305 lines
31 KiB
XML
305 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.94.26859" ID-GBIF-Dataset="30d1d279-f033-4868-b0c9-925a406be2c5" ID-GBIF-Taxon="182236657" ID-Pensoft-Pub="1860-0743-2-369" ID-Pensoft-UUID="83D45F0665FE5C729C47210ACD710944" ID-Zenodo-Dep="1344764" ID-ZooBank="C354D7DF198F4D4FA2A8F763FD03018D" ModsDocID="1860-0743-94-2-369" checkinTime="1621423395062" checkinUser="pensoft" docAuthor="Song, Zhi-Shun, Malenovsky, Igor, Chen, Jian-Qin, Deckert, Juergen & Liang, Ai-Ping" docDate="2018" docId="4C5920DEB5A1B6B1D93DC2B717C24DE3" docLanguage="en" docName="ZoosystEvol 94(2): 369-391" docOrigin="Zoosystematics and Evolution 94 (2)" docPubDate="2018-08-06" docSource="http://dx.doi.org/10.3897/zse.94.26859" docTitle="Orthopagus philippinus Melichar 1914" docType="treatment" docVersion="3" id="83D45F0665FE5C729C47210ACD710944" lastPageNumber="377" masterDocId="83D45F0665FE5C729C47210ACD710944" masterDocTitle="Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species" masterLastPageNumber="391" masterPageNumber="369" pageId="8" pageNumber="377" updateTime="1632016297512" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Song, Zhi-Shun</mods:namePart>
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<mods:affiliation>Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry & Chemical Engineering, Ji & angsu Second Normal University, Nanjing 210013, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Malenovsky, Igor</mods:namePart>
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<mods:affiliation>Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
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<mods:nameIdentifier type="email">imalenovsky@mzm.cz</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chen, Jian-Qin</mods:namePart>
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<mods:affiliation>Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry & Chemical Engineering, Ji</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Deckert, Juergen</mods:namePart>
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<mods:affiliation>Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlarska 2, 611 37 Brno, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liang, Ai-Ping</mods:namePart>
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<mods:affiliation>Museum fuer Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity Science, Invalidenstrasse 43, Berlin 10115, Germany & Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">liangap@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-08-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>94</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:extent unit="page">
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<mods:start>369</mods:start>
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<mods:end>391</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.94.26859</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.94.26859</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-369</mods:identifier>
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<mods:identifier type="ZooBank">C354D7DF198F4D4FA2A8F763FD03018D</mods:identifier>
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<mods:identifier type="Pensoft-UUID">83D45F0665FE5C729C47210ACD710944</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1344764</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182236657" LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3" httpUri="http://treatment.plazi.org/id/4C5920DEB5A1B6B1D93DC2B717C24DE3" lastPageNumber="377" pageId="8" pageNumber="377">
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<subSubSection pageId="8" pageNumber="377" type="nomenclature">
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<paragraph pageId="8" pageNumber="377">
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<taxonomicName authority="Melichar, 1914" authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus Melichar, 1914</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="8" pageNumber="377">Figs 2I-J</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="8" pageNumber="377">, 3E</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">, 9A-J</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="377" type="reference_group">
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<paragraph pageId="8" pageNumber="377">
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<taxonomicName authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus</taxonomicName>
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Melichar, 1914: 173, Pl. I, figs 1, 2. Lectotype (here designated) male, Los
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<normalizedToken originalValue="Baños">Banos</normalizedToken>
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, Philippines (MMBC, examined).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="377" type="type material">
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<paragraph pageId="8" pageNumber="377">Type material examined.</paragraph>
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<paragraph pageId="8" pageNumber="377">
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Lectotype male (here designated), (1) Los Banos, P.I. Baker; (2) 1311; (3)
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<taxonomicName authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">
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<emphasis italics="true" pageId="8" pageNumber="377">Orthopagus philippinus</emphasis>
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</taxonomicName>
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[
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<normalizedToken originalValue="Melichar’s">Melichar's</normalizedToken>
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handwriting] det. Melichar; (4) Collectio Dr. L. Melichar,
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<normalizedToken originalValue="Moravské">Moravske</normalizedToken>
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museum Brno; (5)
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<taxonomicName authorityName="Uhler" authorityYear="1897" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="genus">Orthopagus</taxonomicName>
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male
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<taxonomicName authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus</taxonomicName>
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sp.n., L. Melichar, 1914 [
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<normalizedToken originalValue="Lauterer’s">Lauterer's</normalizedToken>
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handwriting], P. Lauterer det. 1991; (6) Syn-typus [red label]; (7) Invent.
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<normalizedToken originalValue="č">c</normalizedToken>
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. 4954/Ent., Mor. muzeum, Brno; (8) Lectotypus male,
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<taxonomicName authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">
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<emphasis italics="true" pageId="8" pageNumber="377">Orthopagus philippinus</emphasis>
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</taxonomicName>
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Melichar, 1914, designated by Z. S. Song & I.
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<normalizedToken originalValue="Malenovský">Malenovsky</normalizedToken>
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, 2018 [newly added red label] (MMBC).
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</paragraph>
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<paragraph pageId="8" pageNumber="377">
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Paralectotypes, 4 females, same locality labels as holotype but Inv. nos 4949-4951, 4953;1 female, Mt. Makiling, Luzon, Baker (MMBC, Inv. no. 4952, all paralectotypes bearing the following label: Paralectotypus female,
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<taxonomicName authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">Orthopagus philippinus</taxonomicName>
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Melichar, 1914, designated by Z. S. Song & I.
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<normalizedToken originalValue="Malenovský">Malenovsky</normalizedToken>
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, 2018).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="377" type="materials_examined">
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<paragraph pageId="8" pageNumber="377">Other material examined.</paragraph>
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<paragraph pageId="8" pageNumber="377">
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<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" country="PHILIPPINES" location="Luzon island" specimenCount="1">
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<emphasis bold="true" pageId="8" pageNumber="377">
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<collectingCountry name="Philippines">PHILIPPINES</collectingCountry>
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</emphasis>
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:
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<location LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3:253D4CDEFB81B050B88DA5A9971A9CF2" country="PHILIPPINES" name="Luzon island">Luzon island</location>
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</materialsCitation>
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:
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<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="Baker" country="Philippines" location="Mt. Makiling" specimenCount="1" specimenCount-male="1">
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<specimenCount type="male">1 male</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3:E654AB8B4E78D6DB1F82D60777A383D1" country="Philippines" name="Mt. Makiling">Mt. Makiling</location>
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,
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<collectorName>Baker</collectorName>
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leg. (USNM)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1913-12-11" collectorName="D. T. Fullaway" country="Philippines" location="Los Banos" specimenCount="1" specimenCount-male="1">
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<specimenCount type="male">1 male</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3:D336589435FE2FCBA17795EA78EFAC75" country="Philippines" name="Los Banos">
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Los
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<normalizedToken originalValue="Baños">Banos</normalizedToken>
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</location>
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,
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<collectingDate value="1913-12-11">11.xii.1913</collectingDate>
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,
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<collectorName>D. T. Fullaway</collectorName>
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leg. (BPBM)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1913-01" collectorName="P. Ledyard" country="Philippines" location="Los Banos" specimenCount="1" specimenCount-male="1">
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<specimenCount type="male">1 male</specimenCount>
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,
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<location LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3:1157D89D8D914952451161102595AFEA" country="Philippines" name="Los Banos">
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Los
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<normalizedToken originalValue="Baños">Banos</normalizedToken>
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</location>
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,
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<collectingDate value="1913-01">i.1913</collectingDate>
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,
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<collectorName>P. Ledyard</collectorName>
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leg (LBOB)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1965-03-06" collectingDateMax="1965-03-17" collectingDateMin="1965-03-06" collectorName="H. M. Torrevillas" country="Philippines" elevation="175" location="Wa-wa Dam" municipality="Mt. Montalban" specimenCount="2" specimenCount-male="2" stateProvince="Rizal">
