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<document ID-DOI="10.1643/i2019299" ID-ISSN="2766-1520" ID-Zenodo-Dep="7846669" IM.illustrations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1681927084425" checkinUser="felipe" docAuthor="Armbruster, Jonathan W., Lujan, Nathan K. &amp; Bloom, Devin D." docDate="2021" docId="0B02895DD564FFF9FF7AFA7D3D750AA5" docLanguage="en" docName="IchthyolHerpetol.109.1.102-122.pdf" docOrigin="Ichthyology &amp; Herpetology 109 (1)" docSource="http://dx.doi.org/10.1643/i2019299" docStyle="DocumentStyle:25DEBD253B07840CEE0E8BE8BE2C6752.1:IchthyolHerpetol.2021-.journal_article" docStyleId="25DEBD253B07840CEE0E8BE8BE2C6752" docStyleName="IchthyolHerpetol.2021-.journal_article" docStyleVersion="1" docTitle="Characidium wangyapoik Armbruster &amp; Lujan &amp; Bloom 2021, new species" docType="treatment" docVersion="3" lastPageNumber="120" masterDocId="F73BF125D56BFFEBFFF4FFFF3F450C62" masterDocTitle="Redescription of the Guiana Shield Darter Species Characidium crandellii and C. declivirostre (Crenuchidae) with Descriptions of Two New Species" masterLastPageNumber="122" masterPageNumber="102" pageNumber="117" updateTime="1682004287626" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Redescription of the Guiana Shield Darter Species Characidium crandellii and C. declivirostre (Crenuchidae) with Descriptions of Two New Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Armbruster, Jonathan W.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lujan, Nathan K.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bloom, Devin D.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Ichthyology &amp; Herpetology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-29</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>109</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>102</mods:start>
<mods:end>122</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1643/i2019299</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1643/i2019299</mods:identifier>
<mods:identifier type="ISSN">2766-1520</mods:identifier>
<mods:identifier type="Zenodo-Dep">7846669</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5" httpUri="http://treatment.plazi.org/id/0B02895DD564FFF9FF7AFA7D3D750AA5" lastPageId="18" lastPageNumber="120" pageId="15" pageNumber="117">
<subSubSection pageId="15" pageNumber="117" type="nomenclature">
<paragraph blockId="15.[142,590,1410,1433]" box="[142,590,1410,1433]" pageId="15" pageNumber="117">
<heading bold="true" box="[142,590,1410,1433]" fontSize="10" level="2" pageId="15" pageNumber="117" reason="2">
<emphasis bold="true" box="[142,590,1410,1433]" pageId="15" pageNumber="117">
<taxonomicName authority="Armbruster &amp; Lujan &amp; Bloom, 2021" authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[142,431,1410,1433]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="15" pageNumber="117" phylum="Chordata" rank="species" species="wangyapoik" status="new species">
<emphasis bold="true" box="[142,431,1410,1433]" italics="true" pageId="15" pageNumber="117">Characidium wangyapoik</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[446,590,1410,1433]" pageId="15" pageNumber="117" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[142,709,1439,1521]" pageId="15" pageNumber="117">
<uri pageId="15" pageNumber="117">
urn:lsid:zoobank.org:act:
<uuid pageId="15" pageNumber="117">1EBB34BE-431E-4D03-A343- 6E8132EE810C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[142,334,1498,1521]" pageId="15" pageNumber="117" type="description">
<paragraph blockId="15.[142,709,1439,1521]" box="[142,334,1498,1521]" pageId="15" pageNumber="117">
<figureCitation box="[142,257,1498,1521]" captionStart="FIG" captionStartId="1.[142,170,670,690]" captionTargetBox="[142,1528,178,641]" captionTargetId="figure-704@1.[142,1528,178,641]" captionTargetPageId="1" captionText="FIG. 1. Thickened, keratinized pads on the undersurface of the anterior pectoral-fin rays of (A) Characidium wangyapoik, new species, CSBD F-3615, holotype, and (B) C. crandellii, AUM 67142. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846671" httpUri="https://zenodo.org/record/7846671/files/figure.png" pageId="15" pageNumber="117">Figures 1A</figureCitation>
,
<figureCitation box="[268,334,1498,1521]" captionStart-0="FIG" captionStart-1="FIG" captionStartId-0="15.[142,170,1319,1339]" captionStartId-1="16.[103,131,1331,1351]" captionTargetBox-0="[142,1530,178,1290]" captionTargetBox-1="[103,1491,178,1301]" captionTargetId-0="figure-339@15.[142,1530,178,1290]" captionTargetId-1="figure-376@16.[103,1491,178,1301]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIG. 11. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, in life, CSBD F-3615, holotype, 72.6 mm SL. Photos by N. K. Lujan." captionText-1="FIG. 12. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, after preservation, CSBD F-3615, holotype, 72.6 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi-0="http://doi.org/10.5281/zenodo.7846691" figureDoi-1="http://doi.org/10.5281/zenodo.7846693" httpUri-0="https://zenodo.org/record/7846691/files/figure.png" httpUri-1="https://zenodo.org/record/7846693/files/figure.png" pageId="15" pageNumber="117">11–12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="117" type="reference_group">
<paragraph blockId="15.[142,812,1543,1594]" pageId="15" pageNumber="117">
<treatmentCitationGroup pageId="15" pageNumber="117">
<taxonomicName ID-CoL="3M73" authority="Armbruster &amp; Lujan &amp; Bloom, 2021" authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[142,276,1543,1565]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="15" pageNumber="117" phylum="Chordata" rank="species" species="undetermined" status="n. sp.">
<emphasis box="[142,276,1543,1565]" italics="true" pageId="15" pageNumber="117">Characidium</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[284,345,1544,1565]" pageId="15" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
‘Ireng’.—
<treatmentCitation author="Lujan, N. K. &amp; J. W. Armbruster &amp; D. C. Werneke &amp; T. Franco Teixeira &amp; N. R. Lovejoy" box="[450,716,1543,1566]" page="1216" pageId="15" pageNumber="117" year="2020">
<bibRefCitation author="Lujan, N. K. &amp; J. W. Armbruster &amp; D. C. Werneke &amp; T. Franco Teixeira &amp; N. R. Lovejoy" box="[450,716,1543,1566]" pageId="15" pageNumber="117" pagination="1213 - 1235" refId="ref19030" refString="Lujan, N. K., J. W. Armbruster, D. C. Werneke, T. Franco Teixeira, and N. R. Lovejoy. 2020. Phylogeny and biogeography of the Brazilian-Guiana Shield endemic Corymbophanes clade of armoured catfishes (Loricariidae). Zoological Journal of the Linnean Society 188: 1213 - 1235." type="journal article" year="2020">Lujan et al., 2020: 1216</bibRefCitation>
</treatmentCitation>
[locality information].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="118" pageId="15" pageNumber="117" type="materials_examined">
<paragraph blockId="15.[142,812,1626,1795]" pageId="15" pageNumber="117">
<materialsCitation collectingDate="2016-01-08" collectionCode="CSBD" collectorName="N. K. Lujan &amp; J. W. Armbruster &amp; D. C. Werneke &amp; M. Ram" country="Guyana" county="Sukwabi Creek" latitude="5.07711" location="Potaro-Siparuni" longLatPrecision="1" longitude="-59.97423" municipality="Ireng River" pageId="15" pageNumber="117" specimenCode="F-3615, AUM 67118" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[142,267,1626,1649]" italics="true" pageId="15" pageNumber="117">
<typeStatus box="[142,239,1626,1649]" pageId="15" pageNumber="117">Holotype</typeStatus>
.—
</emphasis>
<collectionCode box="[266,329,1626,1647]" pageId="15" pageNumber="117">CSBD</collectionCode>
<specimenCode box="[336,411,1626,1648]" collectionCode="F-" pageId="15" pageNumber="117">F-3615</specimenCode>
(ex
<specimenCode box="[458,594,1626,1649]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67118</specimenCode>
), 1 (1mo/me, 1gm),
<quantity box="[142,247,1655,1677]" metricMagnitude="-2" metricUnit="m" metricValue="7.26" pageId="15" pageNumber="117" unit="mm" value="72.6">72.6 mm</quantity>
SL,
<collectingCountry box="[302,388,1656,1677]" name="Guyana" pageId="15" pageNumber="117">Guyana</collectingCountry>
(border with
<collectingCountry box="[553,618,1655,1678]" name="Brazil" pageId="15" pageNumber="117">Brazil</collectingCountry>
),
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FD75F9883C690AEF" box="[641,812,1655,1677]" country="Guyana" county="Sukwabi Creek" latitude="5.07711" longLatPrecision="1" longitude="-59.97423" municipality="Ireng River" name="Potaro-Siparuni" pageId="15" pageNumber="117">Potaro-Siparuni</location>
(Region 8),
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FEEAF96A3E840AC9" box="[286,449,1685,1707]" country="Guyana" county="Sukwabi Creek" latitude="5.07711" longLatPrecision="1" longitude="-59.97423" municipality="Ireng River" name="Amazon River" pageId="15" pageNumber="117">Amazon River</location>
basin,
<collectingMunicipality box="[545,676,1685,1707]" pageId="15" pageNumber="117">Ireng River</collectingMunicipality>
, first shoal upriver from split with
<collectingCounty box="[407,574,1714,1736]" pageId="15" pageNumber="117">Sukwabi Creek</collectingCounty>
,
<geoCoordinate box="[586,677,1714,1736]" degrees="5.07711" direction="north" orientation="latitude" pageId="15" pageNumber="117" precision="1" value="5.07711">5.07711</geoCoordinate>
,
<geoCoordinate box="[690,807,1714,1736]" degrees="59.97423" direction="west" orientation="longitude" pageId="15" pageNumber="117" precision="1" value="-59.97423">–59.97423</geoCoordinate>
,
<collectorName box="[142,279,1743,1765]" pageId="15" pageNumber="117">N. K. Lujan</collectorName>
,
<collectorName box="[294,488,1743,1765]" pageId="15" pageNumber="117">J. W. Armbruster</collectorName>
,
<collectorName box="[503,674,1743,1765]" pageId="15" pageNumber="117">D. C. Werneke</collectorName>
,
<collectorName box="[690,782,1744,1765]" pageId="15" pageNumber="117">M. Ram</collectorName>
,
<date pageId="15" pageNumber="117" value="2016-01-08">
<collectingDate pageId="15" pageNumber="117" value="2016-01-08">8 January 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="15.[142,812,1826,2024]" lastBlockId="16.[820,1490,1422,1561]" lastPageId="16" lastPageNumber="118" pageId="15" pageNumber="117">
<materialsCitation country="Guyana" county="Rio Mau" location="All" municipality="Ireng River" pageId="15" pageNumber="117" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[142,280,1826,1849]" italics="true" pageId="15" pageNumber="117">
<typeStatus box="[142,251,1826,1849]" pageId="15" pageNumber="117">Paratypes</typeStatus>
.—
</emphasis>
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FEECF8DD3E720B5A" box="[280,311,1826,1848]" country="Guyana" county="Rio Mau" municipality="Ireng River" name="All" pageId="15" pageNumber="117">All</location>
specimens
<collectingCountry box="[461,549,1826,1847]" name="Guyana" pageId="15" pageNumber="117">Guyana</collectingCountry>
(border with
<collectingCountry box="[733,800,1826,1849]" name="Brazil" pageId="15" pageNumber="117">Brazil</collectingCountry>
),
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FF7AF8C03E7C0B37" box="[142,313,1855,1877]" country="Guyana" county="Rio Mau" municipality="Ireng River" name="Potaro-Siparuni" pageId="15" pageNumber="117">Potaro-Siparuni</location>
(
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FEBCF8C03ED00B37" box="[328,405,1855,1877]" country="Guyana" county="Rio Mau" municipality="Ireng River" name="Region" pageId="15" pageNumber="117">Region</location>
8),
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FE30F8C03D1A0B37" box="[452,607,1855,1877]" country="Guyana" county="Rio Mau" municipality="Ireng River" name="Amazon River" pageId="15" pageNumber="117">Amazon River</location>
basin,
<collectingMunicipality box="[690,812,1855,1877]" pageId="15" pageNumber="117">Ireng River</collectingMunicipality>
basin (known as the
<collectingCounty box="[392,489,1884,1906]" pageId="15" pageNumber="117">Rio Mau</collectingCounty>
in
<collectingCountry box="[536,602,1884,1907]" name="Brazil" pageId="15" pageNumber="117">Brazil</collectingCountry>
)
</materialsCitation>
:
<materialsCitation collectionCode="ANSP" country="Guyana" pageId="15" pageNumber="117" specimenCode="ANSP 207526, 3" specimenCount="1" typeStatus="paratype">
<specimenCode box="[624,782,1884,1906]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="15" pageNumber="117">ANSP 207526</specimenCode>
,
<specimenCode box="[798,812,1885,1906]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="15" pageNumber="117">3</specimenCode>
(1mo/me, 3gm),
<quantity box="[335,499,1914,1936]" metricMagnitude="-2" metricUnit="m" metricValue="4.97" metricValueMax="5.68" metricValueMin="4.26" pageId="15" pageNumber="117" unit="mm" value="49.7" valueMax="56.8" valueMin="42.6">42.6–56.8 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation collectionCode="AUM" country="Guyana" pageId="15" pageNumber="117" specimenCode="AUM 67036, 26" specimenCount="1" typeStatus="paratype">
<specimenCode box="[552,691,1914,1936]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67036</specimenCode>
,
<specimenCode box="[706,733,1914,1936]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">26</specimenCode>
(5mo/ me, 26gm, 4cs), 22.0–
<quantity box="[382,481,1943,1965]" metricMagnitude="-2" metricUnit="m" metricValue="5.99" pageId="15" pageNumber="117" unit="mm" value="59.9">59.9 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation collectionCode="CSBD" country="Guyana" pageId="15" pageNumber="117" specimenCode="F-3616, 3" specimenCount="1" typeStatus="paratype">
<collectionCode box="[526,589,1943,1964]" pageId="15" pageNumber="117">CSBD</collectionCode>
<specimenCode box="[595,673,1943,1965]" collectionCode="F-" pageId="15" pageNumber="117">F-3616</specimenCode>
,
<specimenCode box="[683,697,1943,1964]" collectionCode="F-" pageId="15" pageNumber="117">3</specimenCode>
(1mo/me, 3gm),
<quantity box="[221,389,1972,1994]" metricMagnitude="-2" metricUnit="m" metricValue="4.495" metricValueMax="5.22" metricValueMin="3.77" pageId="15" pageNumber="117" unit="mm" value="44.95" valueMax="52.2" valueMin="37.7">37.7–52.2 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation collectionCode="INPA" country="Guyana" pageId="15" pageNumber="117" specimenCode="ICT-059496, 3" specimenCount="1" typeStatus="paratype">
<collectionCode box="[452,508,1973,1994]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="15" pageNumber="117">INPA</collectionCode>
<specimenCode box="[523,655,1972,1994]" collectionCode="ICT-" pageId="15" pageNumber="117">ICT-059496</specimenCode>
,
<specimenCode box="[674,688,1973,1994]" collectionCode="ICT-" pageId="15" pageNumber="117">3</specimenCode>
(0mo/me, 3gm),
<quantity box="[217,381,2002,2024]" metricMagnitude="-2" metricUnit="m" metricValue="5.36" metricValueMax="5.46" metricValueMin="5.26" pageId="15" pageNumber="117" unit="mm" value="53.6" valueMax="54.6" valueMin="52.6">52.6–54.6 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation collectingDate="2016-01-02" collectionCode="ROM" collectorName="D. C. Werneke &amp; J. W. Armbruster &amp; N. K. Lujan &amp; Ram" country="Guyana" latitude="4.72536" location="Ireng River" longLatPrecision="1" longitude="-60.03852" municipality="Orinduik Falls" pageId="15" pageNumber="117" specimenCode="ROM 111286, 3" specimenCount="1" typeStatus="paratype">
<specimenCode box="[435,589,2002,2024]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="15" pageNumber="117" type="Museum">ROM 111286</specimenCode>
,
<specimenCode box="[604,618,2002,2023]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="15" pageNumber="117" type="Museum">3</specimenCode>
(0mo/me, 3gm),
<quantity box="[859,1024,1410,1432]" metricMagnitude="-2" metricUnit="m" metricValue="5.075" metricValueMax="5.38" metricValueMin="4.77" pageId="15" pageNumber="117" unit="mm" value="50.75" valueMax="53.8" valueMin="47.7">47.7–53.8 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FBCDFA7C3BFD09FA" box="[1081,1208,1410,1432]" country="Guyana" latitude="4.72536" longLatPrecision="1" longitude="-60.03852" municipality="Orinduik Falls" name="Ireng River" pageId="15" pageNumber="117">Ireng River</location>
, at
<collectingMunicipality box="[1257,1420,1410,1432]" pageId="15" pageNumber="117">Orinduik Falls</collectingMunicipality>
, between upper and lower falls,
<geoCoordinate box="[1107,1199,1440,1462]" degrees="4.72536" direction="north" orientation="latitude" pageId="15" pageNumber="117" precision="1" value="4.72536">4.72536</geoCoordinate>
,
<geoCoordinate box="[1210,1327,1440,1462]" degrees="60.03852" direction="west" orientation="longitude" pageId="15" pageNumber="117" precision="1" value="-60.03852">–60.03852</geoCoordinate>
,
<collectorName box="[1339,1503,1440,1462]" pageId="15" pageNumber="117">D. C. Werneke</collectorName>
,
<collectorName pageId="15" pageNumber="117">J. W. Armbruster</collectorName>
,
<collectorName box="[1037,1169,1469,1491]" pageId="15" pageNumber="117">N. K. Lujan</collectorName>
, M.
<collectorName box="[1220,1272,1470,1491]" pageId="15" pageNumber="117">Ram</collectorName>
,
<date box="[1285,1458,1469,1491]" pageId="15" pageNumber="117" value="2016-01-02">
<collectingDate box="[1285,1458,1469,1491]" pageId="15" pageNumber="117" value="2016-01-02">2 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-03" collectionCode="AUM" collectorName="N. K. Lujan &amp; J. W. Armbruster &amp; D. C. Werneke &amp; Ram" country="Guyana" latitude="4.71898" location="Ireng River" longLatPrecision="1" longitude="-60.03507" municipality="lower Orinduik Falls" pageId="15" pageNumber="117" specimenCode="AUM 67046, 16" specimenCount="1" typeStatus="paratype">
<specimenCode collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67046</specimenCode>
,
<specimenCode box="[946,973,1499,1521]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">16</specimenCode>
(0),
<quantity box="[1032,1196,1499,1521]" metricMagnitude="-2" metricUnit="m" metricValue="3.21" metricValueMax="4.64" metricValueMin="1.78" pageId="15" pageNumber="117" unit="mm" value="32.1" valueMax="46.4" valueMin="17.8">17.8–46.4 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FB16FA233A250993" box="[1250,1376,1499,1521]" country="Guyana" latitude="4.71898" longLatPrecision="1" longitude="-60.03507" municipality="lower Orinduik Falls" name="Ireng River" pageId="15" pageNumber="117">Ireng River</location>
, below
<collectingMunicipality pageId="15" pageNumber="117">lower Orinduik Falls</collectingMunicipality>
,
<geoCoordinate box="[1049,1141,1529,1550]" degrees="4.71898" direction="north" orientation="latitude" pageId="15" pageNumber="117" precision="1" value="4.71898">4.71898</geoCoordinate>
,
<geoCoordinate box="[1161,1280,1528,1550]" degrees="60.03507" direction="west" orientation="longitude" pageId="15" pageNumber="117" precision="1" value="-60.03507">–60.03507</geoCoordinate>
,
<collectorName box="[1300,1448,1529,1551]" pageId="15" pageNumber="117">N. K. Lujan</collectorName>
,
<collectorName pageId="15" pageNumber="117">J. W. Armbruster</collectorName>
,
<collectorName box="[999,1166,1558,1580]" pageId="15" pageNumber="117">D. C. Werneke</collectorName>
, M.
<collectorName box="[1218,1270,1559,1580]" pageId="15" pageNumber="117">Ram</collectorName>
,
<date box="[1284,1458,1558,1580]" pageId="15" pageNumber="117" value="2016-01-03">
<collectingDate box="[1284,1458,1558,1580]" pageId="15" pageNumber="117" value="2016-01-03">3 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-04" collectionCode="AUM" collectorName="D. C. Werneke &amp; J. W. Armbruster &amp; D. I. Brooks &amp; Ram" country="Guyana" county="Orinduik Falls" latitude="4.67585" location="Ireng River" longLatPrecision="1" longitude="-60.06046" municipality="Branana Rapids" pageId="15" pageNumber="117" specimenCode="AUM 67068, 18" specimenCount="1" typeStatus="paratype">
<specimenCode collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67068</specimenCode>
,
<specimenCode box="[947,974,1588,1609]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">18</specimenCode>
(0),
<quantity box="[1037,1202,1588,1610]" metricMagnitude="-2" metricUnit="m" metricValue="3.435" metricValueMax="4.35" metricValueMin="2.52" pageId="15" pageNumber="117" unit="mm" value="34.35" valueMax="43.5" valueMin="25.2">25.2–43.5 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FB18F9CB3A290A28" box="[1260,1388,1588,1610]" country="Guyana" county="Orinduik Falls" latitude="4.67585" longLatPrecision="1" longitude="-60.06046" municipality="Branana Rapids" name="Ireng River" pageId="15" pageNumber="117">Ireng River</location>
, at
<collectingMunicipality pageId="15" pageNumber="117">Branana Rapids</collectingMunicipality>
, shoals downstream of
<collectingCounty box="[1239,1411,1617,1639]" pageId="15" pageNumber="117">Orinduik Falls</collectingCounty>
,
<geoCoordinate box="[1431,1525,1617,1639]" degrees="4.67585" direction="north" orientation="latitude" pageId="15" pageNumber="117" precision="1" value="4.67585">4.67585</geoCoordinate>
,
<geoCoordinate box="[859,976,1647,1669]" degrees="60.06046" direction="west" orientation="longitude" pageId="15" pageNumber="117" precision="1" value="-60.06046">–60.06046</geoCoordinate>
,
<collectorName box="[992,1163,1647,1669]" pageId="15" pageNumber="117">D. C. Werneke</collectorName>
,
<collectorName box="[1178,1371,1647,1669]" pageId="15" pageNumber="117">J. W. Armbruster</collectorName>
,
<collectorName box="[1385,1525,1647,1669]" pageId="15" pageNumber="117">D. I. Brooks</collectorName>
, M.
<collectorName box="[898,950,1677,1698]" pageId="15" pageNumber="117">Ram</collectorName>
,
<date box="[963,1136,1676,1698]" pageId="15" pageNumber="117" value="2016-01-04">
<collectingDate box="[963,1136,1676,1698]" pageId="15" pageNumber="117" value="2016-01-04">4 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-04" collectionCode="AUM" collectorName="D. C. Werneke &amp; J. W. Armbruster &amp; D. I. Brooks &amp; M. Ram &amp; N. K. Lujan" country="Guyana" latitude="4.72536" location="Ireng River" longLatPrecision="1" longitude="-60.03852" municipality="Orinduik Falls" pageId="15" pageNumber="117" specimenCode="AUM 67076, 4" specimenCount="1" typeStatus="paratype">
<specimenCode box="[1149,1286,1676,1698]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67076</specimenCode>
,
<specimenCode box="[1300,1314,1677,1698]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">4</specimenCode>
(0), 36.0–
<quantity box="[1428,1529,1676,1698]" metricMagnitude="-2" metricUnit="m" metricValue="4.04" pageId="15" pageNumber="117" unit="mm" value="40.4">40.4 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FC71F9543B470AA2" box="[901,1026,1706,1728]" country="Guyana" latitude="4.72536" longLatPrecision="1" longitude="-60.03852" municipality="Orinduik Falls" name="Ireng River" pageId="15" pageNumber="117">Ireng River</location>
, at
<collectingMunicipality box="[1069,1229,1706,1728]" pageId="15" pageNumber="117">Orinduik Falls</collectingMunicipality>
, between upper and lower falls,
<geoCoordinate box="[920,1012,1735,1757]" degrees="4.72536" direction="north" orientation="latitude" pageId="15" pageNumber="117" precision="1" value="4.72536">4.72536</geoCoordinate>
,
<geoCoordinate box="[1025,1142,1735,1757]" degrees="60.03852" direction="west" orientation="longitude" pageId="15" pageNumber="117" precision="1" value="-60.03852">–60.03852</geoCoordinate>
,
<collectorName box="[1155,1323,1736,1758]" pageId="15" pageNumber="117">D. C. Werneke</collectorName>
,
<collectorName box="[1336,1525,1736,1758]" pageId="15" pageNumber="117">J. W. Armbruster</collectorName>
,
<collectorName box="[859,1000,1765,1787]" pageId="15" pageNumber="117">D. I. Brooks</collectorName>
,
<collectorName box="[1015,1108,1766,1787]" pageId="15" pageNumber="117">M. Ram</collectorName>
,
<collectorName box="[1124,1261,1765,1787]" pageId="15" pageNumber="117">N. K. Lujan</collectorName>
,
<date box="[1277,1455,1765,1787]" pageId="15" pageNumber="117" value="2016-01-04">
<collectingDate box="[1277,1455,1765,1787]" pageId="15" pageNumber="117" value="2016-01-04">4 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-04" collectionCode="AUM" collectorName="D. C. Werneke &amp; J. W. Armbruster &amp; D. I. Brooks &amp; M. Ram &amp; N. K. Lujan" country="Guyana" latitude="4.72176" location="Ireng River" longLatPrecision="1" longitude="-60.03703" municipality="Orinduik Falls" pageId="15" pageNumber="117" specimenCode="AUM 67077, 20" specimenCount="1" typeStatus="paratype">
<specimenCode collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67077</specimenCode>
,
<specimenCode box="[941,968,1795,1816]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">20</specimenCode>
(0mo/me, 13gm),
<quantity box="[1173,1332,1795,1817]" metricMagnitude="-2" metricUnit="m" metricValue="4.185" metricValueMax="5.1899999999999995" metricValueMin="3.18" pageId="15" pageNumber="117" unit="mm" value="41.85" valueMax="51.9" valueMin="31.8">31.8–51.9 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FA95F8FC3A9F0B7B" box="[1377,1498,1795,1817]" country="Guyana" latitude="4.72176" longLatPrecision="1" longitude="-60.03703" municipality="Orinduik Falls" name="Ireng River" pageId="15" pageNumber="117">Ireng River</location>
, at
<collectingMunicipality box="[859,1023,1824,1846]" pageId="15" pageNumber="117">Orinduik Falls</collectingMunicipality>
, around halfway between upper and lower falls,
<geoCoordinate box="[920,1012,1854,1876]" degrees="4.72176" direction="north" orientation="latitude" pageId="15" pageNumber="117" precision="1" value="4.72176">4.72176</geoCoordinate>
,
<geoCoordinate box="[1025,1142,1854,1876]" degrees="60.03703" direction="west" orientation="longitude" pageId="15" pageNumber="117" precision="1" value="-60.03703">–60.03703</geoCoordinate>
,
<collectorName box="[1155,1323,1854,1876]" pageId="15" pageNumber="117">D. C. Werneke</collectorName>
,
<collectorName box="[1336,1525,1854,1876]" pageId="15" pageNumber="117">J. W. Armbruster</collectorName>
,
<collectorName box="[859,1000,1883,1905]" pageId="15" pageNumber="117">D. I. Brooks</collectorName>
,
<collectorName box="[1015,1108,1884,1905]" pageId="15" pageNumber="117">M. Ram</collectorName>
,
<collectorName box="[1124,1261,1883,1905]" pageId="15" pageNumber="117">N. K. Lujan</collectorName>
,
<date box="[1277,1455,1883,1905]" pageId="15" pageNumber="117" value="2016-01-04">
<collectingDate box="[1277,1455,1883,1905]" pageId="15" pageNumber="117" value="2016-01-04">4 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-05" collectionCode="AUM" collectorName="N. K. Lujan &amp; J. W. Armbruster &amp; D. C. Werneke &amp; D. I. Brooks &amp; Ram" country="Guyana" latitude="4.72798" location="Ireng River" longLatPrecision="1" longitude="-60.03597" municipality="Orinduik Falls" pageId="15" pageNumber="117" specimenCode="AUM 67094, 1" specimenCount="1" typeStatus="paratype">
<specimenCode collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67094</specimenCode>
,
<specimenCode box="[944,958,1913,1934]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">1</specimenCode>
(0),
<quantity box="[1014,1116,1913,1935]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="15" pageNumber="117" unit="mm" value="30.2">30.2 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D564FFE4FB7BF8853A490BED" box="[1167,1292,1913,1935]" country="Guyana" latitude="4.72798" longLatPrecision="1" longitude="-60.03597" municipality="Orinduik Falls" name="Ireng River" pageId="15" pageNumber="117">Ireng River</location>
, just above
<collectingMunicipality pageId="15" pageNumber="117">Orinduik Falls</collectingMunicipality>
,
<geoCoordinate box="[923,1014,1943,1964]" degrees="4.72798" direction="north" orientation="latitude" pageId="15" pageNumber="117" precision="1" value="4.72798">4.72798</geoCoordinate>
,
<geoCoordinate box="[1028,1145,1942,1964]" degrees="60.03597" direction="west" orientation="longitude" pageId="15" pageNumber="117" precision="1" value="-60.03597">–60.03597</geoCoordinate>
,
<collectorName box="[1158,1290,1942,1964]" pageId="15" pageNumber="117">N. K. Lujan</collectorName>
,
<collectorName box="[1303,1491,1942,1964]" pageId="15" pageNumber="117">J. W. Armbruster</collectorName>
,
<collectorName pageId="15" pageNumber="117">D. C. Werneke</collectorName>
,
<collectorName box="[1010,1151,1972,1994]" pageId="15" pageNumber="117">D. I. Brooks</collectorName>
, M.
<collectorName box="[1208,1260,1973,1994]" pageId="15" pageNumber="117">Ram</collectorName>
,
<date box="[1276,1455,1972,1994]" pageId="15" pageNumber="117" value="2016-01-05">
<collectingDate box="[1276,1455,1972,1994]" pageId="15" pageNumber="117" value="2016-01-05">5 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectionCode="AUM" country="Guyana" pageId="15" pageNumber="117" specimenCode="AUM 67118, 14" specimenCount="1" typeStatus="paratype">
<specimenCode collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">AUM 67118</specimenCode>
,
<specimenCode box="[942,969,2002,2023]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="15" pageNumber="117" type="University or college">14</specimenCode>
(4mo/me, 7gm, 2cs),
<quantity box="[1212,1372,2002,2024]" metricMagnitude="-2" metricUnit="m" metricValue="6.275" metricValueMax="7.609999999999999" metricValueMin="4.9399999999999995" pageId="15" pageNumber="117" unit="mm" value="62.75" valueMax="76.1" valueMin="49.4">49.4–76.1 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation collectingDate="2016-01-08" collectionCode="INPA" collectorName="N. K. Lujan &amp; J. W. Armbruster &amp; D. C. Werneke &amp; M. Ram" country="Guyana" lastPageId="16" lastPageNumber="118" latitude="5.07711" location="Sukwabi Creek" longLatPrecision="1" longitude="-59.97423" pageId="15" pageNumber="117" specimenCode="ICT-059497, 2" specimenCount="1" typeStatus="paratype">
<collectionCode box="[1419,1475,2002,2023]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="15" pageNumber="117">INPA</collectionCode>
<specimenCode collectionCode="ICT-" lastPageId="16" lastPageNumber="118" pageId="15" pageNumber="117">ICT-059497</specimenCode>
,
<specimenCode box="[206,220,1422,1443]" collectionCode="ICT-" pageId="16" pageNumber="117">2</specimenCode>
(1mo/me, 2gm),
<quantity box="[432,598,1422,1444]" metricMagnitude="-2" metricUnit="m" metricValue="6.16" metricValueMax="6.26" metricValueMin="6.06" pageId="16" pageNumber="118" unit="mm" value="61.6" valueMax="62.6" valueMin="60.6">60.6–62.6 mm</quantity>
SL, first shoal upriver from split with
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFE84FA533D5209A0" box="[368,535,1452,1474]" country="Guyana" latitude="5.07711" longLatPrecision="1" longitude="-59.97423" name="Sukwabi Creek" pageId="16" pageNumber="118">Sukwabi Creek</location>
,
<geoCoordinate box="[547,638,1452,1474]" degrees="5.07711" direction="north" orientation="latitude" pageId="16" pageNumber="118" precision="1" value="5.07711">5.07711</geoCoordinate>
,
<geoCoordinate box="[651,768,1452,1474]" degrees="59.97423" direction="west" orientation="longitude" pageId="16" pageNumber="118" precision="1" value="-59.97423">–59.97423</geoCoordinate>
,
<collectorName box="[103,240,1483,1505]" pageId="16" pageNumber="118">N. K. Lujan</collectorName>
,
<collectorName box="[255,449,1483,1505]" pageId="16" pageNumber="118">J. W. Armbruster</collectorName>
,
<collectorName box="[464,635,1483,1505]" pageId="16" pageNumber="118">D. C. Werneke</collectorName>
,
<collectorName box="[651,743,1483,1504]" pageId="16" pageNumber="118">M. Ram</collectorName>
,
<date pageId="16" pageNumber="118" value="2016-01-08">
<collectingDate pageId="16" pageNumber="118" value="2016-01-08">8 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-09" collectionCode="AUM" collectorName="N. K. Lujan &amp; J. W. Armbruster &amp; D. C. Werneke" country="Guyana" county="Kaibarupai" latitude="5.04398" location="Ireng River" longLatPrecision="1" longitude="-59.97717" municipality="Monkey Creek" pageId="16" pageNumber="118" specimenCode="AUM 67143, 6" specimenCount="1" typeStatus="paratype">
<specimenCode box="[262,397,1513,1535]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">AUM 67143</specimenCode>
,
<specimenCode box="[408,422,1513,1535]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">6</specimenCode>
(0mo/me, 6gm),
<quantity box="[614,773,1513,1535]" metricMagnitude="-2" metricUnit="m" metricValue="5.625" metricValueMax="6.77" metricValueMin="4.4799999999999995" pageId="16" pageNumber="118" unit="mm" value="56.25" valueMax="67.7" valueMin="44.8">44.8–67.7 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFF66F9F73E550A7C" box="[146,272,1544,1566]" country="Guyana" county="Kaibarupai" latitude="5.04398" longLatPrecision="1" longitude="-59.97717" municipality="Monkey Creek" name="Ireng River" pageId="16" pageNumber="118">Ireng River</location>
, shoals at mouth of
<collectingMunicipality box="[514,681,1544,1566]" pageId="16" pageNumber="118">Monkey Creek</collectingMunicipality>
,
<collectingCounty pageId="16" pageNumber="118">Kaibarupai</collectingCounty>
,
<geoCoordinate box="[167,258,1574,1596]" degrees="5.04398" direction="north" orientation="latitude" pageId="16" pageNumber="118" precision="1" value="5.04398">5.04398</geoCoordinate>
,
<geoCoordinate box="[271,388,1574,1596]" degrees="59.97717" direction="west" orientation="longitude" pageId="16" pageNumber="118" precision="1" value="-59.97717">–59.97717</geoCoordinate>
,
<collectorName box="[401,533,1574,1596]" pageId="16" pageNumber="118">N. K. Lujan</collectorName>
,
<collectorName box="[546,734,1574,1596]" pageId="16" pageNumber="118">J. W. Armbruster</collectorName>
,
<collectorName pageId="16" pageNumber="118">D. C. Werneke</collectorName>
,
<date box="[244,414,1605,1627]" pageId="16" pageNumber="118" value="2016-01-09">
<collectingDate box="[244,414,1605,1627]" pageId="16" pageNumber="118" value="2016-01-09">9 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectionCode="AUM" country="Guyana" pageId="16" pageNumber="118" specimenCode="AUM 67181, 9" specimenCount="1" typeStatus="paratype">
<specimenCode box="[426,561,1605,1627]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">AUM 67181</specimenCode>
,
<specimenCode box="[573,587,1605,1626]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">9</specimenCode>
(3mo/me, 9gm),
<quantity box="[103,261,1635,1657]" metricMagnitude="-2" metricUnit="m" metricValue="5.895" metricValueMax="6.88" metricValueMin="4.91" pageId="16" pageNumber="118" unit="mm" value="58.95" valueMax="68.8" valueMin="49.1">49.1–68.8 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation collectingDate="2016-01-12" collectionCode="ROM" collectorName="J. W. Armbruster &amp; N. K. Lujan &amp; D. I. Brooks" country="Guyana" county="Kaibarupai" latitude="5.04398" location="Ireng River" longLatPrecision="1" longitude="-59.97717" municipality="Monkey Creek" pageId="16" pageNumber="118" specimenCode="ROM 111287, 2" specimenCount="1" typeStatus="paratype">
<specimenCode box="[305,453,1636,1657]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="16" pageNumber="118" type="Museum">ROM 111287</specimenCode>
,
<specimenCode box="[462,476,1636,1657]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="16" pageNumber="118" type="Museum">2</specimenCode>
(2mo/me, 2gm),
<quantity metricMagnitude="-2" metricUnit="m" metricValue="6.095" metricValueMax="6.13" metricValueMin="6.06" pageId="16" pageNumber="118" unit="mm" value="60.95" valueMax="61.3" valueMin="60.6">60.6–61.3 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFF24F97D3E170AFA" box="[208,338,1666,1688]" country="Guyana" county="Kaibarupai" latitude="5.04398" longLatPrecision="1" longitude="-59.97717" municipality="Monkey Creek" name="Ireng River" pageId="16" pageNumber="118">Ireng River</location>
, shoals at mouth of
<collectingMunicipality box="[598,769,1666,1688]" pageId="16" pageNumber="118">Monkey Creek</collectingMunicipality>
,
<collectingCounty box="[103,224,1696,1718]" pageId="16" pageNumber="118">Kaibarupai</collectingCounty>
,
<geoCoordinate box="[243,334,1696,1718]" degrees="5.04398" direction="north" orientation="latitude" pageId="16" pageNumber="118" precision="1" value="5.04398">5.04398</geoCoordinate>
,
<geoCoordinate box="[353,471,1696,1718]" degrees="59.97717" direction="west" orientation="longitude" pageId="16" pageNumber="118" precision="1" value="-59.97717">–59.97717</geoCoordinate>
,
<collectorName box="[490,690,1696,1718]" pageId="16" pageNumber="118">J. W. Armbruster</collectorName>
,
<collectorName pageId="16" pageNumber="118">N. K. Lujan</collectorName>
,
<collectorName box="[182,318,1727,1749]" pageId="16" pageNumber="118">D. I. Brooks</collectorName>
,
<date box="[331,518,1727,1749]" pageId="16" pageNumber="118" value="2016-01-12">
<collectingDate box="[331,518,1727,1749]" pageId="16" pageNumber="118" value="2016-01-12">12 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectionCode="AUM" collectorName="J. W. Armbruster &amp; N. K. Lujan &amp; D. I. Brooks &amp; D. C. Werneke &amp; Peters" country="Guyana" county="Wotowanda Falls" latitude="5.08867" location="Sukwabi Creek" longLatPrecision="1" longitude="-59.96952" municipality="East Fork" pageId="16" pageNumber="118" specimenCode="AUM 67189, 2" specimenCount="1" typeStatus="paratype">
<specimenCode box="[531,669,1727,1749]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">AUM 67189</specimenCode>
,
<specimenCode box="[682,696,1727,1748]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">2</specimenCode>
(0mo/ me, 2gm),
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFF03F9223EE80A91" box="[247,429,1757,1779]" country="Guyana" county="Wotowanda Falls" latitude="5.08867" longLatPrecision="1" longitude="-59.96952" municipality="East Fork" name="Sukwabi Creek" pageId="16" pageNumber="118">Sukwabi Creek</location>
,
<collectingMunicipality box="[451,568,1757,1779]" pageId="16" pageNumber="118">East Fork</collectingMunicipality>
, downstream of
<collectingCounty box="[103,296,1788,1810]" pageId="16" pageNumber="118">Wotowanda Falls</collectingCounty>
,
<geoCoordinate box="[309,400,1788,1810]" degrees="5.08867" direction="north" orientation="latitude" pageId="16" pageNumber="118" precision="1" value="5.08867">5.08867</geoCoordinate>
,
<geoCoordinate box="[414,531,1788,1810]" degrees="59.96952" direction="west" orientation="longitude" pageId="16" pageNumber="118" precision="1" value="-59.96952">–59.96952</geoCoordinate>
,
<collectorName box="[545,734,1788,1810]" pageId="16" pageNumber="118">J. W. Armbruster</collectorName>
,
<collectorName pageId="16" pageNumber="118">N. K. Lujan</collectorName>
,
<collectorName box="[216,354,1819,1841]" pageId="16" pageNumber="118">D. I. Brooks</collectorName>
,
<collectorName box="[369,538,1819,1841]" pageId="16" pageNumber="118">D. C. Werneke</collectorName>
, P.
<collectorName box="[580,647,1819,1840]" pageId="16" pageNumber="118">Peters</collectorName>
</materialsCitation>
,
<materialsCitation collectingDate="2016-01-13" collectionCode="R" country="Guyana" location="Daniel" pageId="16" pageNumber="118" specimenCount="1" typeStatus="paratype">
<collectionCode box="[661,679,1819,1840]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="118">R</collectionCode>
.
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFD41F8E43C450B53" box="[693,768,1819,1841]" country="Guyana" name="Daniel" pageId="16" pageNumber="118">Daniel</location>
, local fishermen,
<date box="[287,471,1849,1871]" pageId="16" pageNumber="118" value="2016-01-13">
<collectingDate box="[287,471,1849,1871]" pageId="16" pageNumber="118" value="2016-01-13">13 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-14" collectionCode="AUM" collectorName="D. C. Werneke &amp; J. W. Armbruster &amp; N. K. Lujan &amp; M. Ram &amp; D. I. Brooks" country="Guyana" latitude="5.02404" location="Ireng River" longLatPrecision="1" longitude="-59.97763" municipality="Kaibarupai" pageId="16" pageNumber="118" specimenCode="AUM 67196, 12" specimenCount="1" typeStatus="paratype">
<specimenCode box="[483,618,1849,1871]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">AUM 67196</specimenCode>
,
<specimenCode box="[630,657,1849,1870]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">12</specimenCode>
(0mo/me, 11gm),
<quantity box="[199,371,1880,1902]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" metricValueMax="6.79" metricValueMin="2.81" pageId="16" pageNumber="118" unit="mm" value="48.0" valueMax="67.9" valueMin="28.1">28.1–67.9 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFE42F8A73D780B0C" box="[438,573,1880,1902]" country="Guyana" latitude="5.02404" longLatPrecision="1" longitude="-59.97763" municipality="Kaibarupai" name="Ireng River" pageId="16" pageNumber="118">Ireng River</location>
, downstream of
<collectingMunicipality box="[103,227,1910,1932]" pageId="16" pageNumber="118">Kaibarupai</collectingMunicipality>
,
<geoCoordinate box="[248,342,1910,1932]" degrees="5.02404" direction="north" orientation="latitude" pageId="16" pageNumber="118" precision="1" value="5.02404">5.02404</geoCoordinate>
,
<geoCoordinate box="[363,484,1910,1932]" degrees="59.97763" direction="west" orientation="longitude" pageId="16" pageNumber="118" precision="1" value="-59.97763">–59.97763</geoCoordinate>
,
<collectorName box="[505,690,1910,1932]" pageId="16" pageNumber="118">D. C. Werneke</collectorName>
,
<collectorName pageId="16" pageNumber="118">J. W. Armbruster</collectorName>
,
<collectorName box="[243,378,1941,1963]" pageId="16" pageNumber="118">N. K. Lujan</collectorName>
,
<collectorName box="[392,484,1941,1962]" pageId="16" pageNumber="118">M. Ram</collectorName>
,
<collectorName box="[498,636,1941,1963]" pageId="16" pageNumber="118">D. I. Brooks</collectorName>
,
<date pageId="16" pageNumber="118" value="2016-01-14">
<collectingDate pageId="16" pageNumber="118" value="2016-01-14">14 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-14" collectionCode="AUM" collectorName="D. C. Werneke &amp; J. W. Armbruster &amp; N. K. Lujan &amp; M. Ram &amp; D. I. Brooks" country="Guyana" latitude="4.96554" location="Ireng River" longLatPrecision="1" longitude="-59.99411" municipality="Sand Hill" pageId="16" pageNumber="118" specimenCode="AUM 67199, 2" specimenCount="1" typeStatus="paratype">
<specimenCode box="[175,313,1971,1993]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">AUM 67199</specimenCode>
,
<specimenCode box="[327,341,1971,1992]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">2</specimenCode>
(0),
<quantity box="[398,561,1971,1993]" metricMagnitude="-2" metricUnit="m" metricValue="2.145" metricValueMax="2.32" metricValueMin="1.97" pageId="16" pageNumber="118" unit="mm" value="21.45" valueMax="23.2" valueMin="19.7">19.7–23.2 mm</quantity>
SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFD91F84B3DA70BAB" box="[613,738,1971,1993]" country="Guyana" latitude="4.96554" longLatPrecision="1" longitude="-59.99411" municipality="Sand Hill" name="Ireng River" pageId="16" pageNumber="118">Ireng River</location>
, at
<collectingMunicipality box="[103,207,2002,2024]" pageId="16" pageNumber="118">Sand Hill</collectingMunicipality>
shoals,
<geoCoordinate box="[301,393,2002,2024]" degrees="4.96554" direction="north" orientation="latitude" pageId="16" pageNumber="118" precision="1" value="4.96554">4.96554</geoCoordinate>
,
<geoCoordinate box="[406,523,2002,2024]" degrees="59.99411" direction="west" orientation="longitude" pageId="16" pageNumber="118" precision="1" value="-59.99411">–59.99411</geoCoordinate>
,
<collectorName box="[537,705,2002,2024]" pageId="16" pageNumber="118">D. C. Werneke</collectorName>
,
<collectorName box="[718,946,1422,2023]" pageId="16" pageNumber="118">J. W. Armbruster</collectorName>
,
<collectorName box="[960,1095,1422,1444]" pageId="16" pageNumber="118">N. K. Lujan</collectorName>
,
<collectorName box="[1109,1201,1422,1443]" pageId="16" pageNumber="118">M. Ram</collectorName>
,
<collectorName box="[1215,1353,1422,1444]" pageId="16" pageNumber="118">D. I. Brooks</collectorName>
,
<date pageId="16" pageNumber="118" value="2016-01-14">
<collectingDate pageId="16" pageNumber="118" value="2016-01-14">14 January 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation collectingDate="2016-01-14" collectionCode="AUM" collectorName="D. C. Werneke &amp; J. W. Armbruster &amp; N. K. Lujan &amp; M. Ram &amp; D. I. Brooks" country="Guyana" latitude="4.93345" location="Ireng River" longLatPrecision="1" longitude="-59.99514" municipality="Waipa Landing" pageId="16" pageNumber="118" specimenCode="AUM 67204, 12" specimenCount="1" typeStatus="paratype">
<specimenCode box="[893,1033,1451,1473]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">AUM 67204</specimenCode>
,
<specimenCode box="[1048,1075,1451,1472]" collectionCode="AUM" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="16" pageNumber="118" type="University or college">12</specimenCode>
(5mo/me), 17.1–72.0 mm SL,
<location LSID="urn:lsid:plazi:treatment:0B02895DD564FFF9FF7AFA7D3D750AA5:86746E90D57BFFFBFA6DFA543C2B09BC" country="Guyana" latitude="4.93345" longLatPrecision="1" longitude="-59.99514" municipality="Waipa Landing" name="Ireng River" pageId="16" pageNumber="118">Ireng River</location>
, at
<collectingMunicipality box="[916,1085,1480,1502]" pageId="16" pageNumber="118">Waipa Landing</collectingMunicipality>
,
<geoCoordinate box="[1095,1186,1480,1502]" degrees="4.93345" direction="north" orientation="latitude" pageId="16" pageNumber="118" precision="1" value="4.93345">4.93345</geoCoordinate>
,
<geoCoordinate box="[1197,1314,1480,1502]" degrees="59.99514" direction="west" orientation="longitude" pageId="16" pageNumber="118" precision="1" value="-59.99514">–59.99514</geoCoordinate>
,
<collectorName box="[1324,1486,1480,1502]" pageId="16" pageNumber="118">D. C. Werneke</collectorName>
,
<collectorName box="[820,1017,1509,1531]" pageId="16" pageNumber="118">J. W. Armbruster</collectorName>
,
<collectorName box="[1033,1174,1509,1531]" pageId="16" pageNumber="118">N. K. Lujan</collectorName>
,
<collectorName box="[1191,1285,1510,1531]" pageId="16" pageNumber="118">M. Ram</collectorName>
,
<collectorName box="[1302,1446,1509,1531]" pageId="16" pageNumber="118">D. I. Brooks</collectorName>
,
<date pageId="16" pageNumber="118" value="2016-01-14">
<collectingDate pageId="16" pageNumber="118" value="2016-01-14">14 January 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7846693" ID-Zenodo-Dep="7846693" httpUri="https://zenodo.org/record/7846693/files/figure.png" pageId="16" pageNumber="118" startId="16.[103,131,1331,1351]" targetBox="[103,1491,178,1301]" targetPageId="16" targetType="figure">
<paragraph blockId="16.[103,1491,1330,1377]" pageId="16" pageNumber="118">
FIG. 12. Dorsal, lateral, and ventral views of
<taxonomicName authority="Armbruster &amp; Lujan &amp; Bloom, 2021" authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[546,793,1330,1351]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="wangyapoik" status="new species">
<emphasis bold="true" box="[546,793,1330,1351]" italics="true" pageId="16" pageNumber="118">Characidium wangyapoik</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[807,928,1331,1351]" pageId="16" pageNumber="118" rank="species">new species</taxonomicNameLabel>
, after preservation, CSBD F-3615, holotype, 72.6 mm SL; Kuribrong River population. Photos by J. W. Armbruster.
</paragraph>
</caption>
<subSubSection lastPageId="17" lastPageNumber="119" pageId="16" pageNumber="118" type="diagnosis">
<paragraph blockId="16.[820,1490,1592,2024]" lastBlockId="17.[142,812,179,909]" lastPageId="17" lastPageNumber="119" pageId="16" pageNumber="118">
<emphasis box="[820,1228,1592,1615]" italics="true" pageId="16" pageNumber="118">
<emphasis bold="true" box="[820,954,1592,1615]" italics="true" pageId="16" pageNumber="118">Diagnosis.—</emphasis>
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[954,1228,1592,1614]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="wangyapoik">Characidium wangyapoik</taxonomicName>
</emphasis>
can be distinguished from all crenuchids except
<emphasis box="[1132,1438,1621,1643]" italics="true" pageId="16" pageNumber="118">
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1132,1264,1621,1643]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="crandellii">C. crandellii</taxonomicName>
,
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1277,1434,1621,1643]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="declivirostre">C. declivirostre</taxonomicName>
,
</emphasis>
and
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[820,974,1650,1672]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="duplicatum">
<emphasis box="[820,974,1650,1672]" italics="true" pageId="16" pageNumber="118">C. duplicatum</emphasis>
</taxonomicName>
by having venter without scales from the isthmus to the pelvic girdle (vs. maximally to pectoral insertion) and from most species of
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[1233,1367,1709,1731]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="genus">
<emphasis box="[1233,1367,1709,1731]" italics="true" pageId="16" pageNumber="118">Characidium</emphasis>
</taxonomicName>
by having a very large pectoral fin with first three unbranched pectoral-fin rays thickened and first pectoral-fin ray bent at an oblique angle (vs. pectoral-fin rays not thickened and first pectoral-fin ray either straight or slightly convex).
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="wangyapoik">
<emphasis italics="true" pageId="16" pageNumber="118">Characidium wangyapoik</emphasis>
</taxonomicName>
differs from
<emphasis box="[1096,1404,1855,1877]" italics="true" pageId="16" pageNumber="118">
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1096,1228,1855,1877]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="crandellii">C. crandellii</taxonomicName>
,
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1242,1399,1855,1877]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="declivirostre">C. declivirostre</taxonomicName>
,
</emphasis>
and
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="16" pageNumber="118" phylum="Chordata" rank="species" species="duplicatum">
<emphasis italics="true" pageId="16" pageNumber="118">C. duplicatum</emphasis>
</taxonomicName>
by having at least the ventral region of flank with thin bars becoming contiguous across body at larger sizes (vs. ventral area with almost square blotches), having lateral-line canal in most of scales very short (
<emphasis bold="true" box="[1195,1213,1973,1994]" pageId="16" pageNumber="118">~</emphasis>
25% of scale length), and having canal pores covered by preceding scales (vs. canal at least 33% of scale length and pores not covered by preceding scales); from
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[298,429,208,230]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="119" phylum="Chordata" rank="species" species="crandellii">
<emphasis box="[298,429,208,230]" italics="true" pageId="17" pageNumber="119">C. crandellii</emphasis>
</taxonomicName>
by having 10 circumpeduncular scales (vs. 12), having two or more thin dark bands consisting of spots on rays in dorsal and pectoral fins (vs. a median wide dark band with pigment concentrated on membranes in dorsal fin and pectoral fin either entirely dark or dark with lighter middle), having an almost square dorsal fin with slightly concave distal margin (vs. falcate), having adipose fin located above middle of anal-fin base (vs. anterior edge of adipose fin at or behind vertical through posteriormost anal-fin insertion), by having dentary and premaxillary teeth narrow, peg-like, and maximally tricuspid with only central cusp well developed (vs. wide and tri- to pentacuspid, central cusp longest), by having zero to two (total) teeth in second dentary row (vs. 8 or more); by a preanal length/SL ratio of 77.4–82.3% (vs. 72.9–76.6%), and anal–apex length/SL ratio of 94.6–100.3% (vs. 89.4–94.0%); from
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[613,770,650,672]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="119" phylum="Chordata" rank="species" species="declivirostre">
<emphasis box="[613,770,650,672]" italics="true" pageId="17" pageNumber="119">C. declivirostre</emphasis>
</taxonomicName>
by having the flanks with 10 or more narrow bars (vs. less than 10 almost square blotches); and from
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[552,697,709,731]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="119" phylum="Chordata" rank="species" species="duplicatum">
<emphasis box="[552,697,709,731]" italics="true" pageId="17" pageNumber="119">C. duplicatum</emphasis>
</taxonomicName>
by having one leading unbranched pelvic ray (vs. two).
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="119" phylum="Chordata" rank="species" species="wangyapoik">
<emphasis italics="true" pageId="17" pageNumber="119">Characidium wangyapoik</emphasis>
</taxonomicName>
is most similar in color to
<taxonomicName authorityName="Lujan, Agudelo-Zamora, Taphorn, Booth &amp; Lopez-Fernandez" authorityYear="2013" box="[592,699,768,790]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="119" phylum="Chordata" rank="species" species="amaila">
<emphasis box="[592,699,768,790]" italics="true" pageId="17" pageNumber="119">C. amaila</emphasis>
</taxonomicName>
from the upper Kuribrong River, but it has bands in the dorsal and pectoral fins (vs. no bands in
<taxonomicName authorityName="Lujan, Agudelo-Zamora, Taphorn, Booth &amp; Lopez-Fernandez" authorityYear="2013" box="[468,571,827,849]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="17" pageNumber="119" phylum="Chordata" rank="species" species="amaila">
<emphasis box="[468,571,827,849]" italics="true" pageId="17" pageNumber="119">C. amaila</emphasis>
</taxonomicName>
) and the unbranched pectoral-fin rays are greatly thickened (vs. only slightly thickened).
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="120" pageId="17" pageNumber="119" type="description">
<paragraph blockId="17.[142,812,940,2024]" pageId="17" pageNumber="119">
<emphasis bold="true" box="[142,291,940,963]" italics="true" pageId="17" pageNumber="119">Description.—</emphasis>
Measurements based on
<specimenCount box="[559,706,940,962]" pageId="17" pageNumber="119" type="generic">21 specimens</specimenCount>
(
<tableCitation box="[718,798,940,963]" captionStart="Table 2" captionStartId="13.[142,190,178,198]" captionTargetPageId="13" captionText="Table 2. Morphometrics for Characidium duplicatum, new species (n ¼ 20), and C. wangyapoik, new species (n ¼ 21)." pageId="17" pageNumber="119">Table 2</tableCitation>
); meristics based on
<specimenCount box="[368,522,969,991]" pageId="17" pageNumber="119" type="generic">20 specimens</specimenCount>
(reported below). Dorsal profile of body forms convex arc from tip of snout to posterior end of supraoccipital, then becomes steeper and longer convex arc from supraoccipital to end of dorsal fin (highest point of arc at dorsal-fin origin); dorsal profile relatively straight and ventrally angled from dorsal fin to adipose fin, then forming concave arc to caudal fin. Ventral profile straight to anal fin, then forming concave arc to caudal fin. Body depth greatest at dorsal origin and least at middle of caudal peduncle. Body oval in cross section anteriorly (flattened ventrally) and oval with dorsal and ventral surfaces somewhat flattened on caudal peduncle. Eye diameter 52.5–69.0% snout length, decreasing in size with SL (
<figureCitation box="[149,210,1352,1375]" captionStart="FIG" captionStartId="2.[820,848,1949,1969]" captionTargetBox="[822,1490,1520,1919]" captionTargetId="figure-235@2.[820,1490,1520,1919]" captionTargetPageId="2" captionText="FIG. 3. Plot of the ratio of orbital diameter to head length versus standard length for all measured specimens of fast-water Characidium of the Guiana Shield showing allometry in eye size." figureDoi="http://doi.org/10.5281/zenodo.7846675" httpUri="https://zenodo.org/record/7846675/files/figure.png" pageId="17" pageNumber="119">Fig. 3</figureCitation>
), oval, angled with anterior vertex ventral to posterior vertex; dorsal rim of orbit slightly higher than interorbital surface. Snout broadly rounded. Gill membranes united across isthmus. Tubercles present in both sexes dorsally on head and anterodorsal scales.
</paragraph>
<paragraph blockId="17.[142,812,940,2024]" pageId="17" pageNumber="119">Scales cycloid with most scales having 10–30 short, parallel striae (most on first postdorsal scale). Lateral line complete with canal occupying approximately 1/4 of scale length and pores covered by previous scales; 30 (1), 31 (4), 32 (12), or 33 (3) lateral-line scales; lateral-line scales distinctly smaller anteriorly; naked area between anterior lateral-line scales and pectoral-fin base; lateral line continues onto scales covering caudal base. 4 scales above lateral line and 2 scales below lateral line; 10 circumpeduncular scales. Scales covering anterior 1/5 of caudal fin. 7 (2), 8 (14), or 9 (4) predorsal scales. Venter unscaled on isthmus and posteriorly to 4 to 5 scales anterior to pelvic-fin origin.</paragraph>
<paragraph blockId="17.[142,812,940,2024]" lastBlockId="17.[859,1530,179,1646]" pageId="17" pageNumber="119">
Dorsal fin with 2 unbranched and 8 (1) or 9 (19) branched rays (ii,8–9); first unbranched ray about 1/3 length of second; first unbranched ray longest and last shortest; fin roughly rectangular. Pectoral fin with 3 unbranched and 10 (19) or 11 (1) branched rays (iii,10–11); unbranched rays and first branched ray with thick pads of tissue anteriorly; first unbranched ray strongly curved; first branched ray longest and last shortest; pectoral fin oriented obliquely on body with posteriormost insertion located posterodorsally to origin. Pelvic fin with 1 leading unbranched ray, 6 (19) or 7 (1) branched rays, and 1 posterior unbranched ray (i,6–7,i); first branched ray longest; 2 to 3 pelvic axillary scales present with complex covering
<emphasis bold="true" box="[1118,1136,356,377]" pageId="17" pageNumber="119">~</emphasis>
half of pelvic-fin base. Anal fin with 2 unbranched rays and 5 branched rays (ii,5); first unbranched ray slightly less than 1/3 length of second, closely adhered; fin margin curved with second unbranched ray longest and last branched ray shortest. Caudal fin with 1 unbranched and 8 (4) or 9 (16) branched rays in upper lobe and 8 (16), or 9 (4) branched and 1 unbranched ray in lower lobe (i,8–9,8–9,i); forked with upper and lower lobes coequal. Adipose fin present with base centered on vertical through middle of anal-fin base. Rays of paired, dorsal, and anal fins with thick flaps of skin dorsally (paired fins) and laterally (dorsal and anal fins) that overlap successive rays in adpressed fins; flaps widest and longest anteriorly, decreasing in size and width posteriorly, flaps usually absent on posterior rays.
</paragraph>
<paragraph blockId="17.[859,1530,179,1646]" pageId="17" pageNumber="119">Teeth tricuspid, narrow and peg-like with median cusp round and lateral cusps poorly developed. 6 (4), 7 (12), or 8 (4) premaxillary teeth. 9 (3), 10 (6), 11 (6), 12 (4), or 14 (1) teeth in outer dentary row. Inner dentary row comprising many small, unicuspid teeth; teeth may be poorly visible or absent in smaller specimens. Ectopterygoid with two rows of approximately 10 (lateral) and 5–10 (medial) minute teeth.</paragraph>
<paragraph blockId="17.[859,1530,179,1646]" pageId="17" pageNumber="119">
Branchiostegal rays 4; 1 ray attached to posterior ceratohyal; 3 rays attached to anterior ceratohyal. Gill rakers 2–3 on dorsal limb, 1 on angle, 6–7 on ventral limb of anterior branchial arch. One supraorbital present; crescent shaped; from dorsal midpoint of orbit to dorsal
<emphasis bold="true" box="[1338,1356,1152,1173]" pageId="17" pageNumber="119">~</emphasis>
1/4 of anterior scleral ossicle. All elements of infraorbital series, except infraorbital 1, without laminar component. Parietal branch of supraorbital sensory canal extending less than 1/4 into parietal. Parietal fontanel reduced to tiny triangle at posterior borders of parietals, almost absent in some; smallest specimens with gap between parietals. Frontal foramina above supraorbital canal 3–4 with some posterior foramina sometimes combined, wide, circular.
</paragraph>
<paragraph blockId="17.[859,1530,179,1646]" pageId="17" pageNumber="119">Total number of vertebrae 33 (3). Vertebral centra 2–4 fused, without ventral processes. Rib of centrum 4 distally expanded, extending anteriorly toward lateral process of centrum 2. Posterior chamber of swim bladder extremely reduced, about length of one vertebral centrum. Supraneurals between neural spine of fourth centrum and anterior dorsal-fin pterygiophore 4 (4), 5 (1). Epurals 3 (5). Uroneural present, about 1/2 as long as urostyle.</paragraph>
<paragraph blockId="17.[859,1529,1677,2024]" lastBlockId="18.[103,773,179,1225]" lastPageId="18" lastPageNumber="120" pageId="17" pageNumber="119">
<emphasis bold="true" box="[859,1068,1677,1700]" italics="true" pageId="17" pageNumber="119">Coloration in life.—</emphasis>
(
<figureCitation box="[1075,1150,1677,1700]" captionStart="FIG" captionStartId="15.[142,170,1319,1339]" captionTargetBox="[142,1530,178,1290]" captionTargetId="figure-339@15.[142,1530,178,1290]" captionTargetPageId="15" captionText="FIG. 11. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, in life, CSBD F-3615, holotype, 72.6 mm SL. Photos by N. K. Lujan." figureDoi="http://doi.org/10.5281/zenodo.7846691" httpUri="https://zenodo.org/record/7846691/files/figure.png" pageId="17" pageNumber="119">Fig. 11</figureCitation>
) Base color tan with slight yellow tinge (particularly on head, fins, and dorsally). Dorsal surface with many (
<emphasis bold="true" box="[1001,1019,1737,1758]" pageId="17" pageNumber="119">~</emphasis>
10) diffuse saddles that seem to subdivide in larger specimens. Faint stripe present along lateral line. Numerous narrow bars below lateral line (1–1.5 scale rows wide). These may extend above lateral line and join with dorsal saddles (particularly in larger specimens). Color generally concentrated on scales with each scale having lighter edges, but either anterior or posterior edges may be light. Ventral surface gray with ventral bars almost meeting midventrally. Iridescent yellow-green stripe noticeable in some specimens between midlateral stripe and dorsal surface; stripe located below dark pigment and fades at insertion of posteriormost dorsal-fin ray.
</paragraph>
<paragraph blockId="18.[103,773,179,1225]" pageId="18" pageNumber="120">Head with dorsal saddle that extends to near ventral margin of opercle (posterior margin of opercle gray to yellow). Large dark blotches present below and behind eye. Wide stripe present from eye to snout. Dark bands present between orbits and down snout. Amount of melanin on head varies, some individuals with large gray to yellow patches and some almost entirely dark. Iridescent green spot present in some specimens located dorsal to postorbital dark spot.</paragraph>
<paragraph blockId="18.[103,773,179,1225]" pageId="18" pageNumber="120">
Dorsal fin with dark distal margin, gray band that changes to yellow proximally, then black band made of spots on rays that covers
<emphasis bold="true" box="[238,256,530,551]" pageId="18" pageNumber="120">~</emphasis>
1/4 width of fin (interradial membranes gray along band), then narrow gray to yellow band, and finally proximal black band made of roughly triangular spots with longest edges along anterior margins of rays and sloping posteroventrally to posterior margin of rays (interradial membranes gray to yellow); all bands more diffuse anteriorly. Pectoral fin with gray distal margin changing to yellow proximally; dark spots present in two bands distally; distalmost dark band with spots only on rays, but some spots bleed onto membranes in proximal band; dark band at base of pectoral fin may be present; pectoral-fin colors more diffuse ventrally, generally gray with fleshy pads of unbranched and first branched rays almost white. Pelvic fin colored similarly to pectoral with single median band that includes melanophores on interradial membranes and band at base of fin. Anal fin as pelvic fin, but no basal band. Adipose fin yellow to gray proximally and dark distally with dark color confluent with saddle below it. Base color of caudal fin dark with pigment concentrated at junctions of lepidotrichia; two yellow spots present at base (just above and just below midline), single median spot located along midline just after proximal spots, rest of fin with large yellow blotches proximally and large gray blotches distally (gray blotches may fade into base color).
</paragraph>
<paragraph blockId="18.[103,773,1255,1307]" pageId="18" pageNumber="120">
<emphasis bold="true" box="[103,365,1255,1278]" italics="true" pageId="18" pageNumber="120">Coloration in alcohol.—</emphasis>
(
<figureCitation box="[373,451,1255,1278]" captionStart="FIG" captionStartId="16.[103,131,1331,1351]" captionTargetBox="[103,1491,178,1301]" captionTargetId="figure-376@16.[103,1491,178,1301]" captionTargetPageId="16" captionText="FIG. 12. Dorsal, lateral, and ventral views of Characidium wangyapoik, new species, after preservation, CSBD F-3615, holotype, 72.6 mm SL; Kuribrong River population. Photos by J. W. Armbruster." figureDoi="http://doi.org/10.5281/zenodo.7846693" httpUri="https://zenodo.org/record/7846693/files/figure.png" pageId="18" pageNumber="120">Fig. 12</figureCitation>
) As in life but yellows and grays become tan and iridescence is lost.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="120" type="distribution">
<paragraph blockId="18.[103,773,1338,1565]" pageId="18" pageNumber="120">
<emphasis bold="true" box="[103,258,1338,1361]" italics="true" pageId="18" pageNumber="120">Distribution.—</emphasis>
(
<figureCitation box="[266,327,1338,1361]" captionStart="FIG" captionStartId="9.[1178,1206,791,811]" captionTargetBox="[142,1129,179,912]" captionTargetId="figure-703@9.[142,1129,179,912]" captionTargetPageId="9" captionText="FIG. 9. Distributions of Characidium declivirostre, C. duplicatum, new species, and C. wangyapoik, new species. Type localities indicated by T’s (approximated for C. declivirostre)." figureDoi="http://doi.org/10.5281/zenodo.7846687" httpUri="https://zenodo.org/record/7846687/files/figure.png" pageId="18" pageNumber="120">Fig. 9</figureCitation>
)
<taxonomicName authorityName="Armbruster &amp; Lujan &amp; Bloom" authorityYear="2021" box="[342,606,1338,1360]" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="120" phylum="Chordata" rank="species" species="wangyapoik">
<emphasis box="[342,606,1338,1360]" italics="true" pageId="18" pageNumber="120">Characidium wangyapoik</emphasis>
</taxonomicName>
is only known from the upper Ireng River basin (Amazon River) along the
<collectingCountry box="[103,168,1397,1419]" name="Brazil" pageId="18" pageNumber="120">Brazil</collectingCountry>
/
<collectingCountry box="[176,259,1397,1419]" name="Guyana" pageId="18" pageNumber="120">Guyana</collectingCountry>
border (known as the Rio Mau in
<collectingCountry box="[696,762,1396,1419]" name="Brazil" pageId="18" pageNumber="120">Brazil</collectingCountry>
). Specimens were collected from below Orinduik Falls to the upper falls on the Ireng and its equal tributary, the Sukwabi River, but not above the Uluk Tuwuk or Wotawanda falls of the upper Ireng and Sukwabi Rivers (see
<bibRefCitation author="Lujan, N. K. &amp; J. W. Armbruster &amp; D. C. Werneke &amp; T. Franco Teixeira &amp; N. R. Lovejoy" box="[541,731,1514,1536]" pageId="18" pageNumber="120" pagination="1213 - 1235" refId="ref19030" refString="Lujan, N. K., J. W. Armbruster, D. C. Werneke, T. Franco Teixeira, and N. R. Lovejoy. 2020. Phylogeny and biogeography of the Brazilian-Guiana Shield endemic Corymbophanes clade of armoured catfishes (Loricariidae). Zoological Journal of the Linnean Society 188: 1213 - 1235." type="journal article" year="2020">Lujan et al., 2020</bibRefCitation>
, for a more detailed map and description of this area).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="120" type="etymology">
<paragraph blockId="18.[103,773,1596,1735]" pageId="18" pageNumber="120">
<emphasis box="[103,379,1596,1619]" italics="true" pageId="18" pageNumber="120">
<emphasis bold="true" box="[103,248,1596,1619]" italics="true" pageId="18" pageNumber="120">Etymology.—</emphasis>
Wangyapoik
</emphasis>
is the Patamona word for the species, and it is used as a noun in apposition.
<emphasis box="[632,693,1626,1647]" italics="true" pageId="18" pageNumber="120">Wang</emphasis>
means ‘honey’ and
<emphasis box="[249,316,1654,1676]" italics="true" pageId="18" pageNumber="120">yapoik</emphasis>
means ‘seated,’ perhaps in reference to the yellowish color. The Patamona also refer to the species by the English common name of ‘‘fallsfish.’’
</paragraph>
</subSubSection>
</treatment>
</document>
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