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<document id="793AC360D9628B03F57621BFA5C3284B" ID-CLB-Dataset="20326" ID-DOI="10.11646/zootaxa.5105.4.2" ID-GBIF-Dataset="aa0d1d8e-cd45-426a-ab8f-fc2676335929" ID-ISSN="1175-5326" ID-Zenodo-Dep="6333757" ID-ZooBank="1C355434-BC48-4586-BC6B-E54EFB70A20E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646640119968" checkinUser="plazi" docAuthor="David, Patrick &amp; Deuti, Kaushik" docDate="2022" docId="3451EF4EBE59FFB5048DFB02FDF6F8E8" docLanguage="en" docName="zootaxa.5105.4.2.pdf" docOrigin="Zootaxa 5105 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pareas berdmorei Theobald 1868" docType="treatment" docVersion="8" lastPageNumber="494" masterDocId="C8689736BE5BFFB1041AFFCDFFB0FA6D" masterDocTitle="On the type specimens of Pareas macularius Theobald, 1868 and Pareas berdmorei Theobald, 1868 with the designation of a lectotype for Pareas macularius (Squamata: Serpentes: Pareidae)" masterLastPageNumber="500" masterPageNumber="490" pageNumber="492" updateTime="1699317287852" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6EC3D78EA488C115FE92A1B679153927">On the type specimens of Pareas macularius Theobald, 1868 and Pareas berdmorei Theobald, 1868 with the designation of a lectotype for Pareas macularius (Squamata: Serpentes: Pareidae)</mods:title>
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<mods:namePart id="724B25E68EB7F7963F342C249EBFAC62">David, Patrick</mods:namePart>
<mods:affiliation id="D2F26B201614F69E2B728BC30A33A812">Institut de Systématique, Évolution et Biodiversité (ISYEB), Muséum National dHistoire Naturelle, Sorbonne Université, École Pratique des Hautes Études, Université des Antilles, CNRS, CP 30, 57 rue Cuvier, F- 75005 Paris, France.</mods:affiliation>
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<mods:namePart id="B68298D8BA4CA383C0972530B3AC21F9">Deuti, Kaushik</mods:namePart>
<mods:affiliation id="50375C2ECF18CACC2CDC63F3475FFAEF">Zoological Survey of India, Herpetology Division, 27, Jawaharlal Nehru Road, Kolkata 700016, India.</mods:affiliation>
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<treatment id="3451EF4EBE59FFB5048DFB02FDF6F8E8" ID-DOI="http://doi.org/10.5281/zenodo.6343716" ID-GBIF-Taxon="193892586" ID-Zenodo-Dep="6343716" LSID="urn:lsid:plazi:treatment:3451EF4EBE59FFB5048DFB02FDF6F8E8" httpUri="http://treatment.plazi.org/id/3451EF4EBE59FFB5048DFB02FDF6F8E8" lastPageId="4" lastPageNumber="494" pageId="2" pageNumber="492">
<subSubSection id="F4E20DD3BE59FFB3048DFB02FD85FE85" box="[151,565,1230,1257]" pageId="2" pageNumber="492" type="nomenclature">
<paragraph id="BC475E58BE59FFB3048DFB02FD85FE85" blockId="2.[151,565,1230,1257]" box="[151,565,1230,1257]" pageId="2" pageNumber="492">
<heading id="E70FE934BE59FFB3048DFB02FD85FE85" bold="true" box="[151,565,1230,1257]" fontSize="11" level="1" pageId="2" pageNumber="492" reason="1">
<taxonomicName id="7BF825DBBE59FFB3048DFB02FD85FE85" authority="Theobald, 1868" authorityName="Theobald" authorityYear="1868" box="[151,565,1230,1257]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE59FFB3048DFB02FD85FE85" bold="true" box="[151,565,1230,1257]" pageId="2" pageNumber="492">
<emphasis id="8E8C824ABE59FFB3048DFB02FEDEFE84" bold="true" box="[151,366,1230,1257]" italics="true" pageId="2" pageNumber="492">Pareas berdmorei</emphasis>
Theobald, 1868
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F4E20DD3BE59FFB3048DFADAFEE3FF3B" pageId="2" pageNumber="492" type="description">
<paragraph id="BC475E58BE59FFB3048DFADAFEE3FF3B" blockId="2.[151,1437,1303,1977]" pageId="2" pageNumber="492">
<bibRefCitation id="D86923A9BE59FFB3048DFADAFE3AFF5F" author="Theobald, W." box="[151,394,1303,1330]" pageId="2" pageNumber="492" pagination="8 - 88" refId="ref7466" refString="Theobald, W. (1868 b) Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society of Bengal, 37, Extra number, pp. i - vi + [7] + 8 - 88 + i - iii (Appendix), pls. 1 - 4." type="journal article" year="1868">Theobald (1868b: 63)</bibRefCitation>
described
<taxonomicName id="7BF825DBBE59FFB30618FADAFD7AFF5C" authorityName="Theobald" authorityYear="1868" box="[514,714,1303,1329]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE59FFB30618FADAFD7AFF5C" box="[514,714,1303,1329]" italics="true" pageId="2" pageNumber="492">Pareas berdmorei</emphasis>
</taxonomicName>
by referring to the following combination of taxa and specimens (cited verbatim):
</paragraph>
</subSubSection>
<subSubSection id="F4E20DD3BE59FFB304DDFA92FD3CFFF0" pageId="2" pageNumber="492" type="multiple">
<paragraph id="BC475E58BE59FFB304DDFA92FD6CFF14" blockId="2.[151,1437,1303,1977]" box="[199,732,1375,1401]" pageId="2" pageNumber="492">
<emphasis id="8E8C824ABE59FFB304CEFA92FDE1FF14" box="[212,593,1375,1401]" italics="true" pageId="2" pageNumber="492">
<taxonomicName id="7BF825DBBE59FFB304CEFA92FE3DFF14" box="[212,397,1375,1401]" class="Reptilia" family="Pareatidae" genus="Aplopeltura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="boa">Aplopeltura boa</taxonomicName>
. Schl. apud Blyth
</emphasis>
. The adult.”
</paragraph>
<paragraph id="BC475E58BE59FFB304DDFA4EFD3CFFF0" blockId="2.[151,1437,1303,1977]" box="[199,652,1411,1437]" pageId="2" pageNumber="492">
<taxonomicName id="7BF825DBBE59FFB304CEFA4EFD9DFFF0" authority=", Blyth. The" authorityName="Blyth. The" box="[212,557,1411,1437]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE59FFB304CEFA4EFE17FFF0" box="[212,423,1411,1437]" italics="true" pageId="2" pageNumber="492">Pareas macularius</emphasis>
, Blyth. The
</taxonomicName>
young.”
</paragraph>
</subSubSection>
<subSubSection id="F4E20DD3BE59FFB304DDFA6AFE44FDD4" pageId="2" pageNumber="492" type="materials_examined">
<paragraph id="BC475E58BE59FFB304DDFA6AFB0BFC40" blockId="2.[151,1437,1303,1977]" pageId="2" pageNumber="492">
Namely,
<bibRefCitation id="D86923A9BE59FFB30531FA6AFE43FFAF" author="Theobald, W." box="[299,499,1447,1474]" pageId="2" pageNumber="492" pagination="8 - 88" refId="ref7466" refString="Theobald, W. (1868 b) Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society of Bengal, 37, Extra number, pp. i - vi + [7] + 8 - 88 + i - iii (Appendix), pls. 1 - 4." type="journal article" year="1868">Theobald (1868b)</bibRefCitation>
placed in the synonymy of his new species both “
<taxonomicName id="7BF825DBBE59FFB30001FA6AFAD6FFAC" authority="&quot; sensu Blyth" authorityName="Blyth" box="[1051,1382,1447,1473]" class="Reptilia" family="Pareatidae" genus="Aplopeltura" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="boa">
<emphasis id="8E8C824ABE59FFB30001FA6AFB61FFAC" box="[1051,1233,1447,1473]" italics="true" pageId="2" pageNumber="492">Aplopeltura boa</emphasis>
” sensu Blyth
</taxonomicName>
(nec
<taxonomicName id="7BF825DBBE59FFB3048DFA06FE65FF88" authority="H. Boie" authorityName="H. Boie" box="[151,469,1483,1509]" class="Reptilia" family="Colubridae" genus="Amblycephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="boa">
<emphasis id="8E8C824ABE59FFB3048DFA06FEC3FF88" box="[151,371,1483,1509]" italics="true" pageId="2" pageNumber="492">Amblycephalus boa</emphasis>
H. Boie
</taxonomicName>
, a valid, distinct species), based on the
<specimenCount id="AAFE95D1BE59FFB30795FA06FC48FF88" box="[911,1016,1483,1509]" pageId="2" pageNumber="492" type="adult">two adult</specimenCount>
specimens (ZSI 80218022), and the three young (ZSI 80248026) mentioned previously by
<bibRefCitation id="D86923A9BE59FFB30710FA22FC62FC67" author="Theobald, W." box="[778,978,1519,1546]" pageId="2" pageNumber="492" pagination="4 - 67" refId="ref7393" refString="Theobald, W. (1868 a) Catalogue of the Reptiles of British Birma, embracing the Provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. Journal of the Linnean Society, London, Zoology, 10 (1), 4 - 67. https: // doi. org / 10.1111 / j. 1096 - 3642.1868. tb 02007. x" type="journal article" year="1868">Theobald (1868a)</bibRefCitation>
in the description of
<taxonomicName id="7BF825DBBE59FFB300D8FA22FA25FC64" box="[1218,1429,1519,1545]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE59FFB300D8FA22FA25FC64" box="[1218,1429,1519,1545]" italics="true" pageId="2" pageNumber="492">Pareas macularius</emphasis>
</taxonomicName>
. Therefore, the original reference of
<taxonomicName id="7BF825DBBE59FFB30632F9DEFD41FC40" authorityName="Theobald" authorityYear="1868" box="[552,753,1555,1581]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE59FFB30632F9DEFD41FC40" box="[552,753,1555,1581]" italics="true" pageId="2" pageNumber="492">Pareas berdmorei</emphasis>
</taxonomicName>
, with its original
<typeStatus id="6343E0FABE59FFB307ACF9DEFBABFC40" box="[950,1051,1555,1581]" pageId="2" pageNumber="492" type="syntype">syntypes</typeStatus>
, is as follows:
</paragraph>
<paragraph id="BC475E58BE59FFB304DDF9FAFC50FCD0" blockId="2.[151,1437,1303,1977]" pageId="2" pageNumber="492">
<treatmentCitationGroup id="9CE87976BE59FFB304DDF9FAFC50FCD0" pageId="2" pageNumber="492">
<treatmentCitation id="3D597849BE59FFB304DDF9FAFDCAFC3C" author="Theobald, W." box="[199,634,1591,1617]" page="63" pageId="2" pageNumber="492" year="1868">
<taxonomicName id="7BF825DBBE59FFB304DDF9FAFDCAFC3C" authority="Theobald, 1868 b: 63" authorityName="Theobald" authorityPageNumber="63" authorityYear="1868" box="[199,634,1591,1617]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE59FFB304DDF9FAFE3FFC3C" box="[199,399,1591,1617]" italics="true" pageId="2" pageNumber="492">Pareas berdmorei</emphasis>
<materialsCitation id="0C905405BE59FFB3058EF9FAFDCAFC3C" ID-GBIF-Occurrence="3698899301" box="[404,634,1591,1617]" collectionCode="ZSI" collectorName="Major T. M. Berdmore. &amp; Major T. M. Berdmore" country="Myanmar" location="Mottama" municipality="Mon State" pageId="2" pageNumber="492" specimenCode="ZSI 8021, ZSI 8022, ZSI 8024-8026" specimenCount="6" specimenCount-juvenile="1" stateProvince="Taninthayi" typeStatus="syntype">
<bibRefCitation id="D86923A9BE59FFB3058EF9FAFDCAFC3C" author="Theobald, W." box="[404,634,1591,1617]" pageId="2" pageNumber="492" pagination="8 - 88" refId="ref7466" refString="Theobald, W. (1868 b) Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society of Bengal, 37, Extra number, pp. i - vi + [7] + 8 - 88 + i - iii (Appendix), pls. 1 - 4." type="journal article" year="1868">Theobald, 1868b: 63</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.—Type locality. “Tenasserim”, now
<collectingRegion id="7E3C90BABE59FFB3000CF9FAFB41FC3C" box="[1046,1265,1591,1617]" country="Myanmar" name="Taninthayi" pageId="2" pageNumber="492">Tanintharyi Region</collectingRegion>
,
<collectingCountry id="C4EF1EC8BE59FFB300E1F9FAFADAFC3C" box="[1275,1386,1591,1617]" name="Myanmar" pageId="2" pageNumber="492">Myanmar</collectingCountry>
, and “Martaban”, now
<location id="B9270883BE59FFB30579F996FE7BFC18" LSID="urn:lsid:plazi:treatment:3451EF4EBE59FFB5048DFB02FDF6F8E8:B9270883BE59FFB30579F996FE7BFC18" box="[355,459,1627,1653]" country="Myanmar" municipality="Mon State" name="Mottama" pageId="2" pageNumber="492" stateProvince="Taninthayi">Mottama</location>
,
<collectingMunicipality id="5C23C422BE59FFB305CDF996FDFFFC18" box="[471,591,1627,1653]" pageId="2" pageNumber="492">Môn State</collectingMunicipality>
,
<collectingCountry id="C4EF1EC8BE59FFB30641F996FD7AFC18" box="[603,714,1627,1653]" name="Myanmar" pageId="2" pageNumber="492">Myanmar</collectingCountry>
.—
<typeStatus id="6343E0FABE59FFB306F1F996FCE1FC18" box="[747,849,1627,1653]" pageId="2" pageNumber="492" type="syntype">Syntypes</typeStatus>
(
<specimenCount id="AAFE95D1BE59FFB3077BF997FC42FC1B" box="[865,1010,1626,1654]" pageId="2" pageNumber="492" type="generic">5 specimens</specimenCount>
). (12)
<specimenCode id="EC5EF623BE59FFB30050F996FB07FC18" box="[1098,1207,1627,1653]" collectionCode="ZSI" pageId="2" pageNumber="492">ZSI 8021</specimenCode>
, adult male and
<specimenCode id="EC5EF623BE59FFB30168F996FF62FCF4" collectionCode="ZSI" pageId="2" pageNumber="492">ZSI 8022</specimenCode>
, adult female, both from “Tenasserim”; deposited by
<collectorName id="110D3B8EBE59FFB3072DF9B2FBF6FCF4" box="[823,1094,1662,1689]" pageId="2" pageNumber="492">Major T. M. Berdmore.</collectorName>
—(35)
<specimenCode id="EC5EF623BE59FFB300BEF9B2FAE7FCF4" box="[1188,1367,1663,1689]" collectionCode="ZSI" pageId="2" pageNumber="492">ZSI 80248026</specimenCode>
,
<specimenCount id="AAFE95D1BE59FFB3017EF9B2FF41FCD0" pageId="2" pageNumber="492" type="juvenile">three juvenile</specimenCount>
specimens from “Martaban”; deposited by
<collectorName id="110D3B8EBE59FFB306C0F96EFC6DFCD0" box="[730,989,1699,1725]" pageId="2" pageNumber="492">Major T. M. Berdmore</collectorName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BC475E58BE59FFB304DDF90AFE44FDD4" blockId="2.[151,1437,1303,1977]" pageId="2" pageNumber="492">
The
<typeStatus id="6343E0FABE59FFB304E6F90AFEEEFC8C" box="[252,350,1735,1761]" pageId="2" pageNumber="492" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="7BF825DBBE59FFB3059DF90AFDE1FC8C" authorityName="Theobald" authorityYear="1868" box="[391,593,1735,1761]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE59FFB3059DF90AFDE1FC8C" box="[391,593,1735,1761]" italics="true" pageId="2" pageNumber="492">Pareas berdmorei</emphasis>
</taxonomicName>