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<specimenCount type="male">2 males</specimenCount>
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,
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<collectingMunicipality>Mt. Montalban</collectingMunicipality>
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,
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<collectingRegion country="Philippines" name="Rizal">Rizal</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3:6614E6F4FFB105F211742DCA6D48965B" country="Philippines" municipality="Mt. Montalban" name="Wa-wa Dam" stateProvince="Rizal">Wa-wa Dam</location>
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||
,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150-200 m</elevation>
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</quantity>
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,
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<collectingDate value="1965-03-06" valueMax="1965-03-17" valueMin="1965-03-06">6. and 17.iii.1965</collectingDate>
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,
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<collectorName>H. M. Torrevillas</collectorName>
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leg. (BPBM)
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||
</materialsCitation>
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||
;
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<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="G. Boettcher" country="Philippines" location="Manila" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Manila">
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<specimenCount type="male">1 male</specimenCount>
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,
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<specimenCount type="female">4 females</specimenCount>
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,
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<collectingRegion country="Philippines" name="Manila">Manila</collectingRegion>
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,
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<collectorName>G. Boettcher</collectorName>
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leg. (MMBC)
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</materialsCitation>
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;
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<specimenCount type="male">1 male</specimenCount>
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,
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<specimenCount type="female">2 females</specimenCount>
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, "B. M. 1925-491", E. M. Ledyard leg.;
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<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="C. S. Banks" country="Philippines" location="Bu. of Sci." municipality="Lot" specimenCount="1" specimenCount-female="1">
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<specimenCount type="female">1 female</specimenCount>
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, "Acc. No. 6625,
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<collectingMunicipality>Lot</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3:F9008B6C6AD8DCC1D2CFDD6557B12E42" country="Philippines" municipality="Lot" name="Bu. of Sci.">Bu. of Sci.</location>
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||
, P. I.,1908-228",
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<collectorName>C. S. Banks</collectorName>
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leg.
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</materialsCitation>
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||
;
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<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="C. S. Banks" country="Philippines" location="Lot" specimenCount="1" specimenCount-male="1">
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||
<specimenCount type="male">1 male</specimenCount>
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, "Acc. No. 5364,
|
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<location LSID="urn:lsid:plazi:treatment:4C5920DEB5A1B6B1D93DC2B717C24DE3:2417BA98015E2F11DA82F0F58407F6D2" country="Philippines" name="Lot">Lot</location>
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, Govt. Lab. Coll., 1908-228",
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<collectorName>C. S. Banks</collectorName>
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leg.
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</materialsCitation>
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(all BMNH).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="377" type="redescription">
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<paragraph pageId="8" pageNumber="377">Redescription.</paragraph>
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<paragraph pageId="8" pageNumber="377">Measurements (4 males, 9 females). Body length (from apex of head to tip of forewings): male 10.6-11.4 mm, female 12.9-13.9 mm; head length (from apex of cephalic process to base of eyes): male 1.25-1.33 mm, female 1.35-1.45 mm; head width (including eyes): male 1.30-1.43 mm, female 1.50-1.60 mm; forewing length: male 8.5-9.1 mm, female 10.3-11.4 mm.</paragraph>
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<paragraph pageId="8" pageNumber="377">
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Coloration. General coloration as in generic description (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="8" pageNumber="377">2I-J</figureCitation>
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). Vertex predominantly dark brown, lateral margins with a pale ochraceous S-shaped streak in posterior three quarters (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9A</figureCitation>
