145 lines
16 KiB
XML
145 lines
16 KiB
XML
<document id="A03DB9750FD287519B9D5F0599F4D1D9" ID-DOI="10.1206/3823.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5367923" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630259683279" checkinUser="carolina" docAuthor="Vea, Isabelle M. & Grimaldi, David A." docDate="2015" docId="03EF7A1BFFD1FFB302FBFC77FCCC4A8E" docLanguage="en" docName="N3823.pdf" docOrigin="American Museum Novitates 2015 (3823)" docSource="http://www.bioone.org/doi/abs/10.1206/3823.1" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Williamsicoccus megalops Vea & Grimaldi 2015, new species" docType="treatment" docVersion="5" lastPageNumber="51" masterDocId="FFD60263FFE3FF870328FFAEFFDC4E35" masterDocTitle="Diverse new scale insects (Hemiptera: Coccoidea) in amber from the Cretaceous and Eocene with a phylogenetic framework for fossil Coccoidea" masterLastPageNumber="80" masterPageNumber="1" pageNumber="51" updateTime="1699200754860" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1B245855F5C90631E4B68FD5443A2C71">Diverse new scale insects (Hemiptera: Coccoidea) in amber from the Cretaceous and Eocene with a phylogenetic framework for fossil Coccoidea</mods:title>
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<mods:name id="765FD7193031745281C39F89EABF088C" type="personal">
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<mods:roleTerm id="63DF04E3EB25270A9C1D45D7F225CB2D">Author</mods:roleTerm>
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<mods:namePart id="72EDE1C28B2C8BDC561D8C974FB636AF">Vea, Isabelle M.</mods:namePart>
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<mods:roleTerm id="70A320C233B32E9F8DFB279E150ABF5D">Author</mods:roleTerm>
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<mods:namePart id="41E13CE78BE4C6FE71210CDA02451F51">Grimaldi, David A.</mods:namePart>
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<mods:typeOfResource id="20332FBB1E98DDE05CC87F4C967036F5">text</mods:typeOfResource>
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<mods:title id="BEEF7A5DD63A5B6531043A61665F4127">American Museum Novitates</mods:title>
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<mods:part id="00D3318FDB2F7614CAA317D69C39EF54">
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<mods:date id="A5F779F6C2DC78954DC1D69ED8F86237">2015</mods:date>
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<mods:number id="F98C14F4CC735F73DC0C89B10DF6D3F3">2015-01-16</mods:number>
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<mods:number id="D723DD50E9AFE9B0A34E04A21B7BEFC8">2015</mods:number>
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<mods:number id="4AE9436E0990D2210E6234E909E18093">3823</mods:number>
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<mods:url id="95AE68062FBF954297AC29553681E8B7">http://www.bioone.org/doi/abs/10.1206/3823.1</mods:url>
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<mods:classification id="408F14ABF2DF9972610F4CE6A20488FF">journal article</mods:classification>
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<mods:identifier id="F3896D7C53EE3B53B8B8ED2680CFD466" type="DOI">10.1206/3823.1</mods:identifier>
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<treatment id="03EF7A1BFFD1FFB302FBFC77FCCC4A8E" ID-DOI="http://doi.org/10.5281/zenodo.5460530" ID-Zenodo-Dep="5460530" LSID="urn:lsid:plazi:treatment:03EF7A1BFFD1FFB302FBFC77FCCC4A8E" httpUri="http://treatment.plazi.org/id/03EF7A1BFFD1FFB302FBFC77FCCC4A8E" lastPageId="52" lastPageNumber="51" pageId="50" pageNumber="51">
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<subSubSection id="C35C9886FFD1FFB502FBFC77FC414DC6" box="[467,925,985,1012]" pageId="50" pageNumber="51" type="nomenclature">
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<paragraph id="8BF9CB0DFFD1FFB502FBFC77FC414DC6" blockId="50.[467,925,985,1012]" box="[467,925,985,1012]" pageId="50" pageNumber="51">
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<heading id="D0B17C61FFD1FFB502FBFC77FC414DC6" box="[467,925,985,1012]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="2">
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<emphasis id="B932171FFFD1FFB502FBFC77FCD14DC1" bold="true" box="[467,781,985,1012]" italics="true" pageId="50" pageNumber="51">
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<taxonomicName id="4C46B08EFFD1FFB502FBFC77FCDB4DC1" authority="Vea & Grimaldi, 2015" authorityName="Vea & Grimaldi" authorityYear="2015" box="[467,775,985,1012]" class="Insecta" family="Pseudococcidae" genus="Williamsicoccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="megalops" status="sp. nov.">Williamsicoccus megalops</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A201AA64FFD1FFB5003DFC74FC414DC6" box="[789,925,985,1012]" pageId="50" pageNumber="51" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C9886FFD1FFB50166FBA7FCFE4A11" box="[590,802,1033,1060]" pageId="50" pageNumber="51" type="description">
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<paragraph id="8BF9CB0DFFD1FFB50166FBA7FCFE4A11" blockId="50.[590,802,1033,1060]" box="[590,802,1033,1060]" pageId="50" pageNumber="51">
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<heading id="D0B17C61FFD1FFB50166FBA7FCFE4A11" box="[590,802,1033,1060]" fontSize="36" level="3" pageId="50" pageNumber="51" reason="7">
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<figureCitation id="137DD788FFD1FFB50166FBA7FD2B4A11" box="[590,759,1033,1060]" captionStart="FIGURE 18" captionStartId="51.[120,207,1308,1329]" captionTargetBox="[120,1224,224,1282]" captionTargetId="figure-190@51.[120,1224,224,1282]" captionTargetPageId="51" captionText="FIGURE 18. Photomicrographs of (A) dorsal and (B) ventral surfaces of Williamsicoccus megalops, n. sp., holotype 1582. (C) Dorsal and (D) ventral surfaces of Gilderius eukrinops, n. sp., holotype AMNH Bu-1594." figureDoi="http://doi.org/10.5281/zenodo.5367959" httpUri="https://zenodo.org/record/5367959/files/figure.png" pageId="50" pageNumber="51">Figures 18A–B</figureCitation>
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,
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<figureCitation id="137DD788FFD1FFB5002EFBA4FCFE4A11" box="[774,802,1034,1060]" captionStart="FIGURE 19" captionStartId="52.[121,207,1001,1022]" captionTargetBox="[120,1221,224,970]" captionTargetId="figure-271@52.[120,1224,224,975]" captionTargetPageId="52" captionText="FIGURE 19. Details of Williamsicoccus megalops, n. sp. (A) Dorsal view of head. (B) Ventral view of head. (C) Dorsal mesothorax. (D) Wing. (E) Connection of the forewing and hamulohaltere. (F) Leg" figureDoi="http://doi.org/10.5281/zenodo.5367961" httpUri="https://zenodo.org/record/5367961/files/figure.png" pageId="50" pageNumber="51">19</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C9886FFD1FFB50380FB9CFD644B1E" pageId="50" pageNumber="51" type="materials_examined">
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<paragraph id="8BF9CB0DFFD1FFB50380FB9CFDD94A44" blockId="50.[120,1225,1073,1810]" pageId="50" pageNumber="51">
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<materialsCitation id="3B2EC150FFD1FFB50380FB9CFDDD4A44" country="Lebanon" county="Lower Cretaceous" location="Central" municipality="Caza Baabda" pageId="50" pageNumber="51" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54FD75AFFFD1FFB50380FB9CFF3E4A79" box="[168,226,1074,1100]" pageId="50" pageNumber="51">TYPE</typeStatus>
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LOCALITY:
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<collectingCountry id="F3518B9DFFD1FFB50244FB9FFE084A7E" box="[364,468,1073,1099]" name="Lebanon" pageId="50" pageNumber="51">Lebanon</collectingCountry>
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:
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<location id="8E999DD6FFD1FFB502F7FB9FFDEA4A7E" LSID="urn:lsid:plazi:treatment:03EF7A1BFFD1FFB302FBFC77FCCC4A8E:8E999DD6FFD1FFB502F7FB9FFDEA4A7E" box="[479,566,1073,1099]" country="Lebanon" county="Lower Cretaceous" municipality="Caza Baabda" name="Central" pageId="50" pageNumber="51">Central</location>
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<collectingCountry id="F3518B9DFFD1FFB50115FB9FFD794A7E" box="[573,677,1073,1099]" name="Lebanon" pageId="50" pageNumber="51">Lebanon</collectingCountry>
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:
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<location id="8E999DD6FFD1FFB50198FB9FFCD34A7E" LSID="urn:lsid:plazi:treatment:03EF7A1BFFD1FFB302FBFC77FCCC4A8E:8E999DD6FFD1FFB50198FB9FFCD34A7E" box="[688,783,1073,1099]" country="Lebanon" county="Lower Cretaceous" municipality="Caza Baabda" name="Mdeyrij" pageId="50" pageNumber="51">Mdeyrij</location>
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/
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<location id="8E999DD6FFD1FFB50030FB9FFC4E4A7E" LSID="urn:lsid:plazi:treatment:03EF7A1BFFD1FFB302FBFC77FCCC4A8E:8E999DD6FFD1FFB50030FB9FFC4E4A7E" box="[792,914,1073,1099]" country="Lebanon" county="Lower Cretaceous" municipality="Caza Baabda" name="Hammana" pageId="50" pageNumber="51">Hammana</location>
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:
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<collectingMunicipality id="6B9D5177FFD1FFB500B4FB9CFBE84A7E" box="[924,1076,1073,1100]" pageId="50" pageNumber="51">Caza Baabda</collectingMunicipality>
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: Mouhafazet Jabal Loubnon.
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<collectingCounty id="6298B381FFD1FFB50207FBF9FDDD4A44" box="[303,513,1111,1137]" pageId="50" pageNumber="51">Lower Cretaceous</collectingCounty>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BF9CB0DFFD1FFB50380FBD3FD644B1E" blockId="50.[120,1225,1073,1810]" pageId="50" pageNumber="51">
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<materialsCitation id="3B2EC150FFD1FFB50380FBD3FD644B1E" country="Canada" location="Dany Azar" pageId="50" pageNumber="51" specimenCount="1" typeStatus="holotype">
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TYPE:
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<typeStatus id="54FD75AFFFD1FFB503D9FBD2FE804AA3" box="[241,348,1148,1174]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
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1582, alate male in a 4 × 4 ×
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<quantity id="4CBE66E8FFD1FFB501EDFBD2FCFC4AA2" box="[709,800,1148,1175]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="mm" value="0.5">0.5 mm</quantity>
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yellow, transparent, polished amber piece, embedded in
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<collectingCountry id="F3518B9DFFD1FFB50249FB0FFE654A8E" box="[353,441,1185,1211]" name="Canada" pageId="50" pageNumber="51">Canada</collectingCountry>
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balsam and mounted in epoxy with two coverslips, with a few particles of debris; specimen is well preserved, although the wings are hardly visible and antennae truncated. accessible views: ventral and dorsal;
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<location id="8E999DD6FFD1FFB501B9FB43FCD04B33" LSID="urn:lsid:plazi:treatment:03EF7A1BFFD1FFB302FBFC77FCCC4A8E:8E999DD6FFD1FFB501B9FB43FCD04B33" box="[657,780,1260,1287]" country="Canada" name="Dany Azar" pageId="50" pageNumber="51">Dany Azar</location>
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amber collection, provisionally deposited in the Muséum National d’Histoire Naturelle.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C9886FFD1FFB50380FA99FBBD4B43" pageId="50" pageNumber="51" type="etymology">
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<paragraph id="8BF9CB0DFFD1FFB50380FA99FBBD4B43" blockId="50.[120,1225,1073,1810]" pageId="50" pageNumber="51">
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ETYMOLOGY: The epithet
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<taxonomicName id="4C46B08EFFD1FFB502E5FA99FDE84B64" authorityName="Vea & Grimaldi" authorityYear="2015" box="[461,564,1335,1361]" class="Insecta" family="Pseudococcidae" genus="Williamsicoccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="megalops">
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<emphasis id="B932171FFFD1FFB502E5FA99FDE84B64" box="[461,564,1335,1361]" italics="true" pageId="50" pageNumber="51">megalops</emphasis>
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</taxonomicName>
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comes from Greek
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<emphasis id="B932171FFFD1FFB5003FFA99FCAC4B64" box="[791,880,1335,1361]" italics="true" pageId="50" pageNumber="51">megalos</emphasis>
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, meaning “large or great” and
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<emphasis id="B932171FFFD1FFB50350FAF3FF414B43" box="[120,157,1373,1398]" italics="true" pageId="50" pageNumber="51">ops</emphasis>
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, “eye,” referring to the large ventral eyes in this species and is a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C9886FFD1FFB50380FA2CFE114BA9" box="[168,461,1409,1436]" pageId="50" pageNumber="51" type="diagnosis">
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<paragraph id="8BF9CB0DFFD1FFB50380FA2CFE114BA9" blockId="50.[120,1225,1073,1810]" box="[168,461,1409,1436]" pageId="50" pageNumber="51">DIAGNOSIS: As for genus.</paragraph>
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</subSubSection>
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<subSubSection id="C35C9886FFD1FFB30380FA09FCCC4A8E" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51" type="description">
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<paragraph id="8BF9CB0DFFD1FFB30380FA09FCCC4A8E" blockId="50.[120,1225,1073,1810]" lastBlockId="52.[120,1224,1073,1212]" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
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DESCRIPTION: Body minute, total length 775 μm, widest at mesothorax, ca. 235 μm.
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<emphasis id="B932171FFFD1FFB507A0FA09FB154BF4" bold="true" box="[1160,1225,1447,1473]" pageId="50" pageNumber="51">Head</emphasis>
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(fig. 19AB): Round, ca. 210 μm wide, ca. 170 μm long. Each ocular sclerite with two pairs of simple eyes (a pair dorsally and a pair ventrally), each dorsal eye ca. 25 μm, ventral eye larger than dorsal eye (ca. 40 μm) and positioned closer together; with a pair of lateral ocelli (10 μm wide).
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<emphasis id="B932171FFFD1FFB503EEF992FEF24863" bold="true" box="[198,302,1596,1622]" pageId="50" pageNumber="51">Antenna</emphasis>
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(fig. 19A): At least 6-segmented, but both antennae broken on
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<typeStatus id="54FD75AFFFD1FFB5072BF992FBB64863" box="[1027,1130,1596,1622]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
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; lengths of segments (in μm): scape 25; pedicel 40; flagellar segments III to VI all filiform, subequal in length (ca 100 μm). Setae longer than segment width (ca. 35–60 μm).
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<emphasis id="B932171FFFD1FFB50088F929FC224894" bold="true" box="[928,1022,1671,1697]" pageId="50" pageNumber="51">Thorax:</emphasis>
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Head and thorax slightly separated. Prothorax membranous, ridges not observable. Prosternal structure not visible. Mesothorax
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<emphasis id="B932171FFFD1FFB5022FF97CFE8248D9" box="[263,350,1746,1772]" italics="true" pageId="50" pageNumber="51">dorsally</emphasis>
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: prescutum oval, bulging (85 μm wide, 72 μm long); scutum without a membranous area; scutellum oval, almost as wide as prescutum (fig. 19C). Setae not detected.
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<emphasis id="B932171FFFD1FFB407BBF956FF6E4BA4" italics="true" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">Ventrally</emphasis>
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: basisternum well developed with a median ridge.
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<emphasis id="B932171FFFD0FFB401C7FAD8FCE04BA5" bold="true" box="[751,828,1398,1424]" pageId="51" pageNumber="52">Wings</emphasis>
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(fig. 19D): Forewings elongate and thin, base narrow and round distally, ca. 885 μm long, 290 μm wide; subcostal ridge extending from the base of the wing to a little less than 3/4 wing length. Cubital ridge present, starting at wing base; alar setae and sensoria absent, microtrichia present. Alar lobe present. Hamulohalteres narrow, ca. 70 μm long; each with one hamulus (fig. 19E).
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<emphasis id="B932171FFFD0FFB40008F9A2FC8B4813" bold="true" box="[800,855,1548,1574]" pageId="51" pageNumber="52">Legs</emphasis>
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(fig. 19F): Long and slender, all subequal in length and shape, cuticular reticulations absent; measurements of foreleg: coxa 50 μm long; trochanter and femur, 150–160 μm long, 25–30 μm wide, with few short, hairlike setae; tibia: 130–160 μm long, 20 μm wide, with short hairlike setae, becoming numerous and spinose distally; tarsus 2-segmented, 45–50 μm long, 15 μm wide, tarsal digitules finely clavate; claw thin 10 μm, almost straight, digitules and denticles not observed.
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<emphasis id="B932171FFFD0FFB40069F968FC6348D5" bold="true" box="[833,959,1734,1760]" pageId="51" pageNumber="52">Abdomen:</emphasis>
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Ca. 345 μm long (315 μm without genital segment), 215 μm widest. Tergites and sternites well developed and segmentation easy to delineate. Abdominal setae not visible except for pleural setae on posteriormost segments. Tergite VII and VIII each side having wax filaments protruding out posteriad, indicating glandular pouches.
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<emphasis id="B932171FFFD7FFB302AFFBD2FD884AA3" bold="true" box="[391,596,1148,1174]" pageId="52" pageNumber="53">Genital segment:</emphasis>
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Penial sheath short subtubular (40 μm long, 50 μm at base). Aedeagus not visible. Eversible endophallus absent.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |