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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2011" docId="4C90D5C38BBF0374B097285C205F23E6" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha coarctata De Saussure 1860" docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="10" updateTime="1668152297771" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>131</mods:number>
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<mods:start>1</mods:start>
<mods:end>161</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3502753" ID-GBIF-Taxon="152031137" ID-Zenodo-Dep="3502753" LSID="urn:lsid:plazi:treatment:4C90D5C38BBF0374B097285C205F23E6" httpUri="http://treatment.plazi.org/id/4C90D5C38BBF0374B097285C205F23E6" lastPageId="14" lastPageNumber="15" pageId="9" pageNumber="10">
<subSubSection pageId="9" pageNumber="10" type="nomenclature">
<paragraph pageId="9" pageNumber="10">
<taxonomicName LSID="http://species-id.net/wiki/Orthomorpha_coarctata" authority="De Saussure, 1860" authorityName="De Saussure" authorityYear="1860" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata (De Saussure, 1860)</taxonomicName>
Figs 48
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="reference_group">
<paragraph pageId="9" pageNumber="10">
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus coarctatus" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coarctatus">Polydesmus coarctatus</taxonomicName>
<bibRefCitation author="De Saussure, H" journalOrPublisher="Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve" pageId="114" pageNumber="115" pagination="259 - 394" title="Essai d'une faune de Myriapodes du Mexique, avec la description de quelques especes des autres parties de l'Amerique." volume="15" year="1860">De Saussure 1860</bibRefCitation>
: 297 (D).
</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Paradesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paradesmus flavocarinatus" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="flavocarinatus">Paradesmus flavocarinatus</taxonomicName>
<bibRefCitation author="Daday, E" journalOrPublisher="Termeszetrajzi Fuezetek" pageId="114" pageNumber="115" pagination="115 - 156" title="Myriopoda extranea Musaei nationalis Hungarici." volume="12" year="1889">Daday 1889</bibRefCitation>
: 136 (D).
</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
-
<bibRefCitation pageId="9" pageNumber="10">Attems 1937</bibRefCitation>
: 62 (D);
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
: 45 (M) et auctorum.
</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata subsp. gigas" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="coarctata" subSpecies="gigas">Orthomorpha coarctata gigas</taxonomicName>
<bibRefCitation author="Attems, C" journalOrPublisher="Zooelogische Mededeelingen" pageId="114" pageNumber="115" pagination="61 - 70" title="Myriopoden von Ambon und anderen Inseln der Banda-See." volume="10" year="1927">Attems 1927</bibRefCitation>
: 63 (D).
</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Asiomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asiomorpha coarctata" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="coarctata">Asiomorpha coarctata</taxonomicName>
-
<bibRefCitation pageId="9" pageNumber="10">Verhoeff 1939</bibRefCitation>
: 117 (D) et auctorum.
</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata subsp. gigas" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="coarctata" subSpecies="gigas">Orthomorpha coarctata gigas</taxonomicName>
-
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
: 45 (M);
<bibRefCitation pageId="9" pageNumber="10">Golovatch 1998</bibRefCitation>
: 43 (D) et auctorum.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10" type="material examined">
<paragraph pageId="9" pageNumber="10">Material examined.</paragraph>
<paragraph lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10">
1 ♂, 2 ♀(CUMZ), Thailand, Chiang Mai Prov., Mae Rim Distr., Queen Sirikit Botanical Garden,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="19.504723">19°30'17&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="99.44138">99°25'89&quot;E</geoCoordinate>
, 03.03.2007, leg. N. Likhitrakarn. 1 ♀ (CUMZ), Nan Prov., Mueang Nan Distr., Nan Resort, ca 240 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="18.905832">18°54'21&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.7625">100°45'45&quot;E</geoCoordinate>
, 11.10.2009, leg. N. Likhitrakarn. 1 ♂, 1 ♀ (CUMZ), same Prov., Pua Distr., Wora Nakhom Subdistr., ca 270 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="19.92861">19°55'43&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.920555">100°55'14&quot;E</geoCoordinate>
, 29.01.2010, leg. R. Chanabun. 1 ♂ (CUMZ), Phrae Prov., Rong Kwang Distr., Hoylong Waterfall,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="18.744167">18°44'39&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.75972">100°44'95&quot;E</geoCoordinate>
, 09.10.2009, leg. R. Chanabun. 1 ♂ (CUMZ), Phitsanulok Prov., Wang Thong Distr., Sukunotayan Waterfall,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="17.40361">17°23'73&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.88472">100°53'5&quot;E</geoCoordinate>
, 09.10.2009, leg. N. Likhitrakarn. 3 ♂ (CUMZ), Tak Prov., Tha Song Yong Distr., Mae Usu Cave, ca 140 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="17.304443">17°18'16&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="98.15584">98°09'21&quot;E</geoCoordinate>
, 30.05.2009, leg.
<pageBreakToken pageId="10" pageNumber="11" start="start">N</pageBreakToken>
. Likhitrakarn. 1 ♀ (CUMZ), same Prov., Umphang Distr., Thee Lor Su Riverside Resort, ca 470 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.046389">16°02'47&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="98.8525">98°51'9&quot;E</geoCoordinate>
, 06.06.2009, leg. R. Chanabun. 7 ♂, 5 ♀ (CUMZ), Kamphaeng Phet Prov., Mueang Kamphaeng Phet Distr., Thai Restaurant, 17.01.2011, leg. N. Likhitrakarn &amp; R. Chanabun. 1 ♀ (CUMZ), Ubon Ratchathani Prov., Khongchiam Distr., Tadton Waterfall, 14.05.2011, leg. R. Chanabun. 1 ♀ (CUMZ), Saraburi Prov., Chaloem Phra Kiat Distr., Siwilai Cave,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="15.200833">15°12'03&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.453606">101°27'13&quot;E</geoCoordinate>
, 31.09.2006, leg. R. Chanabun. 2 ♂, 1 ♀ (CUMZ), Phra Nakhon Si Ayutthaya Prov., Mueang Phra Nakhon Si Ayutthaya Distr., 15.11.2009, leg. C. Sutcharit. 1 ♂ (CUMZ), same Prov., Maharat Distr., Seafood Restaurant, 20.09.2009,
<pageBreakToken pageId="11" pageNumber="12" start="start">leg</pageBreakToken>
. N. Likhitrakarn. 1 ♂ (CUMZ), Nakhon Nayok Prov., Ban Na Distr., near house, 17.05.1952, leg. K. Isarankura. 1 ♂ (CUMZ), Sa Kaeo Prov., Khlong Hat Distr., Thamphet Temple, ca 170 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="13.354167">13°21'15&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="102.31306">102°18'47&quot;E</geoCoordinate>
, 17.09.2009, leg. R. Chanabun. 1 ♂, 1 ♀ (CUMZ), Bangkok Prov., Pattum Wan Distr., Chulalongkorn University campus,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="14.183333">14°11'0&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.88389">100°53'02&quot;E</geoCoordinate>
, 03.07.2009, leg. N. Likhitrakarn. 1 ♂ (CUMZ), Phetchaburi Prov., Ban Laem Distr., 20.07.2009, leg. R. Chanabun. 1 ♂ (CUMZ), Chonburi Prov., Si Racha Distr., Khao Kheow Open Zoo,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="13.365001">13°21'54&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.08806">101°05'17&quot;E</geoCoordinate>
, 03.04.2007, leg. R. Chanabun. 1 ♀ (CUMZ), same Prov., Bothong Distr., Bothong Waterfall, ca 90 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="13.2502775">13°15'01&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.37583">101°22'33&quot;E</geoCoordinate>
, 15.09.2009, leg. R. Chanabun. 1 ♀ (CUMZ), Chanthaburi Prov., Khlung Distr., Makok Waterfall,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="12.987778">12°59'16&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="102.43389">102°26'02&quot;E</geoCoordinate>
, 03.09.2007, leg. C. Sutcharit. 1 ♀ (CUMZ), same Prov.,
<pageBreakToken pageId="12" pageNumber="13" start="start">Khaeng</pageBreakToken>
Hang Maeo Distr., Khao Sibhachan,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="13.56">13°33'36&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="102.196945">102°11'49&quot;E</geoCoordinate>
, 30.12.2008, leg. N. Likhitrakarn. 1 ♀ (CUMZ), Ranong Prov., Mueang Ranong Distr., 06.10.2008, leg. C. Sutcharit. 8 ♂, 13 ♀, 3 juv. (CUMZ), Phang Nga Prov., Khura Buri Distr., Similan National Park, Ko Similan, Island 8,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="8.6669445">8°40'01&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="97.64833">97°38'54&quot;E</geoCoordinate>
, 07.04.2010, leg. S. Panha &amp; N. Likhitrakarn. 1 ♂ (CUMZ), Surat Thani Prov., Ban Ta Khun Distr., Ratchaprapa Dam,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="9.622222">9°37'20&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="99.35">99°21'0&quot;E</geoCoordinate>
, 08.10.2008, leg. N. Likhitrakarn. 2 ♀ (CUMZ), Phatthalung Prov., Mueang Phatthalung Distr., Tham Malai Thep Nimit, ca 20 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="7.6358333">7°38'09&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.08445">100°05'04&quot;E</geoCoordinate>
, 11.01.2009, leg. R. Chanabun. 2 ♀ (CUMZ), Satun Prov., Mueang Satun Distr., Tarutao National Park, Ao Talo Wow,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="7.0416665">7°02'30&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.138885">100°08'20&quot;E</geoCoordinate>
, 09.04.2008, leg. R. Chanabun. 1 ♀ (CUMZ), Malaysia, Terengganu, Kenyir Lake, ca 170 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.845556">4°50'44&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="102.72083">102°43'15&quot;E</geoCoordinate>
, 25.05.2011, leg. R. Chanabun. 7
<pageBreakToken pageId="13" pageNumber="14" start="start"></pageBreakToken>
, 18 ♀ (CUMZ), Johor, Yong Peng, ca 20 m,
<geoCoordinate direction="north" orientation="latitude" precision="15" value="2.011111">2°0'40&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="103.056946">103°03'25&quot;E</geoCoordinate>
, 21.05.2011, leg. R. Chanabun. Syntypes of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata subsp. gigas" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="coarctata" subSpecies="gigas">Orthomorpha coarctata gigas</taxonomicName>
: 1 ♂, 1 ♀ (NHMW-3504), Indonesia, Maluku Prov., Banda Sea, Maluku Barat Regency, Poelau Island, Teoen (= Teun), leg. Kopstein.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="descriptive notes">
<paragraph pageId="13" pageNumber="14">Descriptive notes.</paragraph>
<paragraph lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Length 14.5-20.5 (♂) or 16.5-27.5 mm (♀), width of midbody pro- and metazona 1.1-1.7 and 1.5-2.7 mm (♂), 1.1-2.5 and 1.6-3.2 mm (♀), respectively.Coloration, texture, all main somatic and gonopod characters as in Figs 4-8.
<pageBreakToken pageId="14" pageNumber="15" start="start">Neither</pageBreakToken>
sternal lobe nor cone(s) between ♂ coxae 4 (Fig. 4H &amp; I), at most only traces of poor knobs (Fig. 7H &amp; I).
</paragraph>
<paragraph pageId="14" pageNumber="15">Gonopods (Figs 5, 6 &amp; 8) with tip of solenophore produced into a single lobe, all other spikes or denticles being either missing or nearly missing.</paragraph>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
Figure 4.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
(De Saussure, 1860), ♂ (
<normalizedToken originalValue="AI">A-I</normalizedToken>
) and ♀ (-L) from Thai Restaurant. A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D, K segments 10 and 11, dorsal, lateral and dorsal views, respectively E-G, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
Figure 5.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
(De Saussure, 1860), ♂ from Chulalongkorn University campus. A, B right gonopod, mesal and lateral views, respectively C-F distal part of right gonopod, mesal, lateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
Figure 6.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
(De Saussure, 1860), ♂ from Mueang Kamphaeng Phet Distr., Thai Restaurant. A, B right gonopod, lateral and mesal views, respectively.Scale bar: 0.2 mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
Figure 7.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
(De Saussure, 1860), ♂ (
<normalizedToken originalValue="AI">A-I</normalizedToken>
) and ♀ (
<normalizedToken originalValue="JL">J-L</normalizedToken>
) syntype of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata subsp. gigas" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="coarctata" subSpecies="gigas">Orthomorpha coarctata gigas</taxonomicName>
Attems, 1927. A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D, K segments 10 and 11, dorsal, lateral and dorsal views, respectively
<normalizedToken originalValue="EG">E-G</normalizedToken>
, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
</paragraph>
</caption>
<caption pageId="14" pageNumber="15">
<paragraph pageId="14" pageNumber="15">
Figure 8.
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
(De Saussure, 1860), ♂ syntype of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata subsp. gigas" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="coarctata" subSpecies="gigas">Orthomorpha coarctata gigas</taxonomicName>
Attems, 1927. A, B right gonopod, lateral and mesal views, respectively.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="14" pageNumber="15" type="remarks">
<paragraph pageId="14" pageNumber="15">Remarks.</paragraph>
<paragraph pageId="14" pageNumber="15">
This pantropical species has been described and redescribed several times, often under different names (e.g.
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
). From Thailand, it had been described as
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Paradesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paradesmus flavocarinatus" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="flavocarinatus">Paradesmus flavocarinatus</taxonomicName>
until
<bibRefCitation author="Enghoff, H" journalOrPublisher="Steenstrupia" pageId="114" pageNumber="115" pagination="87 - 103" title="The millipedes of Thailand (Diplopoda)." volume="29" year="2005">Enghoff (2005)</bibRefCitation>
showed it to actually represent still another synonym of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
. No full catalogue is attempted here as being superfluous, with hundreds of references involved.
</paragraph>
<paragraph pageId="14" pageNumber="15">
Our restudy of the type material of
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata subsp. gigas" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="coarctata" subSpecies="gigas">Orthomorpha coarctata gigas</taxonomicName>
, kept in NHMW (Figs 7 &amp; 8), confirms
<normalizedToken originalValue="Jeekels">Jeekel's</normalizedToken>
(1968) synonymization of this subspecies with
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="coarctata">Orthomorpha coarctata</taxonomicName>
s. str.
</paragraph>
<subSection pageId="14" pageNumber="15" type="species with a bifid gonopod tip, with only a terminal and a subterminal denticle or lobule">
<paragraph pageId="14" pageNumber="15">Species with a bifid gonopod tip, with only a terminal and a subterminal denticle or lobule</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>