are thus identical to those of
<taxonomicName id="7BF825DBBE59FFB307B0F90AFBCFFC8C" box="[938,1151,1735,1761]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE59FFB307B0F90AFBCFFC8C" box="[938,1151,1735,1761]" italics="true" pageId="2" pageNumber="492">Pareas macularius</emphasis>
</taxonomicName>
. Such a case is accepted by Art. 72.6 of the Code (I. C. Z. N. 1999). For this reason,
<taxonomicName id="7BF825DBBE59FFB3075BF926FC61FD68" authorityName="Theobald" authorityYear="1868" box="[833,977,1771,1797]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE59FFB3075BF926FC61FD68" box="[833,977,1771,1797]" italics="true" pageId="2" pageNumber="492">P. berdmorei</emphasis>
</taxonomicName>
and
<taxonomicName id="7BF825DBBE59FFB30012F926FB13FD68" box="[1032,1187,1771,1797]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE59FFB30012F926FB13FD68" box="[1032,1187,1771,1797]" italics="true" pageId="2" pageNumber="492">P. macularius</emphasis>
</taxonomicName>
have sometimes been considered synonyms in the literature. However, these
<specimenCount id="AAFE95D1BE59FFB30713F8C2FC13FD44" box="[777,931,1807,1833]" pageId="2" pageNumber="492" type="juvenile">three juvenile</specimenCount>
individuals are obviously distinct at species level from the
<specimenCount id="AAFE95D1BE59FFB30525F8FEFE19FD20" box="[319,425,1843,1869]" pageId="2" pageNumber="492" type="adult">two adult</specimenCount>
that are referable to the
<taxonomicName id="7BF825DBBE59FFB306D9F8FEFC33FD21" box="[707,899,1843,1868]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="8E8C824ABE59FFB306D9F8FEFC33FD21" box="[707,899,1843,1868]" italics="true" pageId="2" pageNumber="492">Pareas carinatus</emphasis>
</taxonomicName>
species complex. Consequently,
<bibRefCitation id="D86923A9BE59FFB300E1F8FEFA3BFD20" author="Smith, M. A." box="[1275,1419,1843,1869]" pageId="2" pageNumber="492" pagination="1 - 135" refId="ref7283" refString="Smith, M. A. (1930) The Reptilia and Amphibia of the Malay Peninsula from the Isthmus of Kra to Singapore (A supplement to Dr. G. A. Boulenger's Reptilia and Batrachia, 1912). Bulletin of the Raffles Museum, 3, i - xviii + 1 - 135." type="journal article" year="1930">Smith (1930</bibRefCitation>
a;
<bibRefCitation id="D86923A9BE59FFB3048DF89AFF60FD1C" author="Smith, M. A." box="[151,208,1879,1905]" pageId="2" pageNumber="492" refId="ref7341" refString="Smith, M. A. (1943) The fauna of British India, Ceylon and Burma, including the whole of the Indo-chinese subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor &amp; Francis, London, xii + 583 pp., folded map." type="book" year="1943">1943</bibRefCitation>
) placed
<taxonomicName id="7BF825DBBE59FFB30528F89AFE4BFD1C" authorityName="Theobald" authorityYear="1868" box="[306,507,1879,1905]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE59FFB30528F89AFE4BFD1C" box="[306,507,1879,1905]" italics="true" pageId="2" pageNumber="492">Pareas berdmorei</emphasis>
</taxonomicName>
in the synonymy of
<taxonomicName id="7BF825DBBE59FFB306F2F89AFCDEFD1D" box="[744,878,1879,1904]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="8E8C824ABE59FFB306F2F89AFCDEFD1D" box="[744,878,1879,1904]" italics="true" pageId="2" pageNumber="492">P. carinatus</emphasis>
</taxonomicName>
, a synonymy long accepted in the literature (see, for example,
<bibRefCitation id="D86923A9BE59FFB30537F8B6FDBEFDF8" author="Wallach, V. &amp; Williams, K. L. &amp; Boundy, J." box="[301,526,1915,1941]" pageId="2" pageNumber="492" refId="ref7794" refString="Wallach, V., Williams, K. L. &amp; Boundy, J. (2014) Snakes of the world. A catalogue of living and extinct species. CRC Press, Boca Raton, Florida, xxvii + 1209 pp." type="book" year="2014">
Wallach
<emphasis id="8E8C824ABE59FFB3058AF8B6FE72FDF8" box="[400,450,1915,1941]" italics="true" pageId="2" pageNumber="492">et al</emphasis>
. 2014
</bibRefCitation>
). Eventually,
<bibRefCitation id="D86923A9BE59FFB306BDF8B6FC16FDF8" author="Poyarkov, N. A. &amp; Nguyen, T. V. &amp; Pawangkhanant, P. &amp; Yushchenko, P. V. &amp; Brakels, P. &amp; Nguyen, L. H. &amp; Nguyen, H. N. &amp; Suwannapoom, C. &amp; Orlov, N. &amp; Vogel, G." box="[679,934,1915,1941]" pageId="2" pageNumber="492" pagination="1 - 99" refId="ref7137" refString="Poyarkov, N. A., Nguyen, T. V., Pawangkhanant, P., Yushchenko, P. V., Brakels, P., Nguyen, L. H., Nguyen, H. N., Suwannapoom, C., Orlov, N. &amp; Vogel, G. (2022) An integrative taxonomic revision of Slug-eating Snakes (Squamata: Pareidae: Pareineae) reveals unprecedented diversity in Indochina. PeerJ, 10 (e 12713) 1 - 99. https: // doi. org / 10.7717 / peerj. 12713" type="journal article" year="2022">
Poyarkov
<emphasis id="8E8C824ABE59FFB30700F8B6FCFCFDF8" box="[794,844,1915,1941]" italics="true" pageId="2" pageNumber="492">et al</emphasis>
. (2022)
</bibRefCitation>
showed that
<taxonomicName id="7BF825DBBE59FFB30027F8B6FB7EFDF8" authorityName="Theobald" authorityYear="1868" box="[1085,1230,1915,1941]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="498" phylum="Chordata" rank="species" species="berdmorei">
P.
<emphasis id="8E8C824ABE59FFB30041F8B6FB7EFDF8" box="[1115,1230,1915,1941]" italics="true" pageId="2" pageNumber="492">berdmorei</emphasis>
</taxonomicName>
is a valid species, distinct from
<taxonomicName id="7BF825DBBE59FFB30534F852FE5DFDD5" box="[302,493,1951,1976]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="492" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="8E8C824ABE59FFB30534F852FE5DFDD5" box="[302,493,1951,1976]" italics="true" pageId="2" pageNumber="492">Pareas carinatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="E8870ED0BE58FFB2048DF881FCC5FD99" ID-DOI="http://doi.org/10.5281/zenodo.6333759" ID-Zenodo-Dep="6333759" httpUri="https://zenodo.org/record/6333759/files/figure.png" pageId="3" pageNumber="493" startId="3.[151,250,1868,1892]" targetBox="[401,1263,181,1844]" targetPageId="3">
<paragraph id="BC475E58BE58FFB2048DF881FCC5FD99" blockId="3.[151,1436,1868,2036]" pageId="3" pageNumber="493">
<emphasis id="8E8C824ABE58FFB2048DF881FEA5FD09" bold="true" box="[151,277,1868,1892]" pageId="3" pageNumber="493">FIGURE 1.</emphasis>
The original syntypes of
<taxonomicName id="7BF825DBBE58FFB2063AF880FD50FD09" box="[544,736,1868,1892]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="493" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE58FFB2063AF880FD50FD09" box="[544,736,1868,1892]" italics="true" pageId="3" pageNumber="493">Pareas macularius</emphasis>
</taxonomicName>
and
<taxonomicName id="7BF825DBBE58FFB2070FF880FC7CFD09" authorityName="Theobald" authorityYear="1868" box="[789,972,1868,1892]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="493" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE58FFB2070FF880FC7CFD09" box="[789,972,1868,1892]" italics="true" pageId="3" pageNumber="493">Pareas berdmorei</emphasis>
</taxonomicName>
. A. The three “young” specimens mentioned by
<bibRefCitation id="D86923A9BE58FFB204A2F8BDFED7FDE5" author="Theobald, W." box="[184,359,1904,1928]" pageId="3" pageNumber="493" pagination="4 - 67" refId="ref7393" refString="Theobald, W. (1868 a) Catalogue of the Reptiles of British Birma, embracing the Provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. Journal of the Linnean Society, London, Zoology, 10 (1), 4 - 67. https: // doi. org / 10.1111 / j. 1096 - 3642.1868. tb 02007. x" type="journal article" year="1868">Theobald (1868a</bibRefCitation>
b), original syntypes of both of
<taxonomicName id="7BF825DBBE58FFB206A1F8BCFCCBFDE5" box="[699,891,1904,1928]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="493" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE58FFB206A1F8BCFCCBFDE5" box="[699,891,1904,1928]" italics="true" pageId="3" pageNumber="493">Pareas macularius</emphasis>
</taxonomicName>
and
<taxonomicName id="7BF825DBBE58FFB207B5F8BCFB29FDE5" authority=", ZSI" authorityName="ZSI" box="[943,1177,1904,1928]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="493" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE58FFB207B5F8BCFBD6FDE5" box="[943,1126,1904,1928]" italics="true" pageId="3" pageNumber="493">Pareas berdmorei</emphasis>
, ZSI
</taxonomicName>
8024, ZSI 8025 and ZSI 8026.—B. The two “adult” specimens mentioned by
<bibRefCitation id="D86923A9BE58FFB206A7F859FCDDFDC1" author="Theobald, W." box="[701,877,1940,1964]" pageId="3" pageNumber="493" pagination="4 - 67" refId="ref7393" refString="Theobald, W. (1868 a) Catalogue of the Reptiles of British Birma, embracing the Provinces of Pegu, Martaban, and Tenasserim; with descriptions of new or little-known species. Journal of the Linnean Society, London, Zoology, 10 (1), 4 - 67. https: // doi. org / 10.1111 / j. 1096 - 3642.1868. tb 02007. x" type="journal article" year="1868">Theobald (1868a</bibRefCitation>
b), ZSI 8021 and ZSI 8022 (top and centre), original syntypes of both of
<taxonomicName id="7BF825DBBE58FFB2057EF874FD94FDBD" box="[356,548,1976,2000]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="493" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE58FFB2057EF874FD94FDBD" box="[356,548,1976,2000]" italics="true" pageId="3" pageNumber="493">Pareas macularius</emphasis>
</taxonomicName>
and
<taxonomicName id="7BF825DBBE58FFB2064CF874FCBCFDBD" box="[598,780,1976,2000]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="493" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE58FFB2064CF874FCBCFDBD" box="[598,780,1976,2000]" italics="true" pageId="3" pageNumber="493">Pareas berdmorei</emphasis>
</taxonomicName>
, as well as specimen ZSI 8023 (bottom) that does not belong to the type series of either of these species. Photographs by Kaushik Deuti.
</paragraph>
</caption>
<subSubSection id="F4E20DD3BE5FFFB504DDFF5AFDF6F8E8" pageId="4" pageNumber="494" type="discussion">
<paragraph id="BC475E58BE5FFFB504DDFF5AFD1BFB08" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="494">
There have been some confusion in the literature about the type series of
<taxonomicName id="7BF825DBBE5FFFB5001DFF5AFB60FADC" box="[1031,1232,151,177]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE5FFFB5001DFF5AFB60FADC" box="[1031,1232,151,177]" italics="true" pageId="4" pageNumber="494">Pareas berdmorei</emphasis>
</taxonomicName>
.
<bibRefCitation id="D86923A9BE5FFFB500C5FF5AFF7AFAB8" author="Das, I. &amp; Datta Gupta, B. &amp; Gayen, N. C." pageId="4" pageNumber="494" pagination="121 - 172" refId="ref6431" refString="Das, I., Datta Gupta, B. &amp; Gayen, N. C. (1998) History and catalogue of reptiles types in the collection of the Zoological Survey of India. Journal of South Asian Natural History, 3 (2), 121 - 172." type="journal article" year="1998">
Das
<emphasis id="8E8C824ABE5FFFB50108FF5AFAF4FADC" box="[1298,1348,151,177]" italics="true" pageId="4" pageNumber="494">et al</emphasis>
. (1998: 150)
</bibRefCitation>
recognized
<specimenCount id="AAFE95D1BE5FFFB5054CFF76FE4BFAB8" box="[342,507,187,213]" pageId="4" pageNumber="494" type="generic" typeStatus="syntypes">three syntypes</specimenCount>
, ZSI 8021 to ZSI 8023. The description of this taxon by
<bibRefCitation id="D86923A9BE5FFFB50098FF76FAFCFABB" author="Theobald, W." box="[1154,1356,187,214]" pageId="4" pageNumber="494" pagination="8 - 88" refId="ref7466" refString="Theobald, W. (1868 b) Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society of Bengal, 37, Extra number, pp. i - vi + [7] + 8 - 88 + i - iii (Appendix), pls. 1 - 4." type="journal article" year="1868">Theobald (1868b)</bibRefCitation>
makes clear that it was based on two “adult” specimens. We examined these specimens, still extant and in good condition. Only ZSI 8021 and ZSI 8022 are adult animals, as correctly stated by
<bibRefCitation id="D86923A9BE5FFFB507ADFECEFB19FB70" author="Bhosale, H. &amp; Phansalkar, P. &amp; Sawant, M. &amp; Gowande, G. &amp; Patel, H. &amp; Mirza, Z. A." box="[951,1193,259,285]" pageId="4" pageNumber="494" pagination="54 - 73" refId="ref6174" refString="Bhosale, H., Phansalkar, P., Sawant, M., Gowande, G., Patel, H. &amp; Mirza, Z. A. (2020) A new species of snail-eating snakes of the genus Pareas Wagler, 1830 (Reptilia: Serpentes) from eastern Himalayas, India. European Journal of Taxonomy, 729 (1), 54 - 73. https: // doi. org / 10.5852 / ejt. 2020.729.1191" type="journal article" year="2020">
Bhosale
<emphasis id="8E8C824ABE5FFFB50003FECEFBFCFB70" box="[1049,1100,259,285]" italics="true" pageId="4" pageNumber="494">et al</emphasis>
. (2020)
</bibRefCitation>
. Specimen ZSI
<quantity id="7B00F3BDBE5FFFB50147FECEFF46FB2C" metricMagnitude="3" metricUnit="m" metricValue="8.023208" pageId="4" pageNumber="494" unit="mm" value="8023208.0">8023, 208 mm</quantity>
long (SVL:
<quantity id="7B00F3BDBE5FFFB5059CFEEAFE58FB2C" box="[390,488,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.66" pageId="4" pageNumber="494" unit="mm" value="166.0">166 mm</quantity>
; TaL:
<quantity id="7B00F3BDBE5FFFB5062BFEEAFD35FB2C" box="[561,645,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="494" unit="mm" value="42.0">42 mm</quantity>
; examined), is a juvenile or perhaps a neonate (
<figureCitation id="24C342DDBE5FFFB500BCFEEAFB4EFB2C" box="[1190,1278,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[401,1263,181,1844]" captionTargetId="figure-30@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURE 1. The original syntypes of Pareas macularius and Pareas berdmorei. A. The three “young” specimens mentioned by Theobald (1868ab), original syntypes of both of Pareas macularius and Pareas berdmorei, ZSI 8024, ZSI 8025 and ZSI 8026.—B. The two “adult” specimens mentioned by Theobald (1868ab), ZSI 8021 and ZSI 8022 (top and centre), original syntypes of both of Pareas macularius and Pareas berdmorei, as well as specimen ZSI 8023 (bottom) that does not belong to the type series of either of these species. Photographs by Kaushik Deuti." figureDoi="http://doi.org/10.5281/zenodo.6333759" httpUri="https://zenodo.org/record/6333759/files/figure.png" pageId="4" pageNumber="494">Fig. 1B</figureCitation>
). It cannot be considered one of the
<typeStatus id="6343E0FABE5FFFB50594FE86FE40FB08" box="[398,496,331,357]" pageId="4" pageNumber="494" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="7BF825DBBE5FFFB5060CFE86FD14FB08" box="[534,676,331,357]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE5FFFB5060CFE86FD14FB08" box="[534,676,331,357]" italics="true" pageId="4" pageNumber="494">P. berdmorei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="BC475E58BE5FFFB504DDFEA2FDF6F8E8" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="494">
According to the way in which
<bibRefCitation id="D86923A9BE5FFFB50633FEA2FD43FBE7" author="Theobald, W." box="[553,755,367,394]" pageId="4" pageNumber="494" pagination="8 - 88" refId="ref7466" refString="Theobald, W. (1868 b) Catalogue of Reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society of Bengal, 37, Extra number, pp. i - vi + [7] + 8 - 88 + i - iii (Appendix), pls. 1 - 4." type="journal article" year="1868">Theobald (1868b)</bibRefCitation>
described
<taxonomicName id="7BF825DBBE5FFFB50776FEA2FB85FBE4" box="[876,1077,367,393]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE5FFFB50776FEA2FB85FBE4" box="[876,1077,367,393]" italics="true" pageId="4" pageNumber="494">Pareas berdmorei</emphasis>
</taxonomicName>
and its type material, this taxon could have been considered either a junior objective synonym of
<taxonomicName id="7BF825DBBE5FFFB50771FE5EFBB4FBC0" box="[875,1028,403,429]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE5FFFB50771FE5EFBB4FBC0" box="[875,1028,403,429]" italics="true" pageId="4" pageNumber="494">P. macularius</emphasis>
</taxonomicName>
or a replacement name for this latter taxon. However, the type series of
<taxonomicName id="7BF825DBBE5FFFB50605FE7AFD1FFBBC" box="[543,687,439,465]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE5FFFB50605FE7AFD1FFBBC" box="[543,687,439,465]" italics="true" pageId="4" pageNumber="494">P. berdmorei</emphasis>
</taxonomicName>
is obviously composed of specimens referable to two distinct species easy to diagnose. In their revision,
<bibRefCitation id="D86923A9BE5FFFB50648FE16FCFEFB98" author="Poyarkov, N. A. &amp; Nguyen, T. V. &amp; Pawangkhanant, P. &amp; Yushchenko, P. V. &amp; Brakels, P. &amp; Nguyen, L. H. &amp; Nguyen, H. N. &amp; Suwannapoom, C. &amp; Orlov, N. &amp; Vogel, G." box="[594,846,475,501]" pageId="4" pageNumber="494" pagination="1 - 99" refId="ref7137" refString="Poyarkov, N. A., Nguyen, T. V., Pawangkhanant, P., Yushchenko, P. V., Brakels, P., Nguyen, L. H., Nguyen, H. N., Suwannapoom, C., Orlov, N. &amp; Vogel, G. (2022) An integrative taxonomic revision of Slug-eating Snakes (Squamata: Pareidae: Pareineae) reveals unprecedented diversity in Indochina. PeerJ, 10 (e 12713) 1 - 99. https: // doi. org / 10.7717 / peerj. 12713" type="journal article" year="2022">
Poyarkov
<emphasis id="8E8C824ABE5FFFB506DEFE16FD45FB98" box="[708,757,475,501]" italics="true" pageId="4" pageNumber="494">et al</emphasis>
. (2022)
</bibRefCitation>
showed that
<taxonomicName id="7BF825DBBE5FFFB507F9FE16FBC1FB98" box="[995,1137,475,501]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE5FFFB507F9FE16FBC1FB98" box="[995,1137,475,501]" italics="true" pageId="4" pageNumber="494">P. berdmorei</emphasis>
</taxonomicName>
is not a replacement name of
<taxonomicName id="7BF825DBBE5FFFB504AFFE32FEFFF874" box="[181,335,511,537]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE5FFFB504AFFE32FEFFF874" box="[181,335,511,537]" italics="true" pageId="4" pageNumber="494">P. macularius</emphasis>
</taxonomicName>
or a junior synonym of this latter species but a valid species, distinct from both
<taxonomicName id="7BF825DBBE5FFFB500D7FE32FAD6F874" box="[1229,1382,511,537]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE5FFFB500D7FE32FAD6F874" box="[1229,1382,511,537]" italics="true" pageId="4" pageNumber="494">P. macularius</emphasis>
</taxonomicName>
and, also
<taxonomicName id="7BF825DBBE5FFFB504D1FDEEFEE0F851" box="[203,336,547,572]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="carinatus">
<emphasis id="8E8C824ABE5FFFB504D1FDEEFEE0F851" box="[203,336,547,572]" italics="true" pageId="4" pageNumber="494">P. carinatus</emphasis>
</taxonomicName>
at species level. In order to fix the nomenclatural status at species level of
<taxonomicName id="7BF825DBBE5FFFB5008FFDEEFEFAF80C" authority=", Poyarkov et al. (2022: 47)" authorityName="Poyarkov" authorityPageNumber="47" authorityYear="2022" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="8E8C824ABE5FFFB5008FFDEEFA94F850" box="[1173,1316,547,573]" italics="true" pageId="4" pageNumber="494">P. berdmorei</emphasis>
,
<bibRefCitation id="D86923A9BE5FFFB5012BFDEEFEFAF80C" author="Poyarkov, N. A. &amp; Nguyen, T. V. &amp; Pawangkhanant, P. &amp; Yushchenko, P. V. &amp; Brakels, P. &amp; Nguyen, L. H. &amp; Nguyen, H. N. &amp; Suwannapoom, C. &amp; Orlov, N. &amp; Vogel, G." pageId="4" pageNumber="494" pagination="1 - 99" refId="ref7137" refString="Poyarkov, N. A., Nguyen, T. V., Pawangkhanant, P., Yushchenko, P. V., Brakels, P., Nguyen, L. H., Nguyen, H. N., Suwannapoom, C., Orlov, N. &amp; Vogel, G. (2022) An integrative taxonomic revision of Slug-eating Snakes (Squamata: Pareidae: Pareineae) reveals unprecedented diversity in Indochina. PeerJ, 10 (e 12713) 1 - 99. https: // doi. org / 10.7717 / peerj. 12713" type="journal article" year="2022">
Poyarkov
<emphasis id="8E8C824ABE5FFFB5048DFD8AFF78F80C" box="[151,200,583,609]" italics="true" pageId="4" pageNumber="494">et al</emphasis>
. (2022: 47)
</bibRefCitation>
</taxonomicName>
selected a
<typeStatus id="6343E0FABE5FFFB505DEFD8AFD9BF80C" box="[452,555,583,609]" pageId="4" pageNumber="494" type="lectotype">lectotype</typeStatus>
for this species (see below). In order to fix the status of
<taxonomicName id="7BF825DBBE5FFFB50081FD8AFADEF80C" box="[1179,1390,583,609]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="494" phylum="Chordata" rank="species" species="macularius">
<emphasis id="8E8C824ABE5FFFB50081FD8AFADEF80C" box="[1179,1390,583,609]" italics="true" pageId="4" pageNumber="494">Pareas macularius</emphasis>
</taxonomicName>
, we here select a
<typeStatus id="6343E0FABE5FFFB50533FDA6FE20F8E8" box="[297,400,619,645]" pageId="4" pageNumber="494" type="lectotype">lectotype</typeStatus>
for this species.
</paragraph>
</subSubSection>
</treatment>
</document>