F., Avila, L. J., Sites, J. W., Jr. &amp; Morando, M. (2011) Lizards from the end of the world: Phylogenetic relationships of the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemini). Molecular Phylogenetics and Evolution, 59, 364 - 376." type="journal article" year="2011">
Breitman 
<emphasis id="BDE9EAA4FFCEFE33FD23C949FD69F977" box="[656,712,1646,1670]" italics="true" pageId="11" pageNumber="43">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
<paragraph id="8F2236B6FFCEFE33FF76C9B4FBC4F8AF" blockId="11.[151,1437,1466,2031]" pageId="11" pageNumber="43">
Morphological, molecular, and geographic distributional data provide evidence to consider 
<taxonomicName id="489D4D35FFCEFE33FB67C9B4FF6EF93F" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="caparensis" status="sp. nov.">
<emphasis id="BDE9EAA4FFCEFE33FB67C9B4FF6EF93F" italics="true" pageId="11" pageNumber="43">Liolaemus caparensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6DA57DFFFCEFE33FF65C990FE91F93F" box="[214,304,1718,1742]" pageId="11" pageNumber="43" rank="species">
<emphasis id="BDE9EAA4FFCEFE33FF65C990FE91F93F" bold="true" box="[214,304,1718,1742]" pageId="11" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
a distinct species most closely related to 
<emphasis id="BDE9EAA4FFCEFE33FCB7C991FC1FF93F" box="[772,958,1718,1742]" italics="true" pageId="11" pageNumber="43">
<taxonomicName id="489D4D35FFCEFE33FCB7C991FC1BF93F" box="[772,954,1718,1742]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">L. magellanicus</taxonomicName>
,
</emphasis>
forming the 
<taxonomicName id="489D4D35FFCEFE33FBE1C990FB46F93F" box="[1106,1255,1718,1742]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="magellanicus">
<emphasis id="BDE9EAA4FFCEFE33FBE1C990FB46F93F" box="[1106,1255,1718,1742]" italics="true" pageId="11" pageNumber="43">magellanicus</emphasis>
</taxonomicName>
group. Because the aim of this work was to describe the new species and not to propose new taxonomical arrangements, we avoid hypothesizing new taxonomies or presenting morphological features that might characterize new classifications. Including the species described here and two more species described in another manuscript (Breitman 
<emphasis id="BDE9EAA4FFCEFE33FAF3C805FF70F8AF" italics="true" pageId="11" pageNumber="43">et al. in press</emphasis>
), the number of species included in the 
<taxonomicName id="489D4D35FFCEFE33FD22C861FCFBF8AF" box="[657,858,1862,1886]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCEFE33FD22C861FCFBF8AF" box="[657,858,1862,1886]" italics="true" pageId="11" pageNumber="43">L. lineomaculatus</emphasis>
</taxonomicName>
section increases to 21.
</paragraph>
<paragraph id="8F2236B6FFCEFE34FF76C84CFCA5FF06" blockId="11.[151,1437,1466,2031]" lastBlockId="12.[151,1436,151,248]" lastPageId="12" lastPageNumber="44" pageId="11" pageNumber="43">
Our research group is studying the 
<taxonomicName id="489D4D35FFCEFE33FDE2C84CFCD9F873" box="[593,888,1898,1922]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="43" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="BDE9EAA4FFCEFE33FDE2C84CFCD9F873" box="[593,888,1898,1922]" italics="true" pageId="11" pageNumber="43">Liolaemus lineomaculatus</emphasis>
</taxonomicName>
section to delimit species boundaries and reconstruct relationships from multiple sources of evidence. Our goal is to present monographic revisions of poorly known groups on the basis of an “integrative taxonomy” approach (
<bibRefCitation id="EB0C4B47FFCEFE33FC3CC894FBF8F83B" author="Padial" box="[911,1113,1970,1994]" pageId="11" pageNumber="43" refString="Padial, J. M., Miralles, A., De la Riva, I. &amp; Vences, M. (2010) The integrative future of taxonomy. Frontiers in Zoology, 7, 1 - 14." type="journal article" year="2010">
Padial 
<emphasis id="BDE9EAA4FFCEFE33FC6EC895FBB7F83B" box="[989,1046,1970,1994]" italics="true" pageId="11" pageNumber="43">et al.</emphasis>
2010
</bibRefCitation>
), based on independent lines of evidence to propose taxonomic re-arrangements and support hypotheses of species limits. In parallel, we are using integrative phylogeographic analyses based on molecular, morphological, ecological, and geographical data, to reconstruct a temporal sequence of demographic histories, and to eventually test for shared patterns of divergence with other Patagonian clades (
<bibRefCitation id="EB0C4B47FFC9FE34FD83CFF9FD59FF06" author="Sersic" box="[560,760,223,248]" pageId="12" pageNumber="44" refString="Sersic, A. N., Cosacov, A., Cocucci, A. A., Johnson, L. A., Pozner, R., Avila, L. J., Sites, J. W., Jr., &amp; Morando, M. (2011) Emerging phylogeographic patterns in plants and terrestrial vertebrates from Patagonia. Biological Journal of the Linnean Society, 110, 475 - 494." type="journal article" year="2011">
Sérsic 
<emphasis id="BDE9EAA4FFC9FE34FDCECFC7FD17FF09" box="[637,694,224,248]" italics="true" pageId="12" pageNumber="44">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>