treatments-xml/data/03/F2/DE/03F2DE0CA93FC025FF5B37C0FE2CE28A.xml
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<document ID-DOI="10.15560/16.4.821" ID-ISSN="1809-127X" approvalRequired="71" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="60" approvalRequired_for_treatments="2" checkinTime="1630327227302" checkinUser="marcus" docAuthor="Cárdenas-Bautista, Johann Stephens, Parada-Alfonso, Jenny Andrea &amp; CarvajalCogollo, Juan E." docDate="2020" docId="03F2DE0CA93FC025FF5B37C0FE2CE28A" docLanguage="en" docName="CheckList.2020.16.4.821-839.pdf" docOrigin="Check List 16 (4)" docSource="http://dx.doi.org/10.15560/16.4.821" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Dichotomius nisus" docType="treatment" docVersion="1" lastPageNumber="831" masterDocId="FFCBA674A935C02FFFF33644FFE2E676" masterDocTitle="Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia" masterLastPageNumber="839" masterPageNumber="821" pageId="10" pageNumber="831" updateTime="1630327251151" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cárdenas-Bautista, Johann Stephens</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parada-Alfonso, Jenny Andrea</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CarvajalCogollo, Juan E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-07-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>821</mods:start>
<mods:end>839</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/16.4.821</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/16.4.821</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F2DE0CA93FC025FF5B37C0FE2CE28A" httpUri="http://treatment.plazi.org/id/03F2DE0CA93FC025FF5B37C0FE2CE28A" lastPageNumber="831" pageId="10" pageNumber="831">
<subSubSection box="[168,545,388,412]" pageId="10" pageNumber="831" type="nomenclature">
<paragraph blockId="10.[168,545,388,412]" box="[168,545,388,412]" pageId="10" pageNumber="831">
<heading box="[168,545,388,412]" fontSize="10" level="2" pageId="10" pageNumber="831" reason="2">
<taxonomicName authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[168,545,388,412]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="831" phylum="Arthropoda" rank="species" species="nisus">
<emphasis bold="true" box="[168,545,388,412]" inLineHeading="true" pageId="10" pageNumber="831" reason="1">
<emphasis bold="true" box="[168,377,388,412]" italics="true" pageId="10" pageNumber="831">Dichotomius nisus</emphasis>
(Olivier, 1789)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="831" type="materials_examined">
<paragraph blockId="10.[167,775,430,723]" pageId="10" pageNumber="831">
<emphasis bold="true" box="[168,412,430,454]" pageId="10" pageNumber="831">Materials examined.</emphasis>
<materialsCitation collectingDate="2019-02-15" collectorName="Vereda El Carmen &amp; Cardenas Johann &amp; Parada Jenny &amp; Crplt" country="Colombia" county="Villavicencio" elevation="648" latitude="4.139364" location="Farm" longLatPrecision="1" longitude="-73.66134" municipality="Vereda El Carmen" pageId="10" pageNumber="831" specimenCount="1" specimenCount-female="1" stateProvince="Meta">
<collectingCountry box="[427,570,430,454]" name="Colombia" pageId="10" pageNumber="831">COLOMBIA</collectingCountry>
<specimenCount box="[608,638,430,454]" pageId="10" pageNumber="831" type="female">1♀</specimenCount>
,
<quantity box="[660,774,430,454]" metricMagnitude="-2" metricUnit="m" metricValue="2.342" pageId="10" pageNumber="831" unit="mm" value="23.42">23.42 mm</quantity>
(mean);
<collectingRegion box="[260,318,464,487]" country="Colombia" name="Meta" pageId="10" pageNumber="831">Meta</collectingRegion>
,
<collectingCounty box="[334,481,464,488]" pageId="10" pageNumber="831">Villavicencio</collectingCounty>
,
<collectorName box="[496,709,464,488]" pageId="10" pageNumber="831">
<collectingMunicipality box="[496,709,464,488]" pageId="10" pageNumber="831">Vereda El Carmen</collectingMunicipality>
</collectorName>
, “Las Pesebreras”
<location LSID="urn:lsid:plazi:treatment:03F2DE0CA93FC025FF5B37C0FE2CE28A:8E8439C1A93FC025FEC337B6FE8EE47F" box="[304,364,498,521]" country="Colombia" county="Villavicencio" latitude="4.139364" longLatPrecision="1" longitude="-73.66134" municipality="Vereda El Carmen" name="Farm" pageId="10" pageNumber="831" stateProvince="Meta">Farm</location>
;
<geoCoordinate box="[377,534,496,521]" degrees="04" direction="north" minutes="08" orientation="latitude" pageId="10" pageNumber="831" precision="1" seconds="21.71" value="4.139364">04°0821.71″N</geoCoordinate>
,
<geoCoordinate box="[547,722,496,521]" degrees="073" direction="west" minutes="39" orientation="longitude" pageId="10" pageNumber="831" precision="1" seconds="40.81" value="-73.66134">073°3940.81″W</geoCoordinate>
;
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.4799999999999995" pageId="10" pageNumber="831" unit="m" value="648.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.4799999999999995" pageId="10" pageNumber="831" unit="m" value="648.0">648 m</quantity>
a.s.l.
</elevation>
;
<date box="[256,391,531,555]" pageId="10" pageNumber="831" value="2019-02-15">
<collectingDate box="[256,391,531,555]" pageId="10" pageNumber="831" value="2019-02-15">15 Feb. 2019</collectingDate>
</date>
;
<collectorName box="[400,583,531,555]" pageId="10" pageNumber="831">Cárdenas Johann</collectorName>
and
<collectorName box="[633,775,531,555]" pageId="10" pageNumber="831">Parada Jenny</collectorName>
leg.; open pasture;
<collectorName box="[376,435,564,588]" pageId="10" pageNumber="831">Crplt</collectorName>
5; UPTC-In-00056
</materialsCitation>
. •
<materialsCitation collectingDate="2019-02-15" collectorName="Vereda El Carmen &amp; Cardenas Johann &amp; Parada Jenny &amp; Crplt" country="Colombia" county="Villavicencio" elevation="648" latitude="4.140108" location="Farm" longLatPrecision="1" longitude="-73.660904" municipality="Vereda El Carmen" pageId="10" pageNumber="831" specimenCount="1" specimenCount-female="1" stateProvince="Meta">
<specimenCount box="[670,700,564,588]" pageId="10" pageNumber="831" type="female">1♀</specimenCount>
,
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.342" pageId="10" pageNumber="831" unit="mm" value="23.42">23.42 mm</quantity>
(mean);
<collectingRegion box="[317,375,598,621]" country="Colombia" name="Meta" pageId="10" pageNumber="831">Meta</collectingRegion>
,
<collectingCounty box="[392,539,598,622]" pageId="10" pageNumber="831">Villavicencio</collectingCounty>
,
<collectorName box="[556,771,598,622]" pageId="10" pageNumber="831">
<collectingMunicipality box="[556,771,598,622]" pageId="10" pageNumber="831">Vereda El Carmen</collectingMunicipality>
</collectorName>
, “Las Pesebreras”
<location LSID="urn:lsid:plazi:treatment:03F2DE0CA93FC025FF5B37C0FE2CE28A:8E8439C1A93FC025FE91343CFE7CE4F9" box="[354,414,632,655]" country="Colombia" county="Villavicencio" latitude="4.140108" longLatPrecision="1" longitude="-73.660904" municipality="Vereda El Carmen" name="Farm" pageId="10" pageNumber="831" stateProvince="Meta">Farm</location>
;
<geoCoordinate box="[423,586,630,655]" degrees="04" direction="north" minutes="08" orientation="latitude" pageId="10" pageNumber="831" precision="1" seconds="24.39" value="4.140108">04°0824.39″N</geoCoordinate>
,
<geoCoordinate box="[594,771,630,655]" degrees="073" direction="west" minutes="39" orientation="longitude" pageId="10" pageNumber="831" precision="1" seconds="39.24" value="-73.660904">073°3939.24″W</geoCoordinate>
;
<elevation box="[168,295,665,689]" metricMagnitude="2" metricUnit="m" metricValue="6.4799999999999995" pageId="10" pageNumber="831" unit="m" value="648.0">
<quantity box="[168,237,665,689]" metricMagnitude="2" metricUnit="m" metricValue="6.4799999999999995" pageId="10" pageNumber="831" unit="m" value="648.0">648 m</quantity>
a.s.l.
</elevation>
;
<date box="[309,449,665,689]" pageId="10" pageNumber="831" value="2019-02-15">
<collectingDate box="[309,449,665,689]" pageId="10" pageNumber="831" value="2019-02-15">15 Feb. 2019</collectingDate>
</date>
;
<collectorName box="[461,646,665,689]" pageId="10" pageNumber="831">Cárdenas Johann</collectorName>
and
<collectorName pageId="10" pageNumber="831">Parada Jenny</collectorName>
leg.; open pasture;
<collectorName box="[443,503,699,723]" pageId="10" pageNumber="831">Crplt</collectorName>
3; UPTC-In-00057
</materialsCitation>
.
</paragraph>
<paragraph blockId="10.[167,776,740,1067]" pageId="10" pageNumber="831">
<emphasis bold="true" box="[168,330,740,764]" pageId="10" pageNumber="831">Identification.</emphasis>
This species can be distinguished from other
<taxonomicName box="[233,459,774,798]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="831" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[233,371,774,798]" italics="true" pageId="10" pageNumber="831">Dichotomius</emphasis>
species
</taxonomicName>
by the head with transverse roughness, anterior edge of the clypeus margined with two rounded teeth, presence of a tubercle at the base of the eyes in the form of a large horn that is simple in males and small and forked towards the apex in females; pronotum strongly dotted with a vertical slope towards the anterior part, and a very conspicuous midline forming two small rounded protuberances (
<bibRefCitation author="Sarmiento-Garces R &amp; Amat-Garcia G" pageId="10" pageNumber="831" refId="ref18606" refString="Sarmiento-Garces R, Amat-Garcia G (2014) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae, Scarabaeinae) en Colombia. Universidad Nacional de Colombia, Bogota, 134 pp." type="book" year="2014">Sarmiento-Gárces and Amat-García 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="831" type="distribution">
<paragraph blockId="10.[167,777,1084,1276]" pageId="10" pageNumber="831">
<emphasis bold="true" box="[168,317,1084,1108]" pageId="10" pageNumber="831">Distribution.</emphasis>
In
<collectingCountry box="[353,457,1084,1108]" name="Colombia" pageId="10" pageNumber="831">Colombia</collectingCountry>
it is found in the Orinoco region, from the flatlands to sectors of the lowland foot- hills (
<bibRefCitation author="Sarmiento-Garces R &amp; Amat-Garcia G" box="[231,691,1151,1175]" pageId="10" pageNumber="831" refId="ref18606" refString="Sarmiento-Garces R, Amat-Garcia G (2014) Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae, Scarabaeinae) en Colombia. Universidad Nacional de Colombia, Bogota, 134 pp." type="book" year="2014">Sarmiento-Gárces and Amat-García 2014</bibRefCitation>
). In the study area, it was observed between
<quantity box="[558,678,1185,1209]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="10" pageNumber="831" unit="m" value="675.0" valueMax="700.0" valueMin="650.0">650700 m</quantity>
a.s.l. and detected only in open grassland habitat, so it is a rare species (seven individuals).
</paragraph>
</subSubSection>
</treatment>
</document>