|
||
). Frons light ochraceous with small dark brown spots along intermediate and lateral carinae and a large roundish dark brown spot at base (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9C</figureCitation>
|
||
). Forewing membrane pattern as in Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="8" pageNumber="377">3E</figureCitation>
|
||
. Hind wing membrane with a narrow dark brown infuscation along the apical portion of CuA1 vein, not extending along hind wing apical margin.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="377">
|
||
Structure. Head with cephalic process moderately elongate (Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9A-B</figureCitation>
|
||
). Vertex (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9A</figureCitation>
|
||
) with ratio of length at midline to width between eyes 1.6-1.8. Transition of vertex to frons blunt, broadly rounded in lateral view, cephalic process thus appearing inflated and bulbous apically (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9B</figureCitation>
|
||
). Frons relatively narrow, with ratio of length at midline to maximum width 2.7-3.1.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="377">
|
||
Male genitalia. Pygofer, in lateral view, with dorso-posterior margin produced into a relatively large but narrow, tooth-like, apically obtuse process (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9E</figureCitation>
|
||
); in ventral view (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9F</figureCitation>
|
||
) much longer than in dorsal view (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9D</figureCitation>
|
||
) with ratio of ventral to dorsal width about 2.5. Gonostyles (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9G</figureCitation>
|
||
) large, broad medially, with dorsal margin weakly sinuate. Aedeagus (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9H-J</figureCitation>
|
||
) with endosomal processes covered with minute superficial spines, extended posteriad and strongly curved laterad and dorso-anteriad but not reaching the base of phallobase; phallobase with one pair of large, strongly inflated dorsolateral lobes, their apex convergent and tapering posteriad (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9I</figureCitation>
|
||
), and one pair of smaller, thumb-like ventral lobes, directed ventro-posteriad (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9I-J</figureCitation>
|
||
). Segment X, in lateral view, relatively narrow basally, widening to apex beyond middle, apex blunt (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9E</figureCitation>
|
||
); in dorsal view, relatively large and broad, widest in apical third, ratio of length to maximum width 1.0-1.1 (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Orthopagus philippinus Melichar. (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, caudal view. (G) Gonostyle; (H) Aedeagus, dorsal view. (I) Aedeagus, lateral view. (J) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure9" httpUri="https://binary.pensoft.net/fig/221406" pageId="8" pageNumber="377">9D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="377">Female genitalia as in generic description.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="377" type="distribution">
|
||
<paragraph pageId="8" pageNumber="377">Distribution.</paragraph>
|
||
<paragraph pageId="8" pageNumber="377">So far only known from the Luzon island, Philippines.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="377" type="remarks">
|
||
<paragraph pageId="8" pageNumber="377">Remarks.</paragraph>
|
||
<paragraph pageId="8" pageNumber="377">
|
||
<taxonomicName authorityName="Melichar" authorityYear="1914" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus philippinus" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="philippinus">
|
||
<emphasis italics="true" pageId="8" pageNumber="377">Orthopagus philippinus</emphasis>
|
||
</taxonomicName>
|
||
can be easily recognized from other species of the genus by the shape of the head and the presence of a relatively large roundish dark spot at base of frons which is present in all specimens studied.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="377">
|
||
<bibRefCitation author="Melichar, L" journalOrPublisher="Philippine Journal of Science" pageId="12" pageNumber="381" pagination="173 - 181" refId="B22" refString="Melichar, L, 1914. Neue Homopteren von den Philippinen. Philippine Journal of Science 9: 173 - 181" title="Neue Homopteren von den Philippinen." volume="9" year="1914">Melichar (1914)</bibRefCitation>
|
||
indicated that he described this species based on five (male and female) specimens from "Luzon, Los
|
||
<normalizedToken originalValue="Baños">Banos</normalizedToken>
|
||
, Mt. Maquiling (C. F. Baker)". Six specimens (1 male, 5 females) preserved in Melichar personal collection in MMBC and labelled as either from Los
|
||
<normalizedToken originalValue="Baños">Banos</normalizedToken>
|
||
or Mt. Makiling and collected by Baker are considered here as the original syntypes. The male specimen is designated here as the lectotype for
|
||
<taxonomicName genus="O." lsidName="O. philippinus" pageId="8" pageNumber="377" rank="species" species="philippinus">
|
||
<emphasis italics="true" pageId="8" pageNumber="377">O. philippinus</emphasis>
|
||
</taxonomicName>
|
||
to stabilize the nomenclature according to Article 74 of
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.50608" author="ICZN" journalOrPublisher="The International Trust for Zoological Nomenclature c / o the Natural History Museum, London" pageId="12" pageNumber="381" refId="B10" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature c/o the Natural History Museum, London, DOI: https://doi.org/10.5962/bhl.title.50608" title="International Code of Zoological Nomenclature. Fourth Edition." url="https://doi.org/10.5962/bhl.title.50608" year="1999">ICZN (1999)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |